taxonID	type	description	language	source
535D8873FFE4FFCA21145885DE39FB24.taxon	type_taxon	Type species Diotima hirsutissima Simon, 1880 (considered a junior synonym of Amaurobius indistinctus O. Pickard-Cambridge, 1872).	en	Wang, Lu-Yu, Yang, Yun-Shan, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Review of the spider genus Devade Simon, 1885 (Araneae: Dictynidae) from China. European Journal of Taxonomy 989: 289-310, DOI: 10.5852/ejt.2025.989.2901, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2901/13141
535D8873FFE4FFCA21145885DE39FB24.taxon	diagnosis	Differential diagnosis Molecular analysis reveals that Saltonia Chamberlin & Ivie, 1942 is the sister taxon to Devade, and together they form a clade that is sister to another group containing Argyroneta Latreille, 1804 (Crews et al. 2020; Gorneau et al. 2023). Devade is most similar to the Palearctic monotypic water spider genus Argyroneta (Zhang & Wang 2017: fig. 7) in having a laminar retrolateral tibial apophysis, lacking distinct copulatory ducts, and having small, spherical spermathecae, but it differs by the distinct fold at the base of the cymbium in dorsal view (vs absent in Argyroneta), two large processes on the posterior arm of the conductor (vs only one), presence of a septum (vs absent), fertilization ducts originating from the posterior part of the spermathecae (vs prolateral), smaller size of less than 6 mm (vs> 8 mm), tracheal spiracle near spinnerets (vs near epigastral fold) and cribellum and calamistrum present (vs absent in Argyroneta). Devade is also similar to the Nearctic monotypic genus Saltonia Chamberlin & Ivie, 1942 (Chamberlin & Ivie 1942: 24, pl. 2 Figs 24 – 25; Crews et al. 2020: 914, fig. 22 a – l) in having a similar habitus, morphology of the male palp, and saline habitat, but it differs in the cymbial basidorsal fold fingernail edge-shaped present dorsally (vs curved, present retrolatrally at the base of cymbium in Saltonia) and epigyne with distinct epigynal septum (vs indistinct in Saltonia).	en	Wang, Lu-Yu, Yang, Yun-Shan, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Review of the spider genus Devade Simon, 1885 (Araneae: Dictynidae) from China. European Journal of Taxonomy 989: 289-310, DOI: 10.5852/ejt.2025.989.2901, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2901/13141
535D8873FFE4FFCA21145885DE39FB24.taxon	description	Description Small size (3 – 6 mm). Carapace yellowish brown to brown. Eight eyes. Fovea longitudinal. Cervical groove indistinct, radial furrows distinct. Chelicerae, yellow-brown. Labium and endites yellow-brown, longer than wide. Sternum yellow-brown and scutellate. Chelicerae with 2 – 3 promarginal and 2 retromarginal teeth. Legs yellow-brown. Leg formula: 4123. Calamistrum absent in male and weak in female (Fig. 1 C – D). Abdomen oval, dorsum yellow-brown to dark brown, with a few light chevrons dorsally. Venter of abdomen yellowish brown, with small and undivided cribellum (Fig. 1 A – B). Male palp (Figs 2 A – C, 3 C – E, 4 A – C, 5 C – E, 6 A – C, 7 C – E, 8 A – C, 9 C – E, 10 C – E). Tibia as long as cymbium or shorter. Retrolateral tibial apophysis laminar and directed ventrally. Cymbium longer than wide, narrow towards tip, with basidorsal fold (BF). Posterior arm of conductor with 2 large processes: mesal process narrow and hook-like, with pointed end; retrolateral process wide and laminar. Embolus originating at about 11: 30 – 12 o’clock position; distal part of embolus resting in long groove of conductor. Epigyne (Figs 2 D – E, 3 F – G, 4 D – E, 5 F – G, 6 D – E, 7 F – G, 8 D – E, 9 F – G, 10 F – G). Septum longer than wide, located in a fovea (D. longa and D. pulla) or reaching epigastral fold (D. mongolica and D. qiemuensis). Copulatory openings slit-like. Copulatory ducts extremely short, located on the ventral or retrolateral side of spermathecae. Spermathecae spherical, ½ times as long as septum. Fertilization ducts hook-like.	en	Wang, Lu-Yu, Yang, Yun-Shan, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Review of the spider genus Devade Simon, 1885 (Araneae: Dictynidae) from China. European Journal of Taxonomy 989: 289-310, DOI: 10.5852/ejt.2025.989.2901, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2901/13141
535D8873FFE4FFCA21145885DE39FB24.taxon	distribution	Distribution From the West Mediterranean to Eastern Transbaikalia and eastern Inner Mongolia (China), south to Algeria, Egypt, northern Pakistan, and ca 37 ° N in China, southern France, Serbia, South Urals (ca 52 ° N) and Tuva (ca 51 ° N) (Esyunin & Marusik 2001; WSC 2025).	en	Wang, Lu-Yu, Yang, Yun-Shan, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Review of the spider genus Devade Simon, 1885 (Araneae: Dictynidae) from China. European Journal of Taxonomy 989: 289-310, DOI: 10.5852/ejt.2025.989.2901, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2901/13141
535D8873FFE4FFCA21145885DE39FB24.taxon	discussion	Comments West Mediterranean species have never been redescribed, and Strinatinella spinosa Denis, 1957 and D. hirsutissima, considered a junior synonyms of Devade indistincta, may represent valid species.	en	Wang, Lu-Yu, Yang, Yun-Shan, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Review of the spider genus Devade Simon, 1885 (Araneae: Dictynidae) from China. European Journal of Taxonomy 989: 289-310, DOI: 10.5852/ejt.2025.989.2901, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2901/13141
535D8873FFE1FFC921BE5AC0DAB6FA6E.taxon	description	urn: lsid: zoobank. org: act: 46 EE 5301 - 5716 - 4 F 18 - AA 81 - F 657959 EE 8 C 9 Figs 2 – 3, 11	en	Wang, Lu-Yu, Yang, Yun-Shan, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Review of the spider genus Devade Simon, 1885 (Araneae: Dictynidae) from China. European Journal of Taxonomy 989: 289-310, DOI: 10.5852/ejt.2025.989.2901, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2901/13141
535D8873FFE1FFC921BE5AC0DAB6FA6E.taxon	vernacular_names	Chinese name 长带 ḍ	en	Wang, Lu-Yu, Yang, Yun-Shan, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Review of the spider genus Devade Simon, 1885 (Araneae: Dictynidae) from China. European Journal of Taxonomy 989: 289-310, DOI: 10.5852/ejt.2025.989.2901, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2901/13141
535D8873FFE1FFC921BE5AC0DAB6FA6E.taxon	diagnosis	Differential diagnosis The new species resembles D. qiemuensis (Figs 8 – 10) in having a longer palpal tibia (longer than half of the length of the cymbium), the fingernail edge-shaped basidorsal fold (BF) of the cymbium and similar spermathecae, but it differs by having the tibia as long as the cymbium (Figs 2 A – C, 3 C – E) (vs shorter, Figs 8 A – C, 9 C – E, 10 C – E), the anterior side of the mesal process of the conductor posterior arm with a notch (Figs 2 B, 3 D) (vs lacking, Figs 8 B, 9 D, 10 D), and the short septum, ca 2 times as long as wide (Figs 2 D, 3 F) (vs 3 times as long as wide Figs 8 D, 9 F, 10 F).	en	Wang, Lu-Yu, Yang, Yun-Shan, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Review of the spider genus Devade Simon, 1885 (Araneae: Dictynidae) from China. European Journal of Taxonomy 989: 289-310, DOI: 10.5852/ejt.2025.989.2901, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2901/13141
535D8873FFE1FFC921BE5AC0DAB6FA6E.taxon	etymology	Etymology The specific epithet comes from the Latin adjective ‘ longus ’, meaning ‘ long’, and refers to the longer tibia of the male palp.	en	Wang, Lu-Yu, Yang, Yun-Shan, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Review of the spider genus Devade Simon, 1885 (Araneae: Dictynidae) from China. European Journal of Taxonomy 989: 289-310, DOI: 10.5852/ejt.2025.989.2901, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2901/13141
535D8873FFE1FFC921BE5AC0DAB6FA6E.taxon	materials_examined	Type material Holotype CHINA • ♂; Xinjiang, Bole City, Aibi Lake; 44 ° 57 ′ 40 ″ N, 82 ° 38 ′ 38 ″ E; elev. 257 m; 12 Jun. 2014; L. Y. Wang and X. K. Jiang leg.; SWUC-T-DI- 20 - 01. Paratypes (5 ♂♂, 8 ♀♀) CHINA • 4 ♂♂, 8 ♀♀; same data as for holotype; SWUC-T-DI- 20 - 02 to SWUC-T-DI- 20 - 13 • 1 ♂; Xinjiang, Alashankou; 45 ° 11 ′ 33 ″ N, 82 ° 36 ′ 02 ″ E; elev. 328 m; 12 Jun. 2014; L. Y. Wang and X. K. Jiang leg.; SWUC-T-DI- 20 - 14.	en	Wang, Lu-Yu, Yang, Yun-Shan, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Review of the spider genus Devade Simon, 1885 (Araneae: Dictynidae) from China. European Journal of Taxonomy 989: 289-310, DOI: 10.5852/ejt.2025.989.2901, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2901/13141
535D8873FFE1FFC921BE5AC0DAB6FA6E.taxon	description	Description Male (holotype SWUC-T-DI- 20 - 01, Fig. 3 A) MEASUREMENTS. Total length 5.21. Carapace 2.34 long, 1.88 wide; abdomen 2.80 long, 1.54 wide. Eye sizes and interdistances: AME 0.11, ALE 0.13, PME 0.11, PLE, 0.11; AME – AME 0.06, AME – ALE 0.02, PME – PME 0.11, PME – PLE 0.08, ALE – PLE 0.05. MOA 0.33 long, anterior width 0.27, posterior width 0.33. Clypeus height 0.15. CARAPACE. Brown, cephalic part darker than thoracic; pattern weakly developed, with dark margins and dark radial stripes. CHELICERAE. With 3 promarginal and 2 retromarginal teeth. LEG MEASUREMENTS. I = 7.46 (2.00, 2.65, 1.73, 1.08); II = 7.25 (1.92, 2.47, 1.74, 1.12); III = 7.08 (1.92, 2.47, 1.74, 1.12); IV = 8.95 (2.30, 2.90, 2.45, 1.30). ABDOMEN. Light brown with ca 8 – 9 dark transverse chevrons. PALP (Figs 2 A – C, 3 C – E). Tibia long, almost as long as cymbium. Retrolateral tibial apophysis laminar and directed ventrally. Basidorsal fold (BF), fingernail edge-shaped in dorsal view. Posterior arm of conductor with 2 large processes: mesal process narrow and hook-like, with pointed tip; retrolateral process wide and laminar. Embolus originating at about 11: 30 o’clock position, distal part hidden by conductor. Female (paratype SWUC-T-DI- 20 - 02, Fig. 3 B) MEASUREMENTS. Total length 6.04. Carapace 2.39 long, 1.80 wide; abdomen 3.50 long, 2.25 wide. Eye sizes and interdistances: AME 0.11, ALE 0.12, PME 0.11, PLE, 0.12; AME – AME 0.07, AME – ALE 0.04, PME – PME 0.13, PME – PLE 0.09, ALE – PLE 0.06. MOA 0.33 long, anterior width 0.28, posterior width 0.35. Clypeus height 0.13. LEG MEASUREMENTS. I = 7.27 (2.11, 2.46, 1.61, 1.09); II = 6.81 (1.98, 2.25, 1.56, 1.02); III = 6.39 (1.66, 2.01, 1.74, 0.98); IV = 8.37 (2.23, 2.84, 2.20, 1.10). CARAPACE AND ABDOMEN. Same morphological pattern as in male except lighter in color. EPIGYNE (Figs 2 D, 3 F). Epigynal plate 1.7 times as wide as long; fovea as long as wide; septum 2 times as long as wide. Copulatory openings fissured, separated by 3 times width of septum. VULVA (Figs 2 E, 3 G). Copulatory ducts indistinct. Spermathecae globular, separated by almost 2 times their diameters, anterior part with glandular shallow extension. Fertilization ducts hook-like and diverging.	en	Wang, Lu-Yu, Yang, Yun-Shan, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Review of the spider genus Devade Simon, 1885 (Araneae: Dictynidae) from China. European Journal of Taxonomy 989: 289-310, DOI: 10.5852/ejt.2025.989.2901, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2901/13141
535D8873FFE1FFC921BE5AC0DAB6FA6E.taxon	distribution	Distribution China (Xinjiang) (Fig. 11).	en	Wang, Lu-Yu, Yang, Yun-Shan, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Review of the spider genus Devade Simon, 1885 (Araneae: Dictynidae) from China. European Journal of Taxonomy 989: 289-310, DOI: 10.5852/ejt.2025.989.2901, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2901/13141
535D8873FFE2FFC421265B99D934FA41.taxon	description	Figs 4 – 5, 11	en	Wang, Lu-Yu, Yang, Yun-Shan, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Review of the spider genus Devade Simon, 1885 (Araneae: Dictynidae) from China. European Journal of Taxonomy 989: 289-310, DOI: 10.5852/ejt.2025.989.2901, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2901/13141
535D8873FFE2FFC421265B99D934FA41.taxon	vernacular_names	Chinese name = 古带 ḍ	en	Wang, Lu-Yu, Yang, Yun-Shan, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Review of the spider genus Devade Simon, 1885 (Araneae: Dictynidae) from China. European Journal of Taxonomy 989: 289-310, DOI: 10.5852/ejt.2025.989.2901, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2901/13141
535D8873FFE2FFC421265B99D934FA41.taxon	diagnosis	Differential diagnosis The male of this species resembles D. pusilla (Lehtinen 1967: 450, fig. 284) in having a short palpal tibia, almost half the length of the cymbium, but it differs from the latter by the mesal process of the conductor posterior arm almost directed towards the ventral in retrolateral view (Figs 4 B, 5 D) (vs towards retrolateral), and the retrolateral process is pointed (Figs 4 B – C, 5 D – E) (vs rounded). The female of this species is most similar to D. qiemuensis (Figs 8 D – E, 9 F – G, 10 F – G) in having a long, narrow septum, almost reaching the epigastral fold, but it differs from the latter by the rounded end of the septum (Figs 4 D, 5 F) (vs truncate, Figs 8 D, 9 F, 10 F).	en	Wang, Lu-Yu, Yang, Yun-Shan, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Review of the spider genus Devade Simon, 1885 (Araneae: Dictynidae) from China. European Journal of Taxonomy 989: 289-310, DOI: 10.5852/ejt.2025.989.2901, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2901/13141
535D8873FFE2FFC421265B99D934FA41.taxon	materials_examined	Material examined CHINA • 13 ♂♂, 24 ♀♀; Xinjiang, Hutubi Co., Xidaqiao; 44 ° 12 ′ 04 ″ N, 86 ° 51 ′ 53 ″ E; elev. 526 m; 19 Apr. 2022; Y. S. Yang leg.; SWUC-DIDM- 01 to SWUC-DIDM- 37.	en	Wang, Lu-Yu, Yang, Yun-Shan, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Review of the spider genus Devade Simon, 1885 (Araneae: Dictynidae) from China. European Journal of Taxonomy 989: 289-310, DOI: 10.5852/ejt.2025.989.2901, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2901/13141
535D8873FFE2FFC421265B99D934FA41.taxon	description	Description Male (SWUC-DIDM- 01, Fig. 5 A) MEASUREMENTS. Total length 3.02. Carapace 1.47 long, 1.12 wide; abdomen 1.57 long, 0.94 wide. Eye sizes and interdistances: AME 0.09, ALE 0.08, PME 0.08, PLE, 0.09; AME – AME 0.05, AME – ALE 0.01, PME – PME 0.06, PME – PLE 0.04, ALE – PLE 0.02. MOA 0.23 long, anterior width 0.22, posterior width 0.25. Clypeus height 0.05. CARAPACE. Light with dark margins and dark radial stripes. CHELICERAE. With 2 promarginal and 2 retromarginal teeth. LEG MEASUREMENTS. I = 4.67 (1.33, 1.64, 1.02, 0.68); II = 4.46 (1.28, 1.51, 1.00, 0.67); III = 3.71 (1.13, 1.23, 0.91, 0.44); IV = 5.23 (1.47, 1.75, 1.34, 0.67). ABDOMEN. Light colored, with transverse bands. PALP (Figs 4 A – C, 5 C – E). Tibia short, almost as long as its distal width, less than half of cymbial length in retrolateral view. Retrolateral tibial apophysis tip directed almost ventrally in dorsal view. Cymbium 2 times as long as wide, tip as long as tibia; cymbial fold indistinct in dorsal view and easily seen in retrolateral view. Bulb about as long as wide; posterior arm of conductor (C) with two large processes: mesal process (MPA) short and hook-like; retrolateral process (RPA) wide and laminar, almost 3 times as long as mesal process. Embolus originating at about 12: 00 o’clock position, distal part hidden by conductor. Female (SWUC-DIDM- 02, Fig. 5 B) MEASUREMENTS. Total length 3.02. Carapace 1.24 long, 0.93 wide; abdomen 1.93 long, 1.45 wide. Eye sizes and interdistances: AME 0.06, ALE 0.08, PME 0.07, PLE, 0.08; AME – AME 0.05, AME – ALE 0.01, PME – PME 0.07, PME – PLE 0.05, ALE – PLE 0.03. MOA 0.20 long, anterior width 0.18, posterior width 0.22. Clypeus height 0.07. LEG MEASUREMENTS. I = 3.24 (0.99, 1.12, 0.67, 0.46); II = 3.10 (0.94, 1.04, 0.61, 0.51); III = 2.78 (0.80, 0.89, 0.63, 0.46); IV = 3.83 (0.99, 1.30, 1.01, 0.53). CARAPACE AND ABDOMEN. Same morphological pattern as in male except lighter in color. EPIGYNE (Figs 4 D, 5 F). Epigynal plate almost 2 times as wide as long, with anterior notch (No). Fovea V-shaped with distinct lateral margins, divided by septum. Septum reaching posterior margin, wider anteriorly. VULVA (Figs 4 E, 5 G). Copulatory ducts extremely short. Spermathecae globular, separated by more than 3.5 times their diameters.	en	Wang, Lu-Yu, Yang, Yun-Shan, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Review of the spider genus Devade Simon, 1885 (Araneae: Dictynidae) from China. European Journal of Taxonomy 989: 289-310, DOI: 10.5852/ejt.2025.989.2901, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2901/13141
535D8873FFE2FFC421265B99D934FA41.taxon	distribution	Distribution China (Xinjiang) (Fig. 11), Mongolia.	en	Wang, Lu-Yu, Yang, Yun-Shan, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Review of the spider genus Devade Simon, 1885 (Araneae: Dictynidae) from China. European Journal of Taxonomy 989: 289-310, DOI: 10.5852/ejt.2025.989.2901, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2901/13141
535D8873FFEFFFC321B25864DAB6F974.taxon	description	urn: lsid: zoobank. org: act: AD 372 ABA-DB 49 - 41 FE-B 25 B- 363 F 11 BA 1 E 7 F Figs 1, 6 – 7, 11	en	Wang, Lu-Yu, Yang, Yun-Shan, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Review of the spider genus Devade Simon, 1885 (Araneae: Dictynidae) from China. European Journal of Taxonomy 989: 289-310, DOI: 10.5852/ejt.2025.989.2901, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2901/13141
535D8873FFEFFFC321B25864DAB6F974.taxon	vernacular_names	Chinese name 黑带 ḍ	en	Wang, Lu-Yu, Yang, Yun-Shan, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Review of the spider genus Devade Simon, 1885 (Araneae: Dictynidae) from China. European Journal of Taxonomy 989: 289-310, DOI: 10.5852/ejt.2025.989.2901, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2901/13141
535D8873FFEFFFC321B25864DAB6F974.taxon	diagnosis	Differential diagnosis The new species resembles D. longa sp. nov. (Figs 2 – 3) in having a long tibia (longer than half the length of the cymbium), short septum (located in fovea), but differs by having the tibia shorter than the length of the cymbium (vs as long as cymbium), stronger and longer mesal process (as long as retrolateral process) (Figs 6 A – C, 7 C – E) (vs shorter than the length of the posterior arm, Figs 2 A – C, 3 C – E), and the copulatory openings located ectally from spermathecae (Figs 6 D, 7 F) (vs located mesally, Figs 2 D, 3 F).	en	Wang, Lu-Yu, Yang, Yun-Shan, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Review of the spider genus Devade Simon, 1885 (Araneae: Dictynidae) from China. European Journal of Taxonomy 989: 289-310, DOI: 10.5852/ejt.2025.989.2901, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2901/13141
535D8873FFEFFFC321B25864DAB6F974.taxon	etymology	Etymology The specific epithet comes from the Latin adjective ‘ pullus ’, meaning ‘ black’ and refers to the body color of the male.	en	Wang, Lu-Yu, Yang, Yun-Shan, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Review of the spider genus Devade Simon, 1885 (Araneae: Dictynidae) from China. European Journal of Taxonomy 989: 289-310, DOI: 10.5852/ejt.2025.989.2901, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2901/13141
535D8873FFEFFFC321B25864DAB6F974.taxon	materials_examined	Type material Holotype CHINA • ♂; Xinjiang, Yuli Co., No. 218 Road, Km 772; 41 ° 06 ′ 28 ″ N, 86 ° 30 ′ 39 ″ E; elev. 868 m; 1 Jun. 2014; L. Y. Wang and X. K. Jiang leg.; SWUC-T-DI- 21 - 01. Paratypes (5 ♂♂, 4 ♀♀) CHINA • 1 ♀; same data as for holotype; SWUC-T-DI- 21 - 02 • 5 ♂♂, 3 ♀♀; Hutubi Co., Liuhao Reservoir; 44 ° 17 ′ 14 ″ N, 86 ° 57 ′ 30 ″ E; elev. 456 m; 21 May 2022; Y. S. Yang leg.; SWUC-T-DI- 21 - 03 to SWUC-T-DI- 21 - 10.	en	Wang, Lu-Yu, Yang, Yun-Shan, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Review of the spider genus Devade Simon, 1885 (Araneae: Dictynidae) from China. European Journal of Taxonomy 989: 289-310, DOI: 10.5852/ejt.2025.989.2901, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2901/13141
535D8873FFEFFFC321B25864DAB6F974.taxon	description	Description Male (holotype SWUC-T-DI- 21 - 01, Fig. 7 A) MEASUREMENTS. Total length 4.28. Carapace 2.03 long, 1.56 wide; abdomen 2.22 long, 1.37 wide. Eye sizes and interdistances: AME 0.11, ALE 0.11, PME 0.11, PLE, 0.11; AME – AME 0.06, AME – ALE 0.02, PME – PME 0.10, PME – PLE 0.07, ALE – PLE 0.05. MOA 0.28 long, anterior width 0.25, posterior width 0.30. Clypeus height 0.11. CARAPACE. Dark brown, with darker margins and dark radial stripes. CHELICERAE. With 3 promarginal and 2 retromarginal teeth. LEG MEASUREMENTS. I = 6.61 (1.86, 2.26, 1.47, 1.02); II = 6.37 (1.78, 2.12, 1.50, 0.97); III = 5.94 (1.64, 1.73, 1.61, 0.96); IV = 7.80 (2.11, 2.45, 2.16, 1.08). ABDOMEN. Uniformly dark gray. PALP (Figs 6 A – C, 7 C – E). Tibia short, about 2 times as long as wide, almost half of cymbium length. Retrolateral tibial apophysis (RTA) laminar and tip almost pointed ventrally in dorsal view. Cymbium 2 times as long as wide. Basidorsal fold (BF) forms right angle, finger-like in retrolateral view. Posterior arm of conductor with 2 large processes: mesal process (MPA) hook-like, tip slightly twisted; retrolateral process (RPA) wide and laminar, as long as mesal process. Embolus originating at about 11: 30 o’clock position, distal part hidden by conductor. Female (paratype SWUC-T-DI- 21 - 02, Fig. 7 B) MEASUREMENTS. Total length 7.83. Carapace 2.95 long, 2.28 wide; abdomen 4.97 long, 3.66 wide. Eye sizes and interdistances: AME 0.11, ALE 0.13, PME 0.11, PLE, 0.12; AME – AME 0.07, AME – ALE 0.03, PME – PME 0.12, PME – PLE 0.08, ALE – PLE 0.05. MOA 0.31 long, anterior width 0.28, posterior width 0.34. Clypeus height 0.13. CARAPACE. Light brown with darker margins and radial stripes. LEG MEASUREMENTS. I = 7.04 (2.05, 2.42, 1.52, 1.05); II = 6.67 (1.93, 2.23, 1.49, 1.02); III = 6.15 (1.70, 1.94, 1.54, 0.97); IV = 8.12 (2.21, 2.74, 2.09, 1.08). ABDOMEN. Uniformly gray-brown. EPIGYNE (Figs 6 D, 7 F). Epigynal plate about 2 times as wide as long; fovea ca 1.3 times as wide as long; septum about ⅓ of fovea width, almost 2 times as long as wide. Copulatory openings (CO) slit-shaped, separated by 2 times the length of septum. VULVA (Figs 6 E, 7 G). Copulatory ducts indistinct, copulatory opening leads to spermathecae, located ectally. Spermathecae small and globular, separated by ca 2.5 times their diameters.	en	Wang, Lu-Yu, Yang, Yun-Shan, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Review of the spider genus Devade Simon, 1885 (Araneae: Dictynidae) from China. European Journal of Taxonomy 989: 289-310, DOI: 10.5852/ejt.2025.989.2901, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2901/13141
535D8873FFEFFFC321B25864DAB6F974.taxon	distribution	Distribution China (Xinjiang) (Fig. 11).	en	Wang, Lu-Yu, Yang, Yun-Shan, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Review of the spider genus Devade Simon, 1885 (Araneae: Dictynidae) from China. European Journal of Taxonomy 989: 289-310, DOI: 10.5852/ejt.2025.989.2901, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2901/13141
535D8873FFE8FFDC212E5891DA33F9E1.taxon	description	Figs 8 – 11	en	Wang, Lu-Yu, Yang, Yun-Shan, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Review of the spider genus Devade Simon, 1885 (Araneae: Dictynidae) from China. European Journal of Taxonomy 989: 289-310, DOI: 10.5852/ejt.2025.989.2901, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2901/13141
535D8873FFE8FFDC212E5891DA33F9E1.taxon	vernacular_names	Chinese name H 末带 ḍ	en	Wang, Lu-Yu, Yang, Yun-Shan, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Review of the spider genus Devade Simon, 1885 (Araneae: Dictynidae) from China. European Journal of Taxonomy 989: 289-310, DOI: 10.5852/ejt.2025.989.2901, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2901/13141
535D8873FFE8FFDC212E5891DA33F9E1.taxon	diagnosis	Differential diagnosis This species resembles D. longa sp. nov. (Figs 2 – 3) in having a longer tibia (longer than half the length of the cymbium), the fingernail edge-shaped basidorsal fold (BF) of the cymbium and similar spermathecae, but it differs by having the palpal tibia shorter than the length of the cymbium (Figs 8 A – C, 9 C – E, 10 C – E) (vs as long as the cymbium, Figs 2 A – C, 3 C – E), the anterior side of the mesal process of the conductor posterior arm with smooth tip (Figs 8 B, 9 D, 10 D) (vs has a notch, Figs 2 B, 3 D), and the septum 3 times as long as wide (Figs 8 D, 9 F, 10 F) (vs 2 times as long as wide Figs 2 D, 3 F).	en	Wang, Lu-Yu, Yang, Yun-Shan, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Review of the spider genus Devade Simon, 1885 (Araneae: Dictynidae) from China. European Journal of Taxonomy 989: 289-310, DOI: 10.5852/ejt.2025.989.2901, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2901/13141
535D8873FFE8FFDC212E5891DA33F9E1.taxon	materials_examined	Material examined CHINA – Hebei • 2 ♀♀; Zhangjiakou City, Zhangbei Co., Zhongdu; 41 ° 18 ′ 02 ″ N, 114 ° 44 ′ 39 ″ E; elev. 1350 m; 16 Jul. 2002; G. M. Tang leg.; SWUC-DIDQ- 01, SWUC-DIDQ- 02. – Inner Mongolia • 3 ♀♀; Alxa League, Ejina Banner, Badaoqiao Desert Park; 42 ° 02 ′ 25 ″ N, 101 ° 18 ′ 32 ″ E; elev. 924 m; 5 Jun. 2015; T. Lu and G. Q. Huang leg.; SWUC-DIDQ- 03 to SWUC-DIDQ- 05 • 1 ♀; Ejina Banner, Tian’e Lake; 42 ° 00 ′ 40 ″ N, 101 ° 35 ′ 14 ″ E; elev. 890 m; 5 Jun. 2015; T. Lu and G. Q. Huang leg.; SWUC-DIDQ- 06 • 1 ♂, 6 ♀♀; Ejina Banner, Juyan Lake; 42 ° 20 ′ 16 ″ N, 101 ° 15 ′ 01 ″ E; elev. 895 m; 5 Jun. 2015; T. Lu and G. Q. Huang leg.; SWUC-DIDQ- 07 to SWUC-DIDQ- 13 • 7 ♂♂, 11 ♀♀; Alxa Left Banner, Jilantai Salt Lake; 39 ° 43 ′ 16 ″ N, 105 ° 44 ′ 42 ″ E; elev. 1017 m; 7 Jun. 2015; T. Lu and G. Q. Huang leg.; SWUC-DIDQ- 14 to SWUC-DIDQ- 31 • 4 ♂♂, 7 ♀♀; Alxa Left Banner, Jilantai, Dongshawo; 39 ° 44 ′ 23 ″ N, 105 ° 46 ′ 29 ″ E; elev. 1024 m; 7 Jun. 2015; T. Lu and G. Q. Huang leg.; SWUC-DIDQ- 32 to SWUC-DIDQ- 42 • 1 ♀; Jilantai; 39 ° 44 ′ 31 ″ N, 105 ° 45 ′ 48 ″ E; elev. 1027 m; 7 Jun. 2015; T. Lu and G. Q. Huang leg.; SWUC-DIDQ- 43 • 2 ♂♂, 11 ♀♀; Alxa Left Banner, Qinggele; 40 ° 17 ′ 03 ″ N, 105 ° 51 ′ 12 ″ E; elev. 1165 m; 11 Jun. 2015; T. Lu and G. Q. Huang leg.; SWUC-DIDQ- 44 to SWUC-DIDQ- 56 • 5 ♀♀; Alxa Left Banner, Liutuan, Dongqing Lake; 40 ° 30 ′ 17 ″ N, 106 ° 30 ′ 23 ″ E; elev. 1030 m; 11 Jun. 2015; T. Lu and G. Q. Huang leg.; SWUC-DIDQ- 57 to SWUC-DIDQ- 61 • 1 ♀; Bayannur, Nalin Lake; 40 ° 31 ′ 34 ″ N, 106 ° 38 ′ 39 ″ E; elev. 1038 m; 11 Jun. 2015; T. Lu and G. Q. Huang leg.; SWUC-DIDQ- 62 • 1 ♂, 8 ♀♀: Bayannur, Urad Front Banner, Wuliangsuhai; 40 ° 51 ′ 34 ″ N, 108 ° 50 ′ 54 E; elev. 1025 m; 14 Jun. 2015; T. Lu and G. Q. Huang leg.; SWUC-DIDQ- 63 to SWUC-DIDQ- 71 • 1 ♀; Baotou City, Bailingmiao Town; 41 ° 40 ′ 32 ″ N, 110 ° 27 ′ 37 ″ E; elev. 1348 m; 17 Jun. 2015; T. Lu and G. Q. Huang leg.; SWUC-DIDQ- 72 • 6 ♂♂, 8 ♀♀; Chifeng City, Hexigten Banner, Dalinuoer; 43 ° 22 ′ 51 ″ N, 116 ° 39 ′ 39 ″ E; elev. 1221 m; 27 Jun. 2015; T. Lu and G. Q. Huang leg; SWUC-DIDQ- 73 to SWUC-DIDQ- 86 • 1 ♂; 3 ♀♀; Dalinuoer; 3 Jul. 2002; Z. S. Zhang leg.; SWUC-DIDQ- 87 to SWUC-DIDQ- 90. – Qinghai • 7 ♀♀; Golmud, Xiao Qaidam Lake; 37 ° 26 ′ 59 ″ N, 95 ° 30 ′ 40 ″ E; elev. 3170 m; 24 Jul. 2018; L. Y. Wang, Z. S. Wu and Y. N. Mu leg.; SWUC-DIDQ- 91 to SWUC-DIDQ- 97. – Xinjiang • 13 ♂♂, 9 ♀♀; Altay City, Fuhai Co., Ulungur Lake; 47 ° 14 ′ 32 ″ N, 87 ° 24 ′ 09 ″ E; elev. 506 m; 15 Jun. 2014; L. Y. Wang and X. K. Jiang leg; SWUC-DIDQ- 98 to SWUC-DIDQ- 119 • 2 ♂♂, 2 ♀♀; Bohu Co., Bosten Lake, Dahekou; 41 ° 54 ′ 29 ″ N, 86 ° 44 ′ 16 ″ E; elev. 1033 m; 12 May 2013; L. Y. Wang leg.; SWUC-DIDQ- 120 to SWUC-DIDQ- 123 • 1 ♂, 4 ♀♀; Korla, Yuli Co.; 40 ° 43 ′ 56 ″ N, 87 ° 20 ′ 08 ″ E: elev. 863 m; 1 Jun. 2014; L. Y. Wang and X. K. Jiang leg.; SWUC-DIDQ- 124 to SWUC-DIDQ- 128 • 24 ♂♂, 74 ♀♀; Ruoqiang Co., Taitema Lake; 39 ° 30 ′ 14 ″ N, 88 ° 18 ′ 54 ″ E; elev. 845 m; 1 Jun. 2014; L. Y. Wang and X. K. Jiang leg.; SWUC-DIDQ- 129 to SWUC-DIDQ- 226 • 11 ♂♂, 24 ♀♀; Taitema Lake; 10 May 2013; L. Y. Wang leg.; SWUC-DIDQ- 227 to SWUC-DIDQ- 261 • 1 ♂, 6 ♀♀; Ruoqiang Co., Washixia Township; 38 ° 41 ′ 30 ″ N, 86 ° 53 ′ 14 ″ E; elev. 1029 m; 10 May 2013; L. Y. Wang leg.; SWUC-DIDQ- 262 to SWUC-DIDQ- 268 • 1 ♂, 2 ♀♀; Qiemo Co.; 38 ° 12 ′ 13 ″ N, 85 ° 31 ′ 08 ″ E; elev. 1241 m; 11 May 2013; L. Y. Wang leg.; SWUC-DIDQ- 269 to SWUC-DIDQ- 271 • 2 ♂♂, 12 ♀♀; Qiemo Co.; 37 ° 57 ′ 47 ″ N, 84 ° 26 ′ 47 ″ E; elev. 1260 m; 2 Jun. 2014; L. Y. Wang and X. K. Jiang leg.; SWUC-DIDQ- 272 to SWUC-DIDQ- 285.	en	Wang, Lu-Yu, Yang, Yun-Shan, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Review of the spider genus Devade Simon, 1885 (Araneae: Dictynidae) from China. European Journal of Taxonomy 989: 289-310, DOI: 10.5852/ejt.2025.989.2901, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2901/13141
535D8873FFE8FFDC212E5891DA33F9E1.taxon	description	Description Male (SWUC-DIDQ- 272, Fig. 9 A) MEASUREMENTS. Total length 4.27. Carapace 1.93 long, 1.47 wide; abdomen 2.46 long, 1.47 wide. Eye sizes and interdistances: AME 0.09, ALE 0.11, PME 0.09, PLE, 0.09; AME – AME 0.09, AME – ALE 0.03, PME – PME 0.10, PME – PLE 0.07, ALE – PLE 0.05. MOA 0.30 long, anterior width 0.26, posterior width 0.28. Clypeus height 0.10. CARAPACE. Brown, cephalic part darker than thoracic part; with dark margins and dark radial stripes. CHELICERAE. With 3 promarginal and 2 retromarginal teeth. LEG MEASUREMENTS. I = 6.14 (1.75, 2.05, 1.36, 0.98); II = 6.11 (1.70, 1.98, 1.45, 0.98); III = 6.09 (1.62, 1.85, 1.60, 1.02); IV = 7.67 (2.04, 2.41, 2.06, 1.16). ABDOMEN. Light brown to gray with ca 7 – 8 dark transversal chevrons. PALP (Figs 8 A – C, 9 C – E, 10 C – E). Tibia long, longer than half the length of cymbium. Retrolateral tibial apophysis laminar and directed ventrally. Cymbium longer than wide, narrow towards tip, basidorsal fold (BF) rounded in dorsal view and fingernail edge-shaped in retrolateral view. Posterior arm of conductor with two large processes: mesal process narrow and hook-like, with pointed tip; retrolateral process wide and laminar, almost 2 times as long as mesal process. Embolus originating at about the 11: 00 o’clock position. Female (SWUC-DIDQ- 274, Fig. 9 B) MEASUREMENTS. Total length 6.13. Carapace 2.01 long, 1.41 wide; abdomen 4.14 long, 2.34 wide. Eye sizes and interdistances: AME 0.10, ALE 0.13, PME 0.10, PLE, 0.11; AME – AME 0.08, AME – ALE 0.02, PME – PME 0.10, PME – PLE 0.06, ALE – PLE 0.04. MOA 0.28 long, anterior width 0.28, posterior width 0.30. Clypeus height 0.10. LEG MEASUREMENTS. I = 5.59 (1.67, 1.89, 1.20, 0.83); II = 5.07 (1.36, 1.65, 1.18, 0.88); III = 4.98 (1.32, 1.46, 1.36, 0.86); IV = 6.59 (1.75, 2.12, 1.82, 0.90). CARAPACE AND ABDOMEN. Same morphological pattern as in male except lighter in color. EPIGYNE (Figs 8 D, 9 F, 10 F). Epigynal plate almost 2 times as wide as long. Septum 3 times as long as wide. Copulatory openings small, separated by two times the length of the septum. VULVA (Figs 8 E, 9 G, 10 G). Copulatory ducts extremely short. Spermathecae small and globular, separated by 2.5 times their diameters.	en	Wang, Lu-Yu, Yang, Yun-Shan, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Review of the spider genus Devade Simon, 1885 (Araneae: Dictynidae) from China. European Journal of Taxonomy 989: 289-310, DOI: 10.5852/ejt.2025.989.2901, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2901/13141
535D8873FFE8FFDC212E5891DA33F9E1.taxon	distribution	Distribution China (Hebei, Inner Mongolia, Qinghai, Xinjiang) (Fig. 11).	en	Wang, Lu-Yu, Yang, Yun-Shan, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Review of the spider genus Devade Simon, 1885 (Araneae: Dictynidae) from China. European Journal of Taxonomy 989: 289-310, DOI: 10.5852/ejt.2025.989.2901, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2901/13141
535D8873FFE8FFDC212E5891DA33F9E1.taxon	discussion	Remarks We presented the photographs of specimens of Devade qiemuensis collected from Qiemo County (type locality) and Inner Mongolia. While variations were observed in the septum of the epigyne among the female specimens, no differences were noted in the male specimens collected alongside the females in Inner Mongolia.	en	Wang, Lu-Yu, Yang, Yun-Shan, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Review of the spider genus Devade Simon, 1885 (Araneae: Dictynidae) from China. European Journal of Taxonomy 989: 289-310, DOI: 10.5852/ejt.2025.989.2901, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2901/13141
