taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
0747A6D3C78753E185657B483AA6560F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1385816	https://doi.org/10.3897/phytokeys.260.162975.figure1	Figure 1. Cyathula achyranthoides. A. Leaves; B. Inflorescence (Nicaragua, El Brujo, 11.97984, - 86.26455). Photographer: Sune Holt.	Figure 1. Cyathula achyranthoides. A. Leaves; B. Inflorescence (Nicaragua, El Brujo, 11.97984, - 86.26455). Photographer: Sune Holt.	2025-08-01	Sukhorukov, Alexander P.;Sennikov, Alexander N.;Cheek, Martin;Nusbaumer, Louis;Kushunina, Maria;Timonin, Alexander C.		Zenodo	biologists	Sukhorukov, Alexander P.;Sennikov, Alexander N.;Cheek, Martin;Nusbaumer, Louis;Kushunina, Maria;Timonin, Alexander C.			
0747A6D3C78753E185657B483AA6560F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1385817	https://doi.org/10.3897/phytokeys.260.162975.figure2	Figure 2. Cymes of studied Cyathula species. A. C. achyranthoides; B. C. brevispicata; C. Sterile flower of C. brevispicata with bracteoles; D. C. fernando-poensis; E. C. geminata; F. C. prostrata. Abbreviations: br 1, br 2 – bracteoles of the first and second orders, sp – sterile part, ff – fertile flower. Scale bars: 1 mm.	Figure 2. Cymes of studied Cyathula species. A. C. achyranthoides; B. C. brevispicata; C. Sterile flower of C. brevispicata with bracteoles; D. C. fernando-poensis; E. C. geminata; F. C. prostrata. Abbreviations: br 1, br 2 – bracteoles of the first and second orders, sp – sterile part, ff – fertile flower. Scale bars: 1 mm.	2025-08-01	Sukhorukov, Alexander P.;Sennikov, Alexander N.;Cheek, Martin;Nusbaumer, Louis;Kushunina, Maria;Timonin, Alexander C.		Zenodo	biologists	Sukhorukov, Alexander P.;Sennikov, Alexander N.;Cheek, Martin;Nusbaumer, Louis;Kushunina, Maria;Timonin, Alexander C.			
0747A6D3C78753E185657B483AA6560F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1385818	https://doi.org/10.3897/phytokeys.260.162975.figure3	Figure 3. Simplified cyme architecture in the Cyathula achyranthoides group and related C. prostrata. A, B. Asymmetric cymes of C. achyranthoides and C. geminata including the common type (A) with a sterile part with one sterile flower (sf) located on one side of the fertile flower (ff), and deviating type (B) with one sessile rudimentary flower or bracteole (s); C. Symmetric cyme of C. aethiopica, C. brevispicata and C. fernando-poensis with one fertile flower (ff) and two sterile flowers (sf) located laterally; D. Symmetric cyme of C. prostrata with three fertile flowers and sterile parts with sterile flowers. Abbreviations: br 1, br 2, br 3, and br 4 – uncinate or non-uncinate bracteoles of different orders, X – rudimentary flowers of the sterile part.	Figure 3. Simplified cyme architecture in the Cyathula achyranthoides group and related C. prostrata. A, B. Asymmetric cymes of C. achyranthoides and C. geminata including the common type (A) with a sterile part with one sterile flower (sf) located on one side of the fertile flower (ff), and deviating type (B) with one sessile rudimentary flower or bracteole (s); C. Symmetric cyme of C. aethiopica, C. brevispicata and C. fernando-poensis with one fertile flower (ff) and two sterile flowers (sf) located laterally; D. Symmetric cyme of C. prostrata with three fertile flowers and sterile parts with sterile flowers. Abbreviations: br 1, br 2, br 3, and br 4 – uncinate or non-uncinate bracteoles of different orders, X – rudimentary flowers of the sterile part.	2025-08-01	Sukhorukov, Alexander P.;Sennikov, Alexander N.;Cheek, Martin;Nusbaumer, Louis;Kushunina, Maria;Timonin, Alexander C.		Zenodo	biologists	Sukhorukov, Alexander P.;Sennikov, Alexander N.;Cheek, Martin;Nusbaumer, Louis;Kushunina, Maria;Timonin, Alexander C.			
0747A6D3C78753E185657B483AA6560F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1385819	https://doi.org/10.3897/phytokeys.260.162975.figure4	Figure 4. Distribution map of Cyathula achyranthoides.	Figure 4. Distribution map of Cyathula achyranthoides.	2025-08-01	Sukhorukov, Alexander P.;Sennikov, Alexander N.;Cheek, Martin;Nusbaumer, Louis;Kushunina, Maria;Timonin, Alexander C.		Zenodo	biologists	Sukhorukov, Alexander P.;Sennikov, Alexander N.;Cheek, Martin;Nusbaumer, Louis;Kushunina, Maria;Timonin, Alexander C.			
D1D743B3E4875475957CD1F752D21BB0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1385820	https://doi.org/10.3897/phytokeys.260.162975.figure5	Figure 5. Holotype of Cyathula aethiopica.	Figure 5. Holotype of Cyathula aethiopica.	2025-08-01	Sukhorukov, Alexander P.;Sennikov, Alexander N.;Cheek, Martin;Nusbaumer, Louis;Kushunina, Maria;Timonin, Alexander C.		Zenodo	biologists	Sukhorukov, Alexander P.;Sennikov, Alexander N.;Cheek, Martin;Nusbaumer, Louis;Kushunina, Maria;Timonin, Alexander C.			
D1D743B3E4875475957CD1F752D21BB0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1385818	https://doi.org/10.3897/phytokeys.260.162975.figure3	Figure 3. Simplified cyme architecture in the Cyathula achyranthoides group and related C. prostrata. A, B. Asymmetric cymes of C. achyranthoides and C. geminata including the common type (A) with a sterile part with one sterile flower (sf) located on one side of the fertile flower (ff), and deviating type (B) with one sessile rudimentary flower or bracteole (s); C. Symmetric cyme of C. aethiopica, C. brevispicata and C. fernando-poensis with one fertile flower (ff) and two sterile flowers (sf) located laterally; D. Symmetric cyme of C. prostrata with three fertile flowers and sterile parts with sterile flowers. Abbreviations: br 1, br 2, br 3, and br 4 – uncinate or non-uncinate bracteoles of different orders, X – rudimentary flowers of the sterile part.	Figure 3. Simplified cyme architecture in the Cyathula achyranthoides group and related C. prostrata. A, B. Asymmetric cymes of C. achyranthoides and C. geminata including the common type (A) with a sterile part with one sterile flower (sf) located on one side of the fertile flower (ff), and deviating type (B) with one sessile rudimentary flower or bracteole (s); C. Symmetric cyme of C. aethiopica, C. brevispicata and C. fernando-poensis with one fertile flower (ff) and two sterile flowers (sf) located laterally; D. Symmetric cyme of C. prostrata with three fertile flowers and sterile parts with sterile flowers. Abbreviations: br 1, br 2, br 3, and br 4 – uncinate or non-uncinate bracteoles of different orders, X – rudimentary flowers of the sterile part.	2025-08-01	Sukhorukov, Alexander P.;Sennikov, Alexander N.;Cheek, Martin;Nusbaumer, Louis;Kushunina, Maria;Timonin, Alexander C.		Zenodo	biologists	Sukhorukov, Alexander P.;Sennikov, Alexander N.;Cheek, Martin;Nusbaumer, Louis;Kushunina, Maria;Timonin, Alexander C.			
D1D743B3E4875475957CD1F752D21BB0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1385821	https://doi.org/10.3897/phytokeys.260.162975.figure6	Figure 6. Distribution map of Cyathula aethiopica.	Figure 6. Distribution map of Cyathula aethiopica.	2025-08-01	Sukhorukov, Alexander P.;Sennikov, Alexander N.;Cheek, Martin;Nusbaumer, Louis;Kushunina, Maria;Timonin, Alexander C.		Zenodo	biologists	Sukhorukov, Alexander P.;Sennikov, Alexander N.;Cheek, Martin;Nusbaumer, Louis;Kushunina, Maria;Timonin, Alexander C.			
D5DE1CF89C1D5144B44CB6560B68C925.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1385822	https://doi.org/10.3897/phytokeys.260.162975.figure7	Figure 7. Cyathula brevispicata. A. General habit; B. Inflorescence (Madagascar, Vohemar Prefecture, forêt de Sorata). Photographer: Alessandra Havinga.	Figure 7. Cyathula brevispicata. A. General habit; B. Inflorescence (Madagascar, Vohemar Prefecture, forêt de Sorata). Photographer: Alessandra Havinga.	2025-08-01	Sukhorukov, Alexander P.;Sennikov, Alexander N.;Cheek, Martin;Nusbaumer, Louis;Kushunina, Maria;Timonin, Alexander C.		Zenodo	biologists	Sukhorukov, Alexander P.;Sennikov, Alexander N.;Cheek, Martin;Nusbaumer, Louis;Kushunina, Maria;Timonin, Alexander C.			
D5DE1CF89C1D5144B44CB6560B68C925.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1385823	https://doi.org/10.3897/phytokeys.260.162975.figure8	Figure 8. Holotype of Cyathula brevispicata.	Figure 8. Holotype of Cyathula brevispicata.	2025-08-01	Sukhorukov, Alexander P.;Sennikov, Alexander N.;Cheek, Martin;Nusbaumer, Louis;Kushunina, Maria;Timonin, Alexander C.		Zenodo	biologists	Sukhorukov, Alexander P.;Sennikov, Alexander N.;Cheek, Martin;Nusbaumer, Louis;Kushunina, Maria;Timonin, Alexander C.			
D5DE1CF89C1D5144B44CB6560B68C925.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1385817	https://doi.org/10.3897/phytokeys.260.162975.figure2	Figure 2. Cymes of studied Cyathula species. A. C. achyranthoides; B. C. brevispicata; C. Sterile flower of C. brevispicata with bracteoles; D. C. fernando-poensis; E. C. geminata; F. C. prostrata. Abbreviations: br 1, br 2 – bracteoles of the first and second orders, sp – sterile part, ff – fertile flower. Scale bars: 1 mm.	Figure 2. Cymes of studied Cyathula species. A. C. achyranthoides; B. C. brevispicata; C. Sterile flower of C. brevispicata with bracteoles; D. C. fernando-poensis; E. C. geminata; F. C. prostrata. Abbreviations: br 1, br 2 – bracteoles of the first and second orders, sp – sterile part, ff – fertile flower. Scale bars: 1 mm.	2025-08-01	Sukhorukov, Alexander P.;Sennikov, Alexander N.;Cheek, Martin;Nusbaumer, Louis;Kushunina, Maria;Timonin, Alexander C.		Zenodo	biologists	Sukhorukov, Alexander P.;Sennikov, Alexander N.;Cheek, Martin;Nusbaumer, Louis;Kushunina, Maria;Timonin, Alexander C.			
D5DE1CF89C1D5144B44CB6560B68C925.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1385818	https://doi.org/10.3897/phytokeys.260.162975.figure3	Figure 3. Simplified cyme architecture in the Cyathula achyranthoides group and related C. prostrata. A, B. Asymmetric cymes of C. achyranthoides and C. geminata including the common type (A) with a sterile part with one sterile flower (sf) located on one side of the fertile flower (ff), and deviating type (B) with one sessile rudimentary flower or bracteole (s); C. Symmetric cyme of C. aethiopica, C. brevispicata and C. fernando-poensis with one fertile flower (ff) and two sterile flowers (sf) located laterally; D. Symmetric cyme of C. prostrata with three fertile flowers and sterile parts with sterile flowers. Abbreviations: br 1, br 2, br 3, and br 4 – uncinate or non-uncinate bracteoles of different orders, X – rudimentary flowers of the sterile part.	Figure 3. Simplified cyme architecture in the Cyathula achyranthoides group and related C. prostrata. A, B. Asymmetric cymes of C. achyranthoides and C. geminata including the common type (A) with a sterile part with one sterile flower (sf) located on one side of the fertile flower (ff), and deviating type (B) with one sessile rudimentary flower or bracteole (s); C. Symmetric cyme of C. aethiopica, C. brevispicata and C. fernando-poensis with one fertile flower (ff) and two sterile flowers (sf) located laterally; D. Symmetric cyme of C. prostrata with three fertile flowers and sterile parts with sterile flowers. Abbreviations: br 1, br 2, br 3, and br 4 – uncinate or non-uncinate bracteoles of different orders, X – rudimentary flowers of the sterile part.	2025-08-01	Sukhorukov, Alexander P.;Sennikov, Alexander N.;Cheek, Martin;Nusbaumer, Louis;Kushunina, Maria;Timonin, Alexander C.		Zenodo	biologists	Sukhorukov, Alexander P.;Sennikov, Alexander N.;Cheek, Martin;Nusbaumer, Louis;Kushunina, Maria;Timonin, Alexander C.			
D5DE1CF89C1D5144B44CB6560B68C925.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1385824	https://doi.org/10.3897/phytokeys.260.162975.figure9	Figure 9. Distribution map of Cyathula brevispicata.	Figure 9. Distribution map of Cyathula brevispicata.	2025-08-01	Sukhorukov, Alexander P.;Sennikov, Alexander N.;Cheek, Martin;Nusbaumer, Louis;Kushunina, Maria;Timonin, Alexander C.		Zenodo	biologists	Sukhorukov, Alexander P.;Sennikov, Alexander N.;Cheek, Martin;Nusbaumer, Louis;Kushunina, Maria;Timonin, Alexander C.			
E047AC9E523E5D689AACB7177195D957.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1385817	https://doi.org/10.3897/phytokeys.260.162975.figure2	Figure 2. Cymes of studied Cyathula species. A. C. achyranthoides; B. C. brevispicata; C. Sterile flower of C. brevispicata with bracteoles; D. C. fernando-poensis; E. C. geminata; F. C. prostrata. Abbreviations: br 1, br 2 – bracteoles of the first and second orders, sp – sterile part, ff – fertile flower. Scale bars: 1 mm.	Figure 2. Cymes of studied Cyathula species. A. C. achyranthoides; B. C. brevispicata; C. Sterile flower of C. brevispicata with bracteoles; D. C. fernando-poensis; E. C. geminata; F. C. prostrata. Abbreviations: br 1, br 2 – bracteoles of the first and second orders, sp – sterile part, ff – fertile flower. Scale bars: 1 mm.	2025-08-01	Sukhorukov, Alexander P.;Sennikov, Alexander N.;Cheek, Martin;Nusbaumer, Louis;Kushunina, Maria;Timonin, Alexander C.		Zenodo	biologists	Sukhorukov, Alexander P.;Sennikov, Alexander N.;Cheek, Martin;Nusbaumer, Louis;Kushunina, Maria;Timonin, Alexander C.			
E047AC9E523E5D689AACB7177195D957.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1385818	https://doi.org/10.3897/phytokeys.260.162975.figure3	Figure 3. Simplified cyme architecture in the Cyathula achyranthoides group and related C. prostrata. A, B. Asymmetric cymes of C. achyranthoides and C. geminata including the common type (A) with a sterile part with one sterile flower (sf) located on one side of the fertile flower (ff), and deviating type (B) with one sessile rudimentary flower or bracteole (s); C. Symmetric cyme of C. aethiopica, C. brevispicata and C. fernando-poensis with one fertile flower (ff) and two sterile flowers (sf) located laterally; D. Symmetric cyme of C. prostrata with three fertile flowers and sterile parts with sterile flowers. Abbreviations: br 1, br 2, br 3, and br 4 – uncinate or non-uncinate bracteoles of different orders, X – rudimentary flowers of the sterile part.	Figure 3. Simplified cyme architecture in the Cyathula achyranthoides group and related C. prostrata. A, B. Asymmetric cymes of C. achyranthoides and C. geminata including the common type (A) with a sterile part with one sterile flower (sf) located on one side of the fertile flower (ff), and deviating type (B) with one sessile rudimentary flower or bracteole (s); C. Symmetric cyme of C. aethiopica, C. brevispicata and C. fernando-poensis with one fertile flower (ff) and two sterile flowers (sf) located laterally; D. Symmetric cyme of C. prostrata with three fertile flowers and sterile parts with sterile flowers. Abbreviations: br 1, br 2, br 3, and br 4 – uncinate or non-uncinate bracteoles of different orders, X – rudimentary flowers of the sterile part.	2025-08-01	Sukhorukov, Alexander P.;Sennikov, Alexander N.;Cheek, Martin;Nusbaumer, Louis;Kushunina, Maria;Timonin, Alexander C.		Zenodo	biologists	Sukhorukov, Alexander P.;Sennikov, Alexander N.;Cheek, Martin;Nusbaumer, Louis;Kushunina, Maria;Timonin, Alexander C.			
E047AC9E523E5D689AACB7177195D957.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1385825	https://doi.org/10.3897/phytokeys.260.162975.figure10	Figure 10. Distribution map of Cyathula fernando-poensis.	Figure 10. Distribution map of Cyathula fernando-poensis.	2025-08-01	Sukhorukov, Alexander P.;Sennikov, Alexander N.;Cheek, Martin;Nusbaumer, Louis;Kushunina, Maria;Timonin, Alexander C.		Zenodo	biologists	Sukhorukov, Alexander P.;Sennikov, Alexander N.;Cheek, Martin;Nusbaumer, Louis;Kushunina, Maria;Timonin, Alexander C.			
1D9D5B31A4B45C4EBE488F43D4C9B43F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1385817	https://doi.org/10.3897/phytokeys.260.162975.figure2	Figure 2. Cymes of studied Cyathula species. A. C. achyranthoides; B. C. brevispicata; C. Sterile flower of C. brevispicata with bracteoles; D. C. fernando-poensis; E. C. geminata; F. C. prostrata. Abbreviations: br 1, br 2 – bracteoles of the first and second orders, sp – sterile part, ff – fertile flower. Scale bars: 1 mm.	Figure 2. Cymes of studied Cyathula species. A. C. achyranthoides; B. C. brevispicata; C. Sterile flower of C. brevispicata with bracteoles; D. C. fernando-poensis; E. C. geminata; F. C. prostrata. Abbreviations: br 1, br 2 – bracteoles of the first and second orders, sp – sterile part, ff – fertile flower. Scale bars: 1 mm.	2025-08-01	Sukhorukov, Alexander P.;Sennikov, Alexander N.;Cheek, Martin;Nusbaumer, Louis;Kushunina, Maria;Timonin, Alexander C.		Zenodo	biologists	Sukhorukov, Alexander P.;Sennikov, Alexander N.;Cheek, Martin;Nusbaumer, Louis;Kushunina, Maria;Timonin, Alexander C.			
1D9D5B31A4B45C4EBE488F43D4C9B43F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1385818	https://doi.org/10.3897/phytokeys.260.162975.figure3	Figure 3. Simplified cyme architecture in the Cyathula achyranthoides group and related C. prostrata. A, B. Asymmetric cymes of C. achyranthoides and C. geminata including the common type (A) with a sterile part with one sterile flower (sf) located on one side of the fertile flower (ff), and deviating type (B) with one sessile rudimentary flower or bracteole (s); C. Symmetric cyme of C. aethiopica, C. brevispicata and C. fernando-poensis with one fertile flower (ff) and two sterile flowers (sf) located laterally; D. Symmetric cyme of C. prostrata with three fertile flowers and sterile parts with sterile flowers. Abbreviations: br 1, br 2, br 3, and br 4 – uncinate or non-uncinate bracteoles of different orders, X – rudimentary flowers of the sterile part.	Figure 3. Simplified cyme architecture in the Cyathula achyranthoides group and related C. prostrata. A, B. Asymmetric cymes of C. achyranthoides and C. geminata including the common type (A) with a sterile part with one sterile flower (sf) located on one side of the fertile flower (ff), and deviating type (B) with one sessile rudimentary flower or bracteole (s); C. Symmetric cyme of C. aethiopica, C. brevispicata and C. fernando-poensis with one fertile flower (ff) and two sterile flowers (sf) located laterally; D. Symmetric cyme of C. prostrata with three fertile flowers and sterile parts with sterile flowers. Abbreviations: br 1, br 2, br 3, and br 4 – uncinate or non-uncinate bracteoles of different orders, X – rudimentary flowers of the sterile part.	2025-08-01	Sukhorukov, Alexander P.;Sennikov, Alexander N.;Cheek, Martin;Nusbaumer, Louis;Kushunina, Maria;Timonin, Alexander C.		Zenodo	biologists	Sukhorukov, Alexander P.;Sennikov, Alexander N.;Cheek, Martin;Nusbaumer, Louis;Kushunina, Maria;Timonin, Alexander C.			
1D9D5B31A4B45C4EBE488F43D4C9B43F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1385826	https://doi.org/10.3897/phytokeys.260.162975.figure11	Figure 11. Distribution map of Cyathula geminata.	Figure 11. Distribution map of Cyathula geminata.	2025-08-01	Sukhorukov, Alexander P.;Sennikov, Alexander N.;Cheek, Martin;Nusbaumer, Louis;Kushunina, Maria;Timonin, Alexander C.		Zenodo	biologists	Sukhorukov, Alexander P.;Sennikov, Alexander N.;Cheek, Martin;Nusbaumer, Louis;Kushunina, Maria;Timonin, Alexander C.			
1D9D5B31A4B45C4EBE488F43D4C9B43F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1385827	https://doi.org/10.3897/phytokeys.260.162975.figure12	Figure 12. A representative specimen of Cyathula geminata.	Figure 12. A representative specimen of Cyathula geminata.	2025-08-01	Sukhorukov, Alexander P.;Sennikov, Alexander N.;Cheek, Martin;Nusbaumer, Louis;Kushunina, Maria;Timonin, Alexander C.		Zenodo	biologists	Sukhorukov, Alexander P.;Sennikov, Alexander N.;Cheek, Martin;Nusbaumer, Louis;Kushunina, Maria;Timonin, Alexander C.			
