identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
309E7C6FC7555525A896C31DA8ACA26B.text	309E7C6FC7555525A896C31DA8ACA26B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hedyotis austroguangxiensis M. D. Yuan & R. J. Wang 2025	<div><p>2. Hedyotis austroguangxiensis M. D. Yuan &amp; R. J. Wang sp. nov.</p><p>Figs 4, 5</p><p>Type.</p><p>CHINA • Guangxi: Chongzuo City, Ningming County, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.52156&amp;materialsCitation.latitude=21.711643" title="Search Plazi for locations around (long 107.52156/lat 21.711643)">Tongmian Town</a>, roadside, 21.711643, 107.521562, ca. 830 m a. s. l., flowering, long-styled flower, 24 April 2021, Ming-Deng Yuan &amp; Yi-Da Xu YS 430 (holotype: IBSC [0868775]); isotype: IBSC [0868776] ).</p><p>Diagnosis.</p><p>Hedyotis austroguangxiensis is similar to H. taishanensis G. T. Wang (Wang et al. 2018) and H. shenzhenensis Tao Chen (2007) in having shortened internodes at the base and terminal, compound-cymose inflorescences. But it can be differed from H. taishanensis by its terete stem and inflorescence rachis, obovate to ovate or oblong leaves, 3–6 secondary veins on each side, and longer corolla tubes; it differs from H. shenzhenensis by its terete stems and inflorescence axes, smaller leaves, and longer corolla tubes (Table 3).</p><p>Description.</p><p>Perennial herbs, erect, 20–50 cm tall. Stem terete, glabrous, internodes 0.5–5.0 cm long at the base and 5.0–13.0 cm long toward top. Leaves opposite, sessile, blade 2.5–15.0 × 1.5–7.0 cm, ovate to oblong, apex acuminate or obtuse, base broadly cuneate to rounded, blade coriaceous, dark green adaxially and grayish green abaxially, glabrous on both sides; midrib concave adaxially and prominent abaxially; secondary veins 4–6 on each side, inconspicuous on both sides. Stipules interpetiolar, 1.5–2.5 mm long, broadly ovate or triangular, apex acuminate, margin toothed with tipped colleters, glabrous or sometimes fimbriate. Inflorescences terminal, compound-cymose; rachis terete; peduncles 6.0–15.0 cm long; bracts ovate-lanceolate to lanceolate, 0.5–3.0 cm long; bracteoles lanceolate, 1.0– 1.5 mm long. Flowers heterostylous, subsessile, pedicels 0–0.5 mm long. Hypanthium ca. 1.0 mm long, obconic-campanulate, glabrous; sepals 4, ca. 1 mm long, triangular, glabrous. Corolla white or purplish on the margin of corolla lobes, infundibuliform, tube 4.2–5.2 mm long, glabrous abaxially, densely pubescent at throat; lobes 4, ca. 3.0 mm long, ovate. Stigma bilobed, ca. 1.0 mm, clavate. Ovary 2 - celled, ovules numerous on axile placentas. Stamens 4, anthers oblong, ca. 1.0 mm long, oblong-lanceolate. Long-styled flower: stamens included, adnate to the middle part of corolla tube; filaments ca. 1.0 mm long; styles exserted, ca. 6.0 mm long. Short-styled flower: stamens exserted, adnate to throat of corolla tube; filaments ca. 2.0 mm long; styles included, ca. 3.0 mm long. Fruits capsular, ca. 2.0 mm in diameter, subglobose or oblate globose, dehiscent at apex and then septicidally along the ventral suture. Seeds 0.5–0.7 mm long, numerous, angular, testa black, reticulate.</p><p>Phenology.</p><p>Flowering occurs from March to June, and fruiting from July to January of the following year.</p><p>Etymology.</p><p>The specific epithet refers to the fact that the species is growing in the south of the Guangxi Zhuang Autonomous Region. Its Chinese name is “ 桂南耳草 ” (Guì Nán Ěr Cǎo).</p><p>Distribution and habitat.</p><p>Up to now, Hedyotis austroguangxiensis is known from Chongzuo City and Fangchenggang City of the Guangxi Zhuang Autonomous Region; it grows mainly on roadsides or wet rocks under secondary forests at an elevation of 800–1100 m. Main associated species are Selaginella doederleinii Hieron. ( Selaginellaceae) and Blechnopsis orientalis (L.) C. Presl ( Blechnaceae).</p><p>Palynology.</p><p>The pollens of the new species are monads, isopolar and spheroidal, with 3 - colporate apertures, the tectum is double microreticulate, with a psilate suprareticulum and a microechinate infrareticulum. The pollen size is 21.7 (20.2–22.7) × 20.6 (19.5–21.4) μm with P / E value of 1.05 in long-styled flower (Fig. 5 A – C) and 23.3 (21.1–24.3) × 22.5 (21.3–24.1) μm with P / E value of 1.04 in short-styled flower (Fig. 5 D – F).</p><p>Additional specimens examined.</p><p>CHINA • Guangxi Zhuang Autonomous Region: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.48589&amp;materialsCitation.latitude=21.672909" title="Search Plazi for locations around (long 107.48589/lat 21.672909)">Chongzuo City</a>, Ningming County, 28 Oct 2020, 21°40'22.476"N, 107°29'09.218"E, 1066.52 m a. s. l., Ming-Deng Yuan &amp; Yi-Da Xu YS 391 (IBSC) ; • ibid., Aidian Town, Luxu Village, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.48589&amp;materialsCitation.latitude=21.672909" title="Search Plazi for locations around (long 107.48589/lat 21.672909)">Gongmu Mountain</a>, 14 Mar 2010, Yu-De Peng, Ying-Feng Huang, Cui-Hong Lu 21786 (GXMG) ; • ibid., Tongmian Town, Kunan Village, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.5234&amp;materialsCitation.latitude=21.70938" title="Search Plazi for locations around (long 107.5234/lat 21.70938)">Liangling</a>, 21°42'33.76"N, 107°31'24.23"E, 951 m a. s. l., 28 May 2013, Dong-Xin Nong, Hai-Feng Ceng, Wei Liu 451422130528052 LY (GXMG) ; • ibid., 7 Jan 2020, Ming-Deng Yuan et al. YS 99 (IBSC); • ibid., 14 Jun 2020, Ming-Deng Yuan, Guo-Bin Jiang YS 340 (IBSC); • ibid., short-styled flower, 24 Apr 2021, Ming-Deng Yuan &amp; Yi-Da Xu YS 431 (IBSC); • Fangchenggang City, Fangcheng District, under thick forest, on the hill, 23 Mar 2010, Shiwandashan Collection Team 2189 (IBK) .</p><p>Conservation status.</p><p>So far, the subpopulations of Hedyotis austroguangxiensis exceed 4 based on the information from specimens and our field expedition, and more than 600 mature individuals have been found in the field (AOO 16 km 2, EOO 91 km 2). According to the criteria D 1 of the IUCN Red List Categories and Criteria (IUCN 2024), the species can be assessed as “ Vulnerable ”. However, this species is widely distributed and may also be found in neighboring countries like Vietnam and Laos. In addition, it has not found any medicinal and ornamental values yet. We thus recommend evaluating it as Least Concern (LC).</p></div>	https://treatment.plazi.org/id/309E7C6FC7555525A896C31DA8ACA26B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Xu, Yi-Da;Yuan, Ming-Deng;Wang, Rui-Jiang	Xu, Yi-Da, Yuan, Ming-Deng, Wang, Rui-Jiang (2025): Two new species of Hedyotis L. (Rubiaceae) from Yunnan and Guangxi, China. PhytoKeys 263: 135-149, DOI: 10.3897/phytokeys.263.163043
5616C4CDFAFD5A2A956B2163E809C8AD.text	5616C4CDFAFD5A2A956B2163E809C8AD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hedyotis jinghongensis M. D. Yuan & R. J. Wang 2025	<div><p>1. Hedyotis jinghongensis M. D. Yuan &amp; R. J. Wang sp. nov.</p><p>Figs 2, 3</p><p>Type.</p><p>CHINA • From Yunnan Province: Dai Autonomous Prefecture of Xishuangbanna, Jinghong City, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.07254&amp;materialsCitation.latitude=21.980637" title="Search Plazi for locations around (long 101.07254/lat 21.980637)">Jinuo Town</a>, roadside, under thick forest, 21°58'50.29"N, 101°04'21.16"E, ca. 1288 m a. s. l., flowering, short-styled flower, 31 July 2019, Xin-Xin Zhou, Guo-Bin Jiang &amp; Ming-Deng Yuan 1406 (holotype: IBSC [0858378]; isotype IBSC [0858377]) .</p><p>Diagnosis.</p><p>The new species is similar to H. communis W. C. Ko (1974: 578) and H. interrupta G. B. Jiang &amp; R. J. Wang (2021) in having lanceolate to narrowly lanceolate leaves and triangular stipules, but it differs from H. communis by having obvious tetragonal stem, smaller leaves, stipules marginally with 3–6 colleter-tipped linear lobes on each side, and short inflorescences with small flowers (Table 2). It is also distinguished from H. interrupta by having small leaves, stipules marginally with 3–6 colleter-tipped linear lobes on each side, and short inflorescences (Table 2).</p><p>Description.</p><p>Perennial herbs, erect, 15–60 (– 80) cm tall. Stem tetragonal, narrowly winged, branched mostly at the upper part, glabrous. Leaves opposite, petioles 2.0–5.0 mm long, glabrous; blade 2.5–9.0 × 0.7–2.0 cm, lanceolate to narrowly lanceolate, apex acuminate to caudate acuminate, base cuneate, blade papyraceous, light green adaxially and grayish green abaxially, glabrous on both sides; midrib depressed adaxially and prominent abaxially, secondary veins 3–7 on each side, distinct on both sides. Stipules interpetiolar, 5.0– 10 mm long, triangular to narrowly triangular, marginally with 3–6 colleter-tipped linear lobes on each side, glabrous. Inflorescences terminal and axillary on lateral branches, 5–20 flowered, cyme or compound cymose; peduncles 0.5–3.0 cm long, tetragonal, narrowly winged, glabrous; bracts 2.0–12.0 mm long, lanceolate or linear. Flowers heterostylous, pedicels 0–2.0 mm long. Hypanthium ca. 1.0 mm long, campanulate, glabrous; sepals 4, 1.0– 1.2 mm long, narrowly triangular, glabrous. Corolla tubular, white; tube 1.7–2.0 mm long, densely pubescent at throat adaxially, glabrous abaxially; lobes 4, 1.8–2.0 mm long, oblong-lanceolate, glabrous abaxially, and pubescence adaxially. Stigma bilobed, ca. 0.5 mm long, clavate. Ovary 2 - celled, ovules numerous on axile placentas. Stamens 4, anthers ca. 0.8 mm long, oblong-linear. Long-styled flowers: corolla tube ca. 2.0 mm long; stamens included, adnate to the middle of corolla tube; filaments ca. 0.8 mm long; styles ca. 3.3 mm long, stigma exserted. Short-styled flowers: corolla tube ca. 1.7 mm long; stamens included, adnate to throat of corolla tube; filaments ca. 1.2 mm long, anthers exserted; styles ca. 2.8 mm long, stigma included or slightly exserted. Fruits capsular, 1.5–2.0 × 2.0– 2.5 mm, subglobose, sepals persistent, glabrous, dehiscent at apex and then septicidally along the ventral suture. Seeds ca. 0.5 mm long, numerous, angular, 0.4–0.6 mm long, testa black, reticulated.</p><p>Phenology.</p><p>Flowering and fruiting individuals were observed between June and December.</p><p>Etymology.</p><p>The specific epithet refers to the type locality of this new species, Jinghong City, Dai Autonomous Prefecture of Xishuangbanna, Yunnan Province. Its Chinese name is “ 景洪耳草 ” (Jǐng Hóng Ěr Cǎo).</p><p>Distribution and habitat.</p><p>Currently, Hedyotis jinghongensis is only known from southern Yunnan Province, China. It grows on roadsides under secondary forests at an elevation of 700–1400 m. The main associated species are Selaginella leptophylla Baker ( Selaginellaceae), Phyllanthus urinaria Linnaeus ( Phyllanthaceae), and Sonerila cantonensis Stapf ( Melastomataceae).</p><p>Palynology.</p><p>The pollens of the new species are monads, isopolar, with 3 - or 4 - colporate apertures, the tectum is double microreticulate, with a psilate suprareticulum and a microechinate infrareticulum. The pollen size is 25.9 (23.6–28.0) × 24.8 (23.4–26.3) μm with P / E value 1.04, and the shape is spheroidal in long-styled flowers (Fig. 3 A – C); the pollen size is 20.0 (17.5–20.2) × 16.9 (16.4–17.1) μm with P / E value 1.18, and the shape is subprolate in short-styled flowers (Fig. 3 D – F).</p><p>Additional specimens examined.</p><p>CHINA • Yunnan: Dai Autonomous Prefecture of Xishuangbanna, Cheli County, Zhalei, under thickets, 1400 m a. s. l., Oct 1936, Qi-Wu, Wang 79170 (KUN) ; • Jinghong City, Jinuo Town, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.01667&amp;materialsCitation.latitude=22.033333" title="Search Plazi for locations around (long 101.01667/lat 22.033333)">Longpa Village</a>, under forest, 22°2'N, 101°1'E, 1250 m a. s. l., 02 Aug 1977, Guo-Da Tao 16483 (HITBC, IBSC) ; • ibid., 04 Aug 1977, Guo-Da Tao 17845 (HITBC); • ibid., 21°59'N, 101°40'E, 1250 m a. s. l., 29 Jul 1977, Guo-Da Tao 17850 (HITBC, KUN); • ibid., 30 Nov. 2018, Xin-Xin Zhou 5469 (IBSC); • ibid., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.88333&amp;materialsCitation.latitude=22.883333" title="Search Plazi for locations around (long 100.88333/lat 22.883333)">Mengyang Town</a>, near river, 22°53'N, 100°53'E, 750 m a. s. l., Guo-Da Tao 16799 (HITBC) ; • ibid., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.86667&amp;materialsCitation.latitude=22.183332" title="Search Plazi for locations around (long 100.86667/lat 22.183332)">Guanping Village</a>, roadside, slopes, 22°11'N, 100°52'E, 800 m a. s. l., 17 Aug 1977, Guo-Da Tao 17577 (KUN, IBSC) ; • ibid., 10 Nov 2019, Ming-Deng Yuan &amp; Jiang-Ping Shu YS 26 (IBSC); • ibid., Puwen Town, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.03333&amp;materialsCitation.latitude=22.583334" title="Search Plazi for locations around (long 101.03333/lat 22.583334)">Dakaihe Village</a>, under forest, 22°35'N, 101°2'E, 14 Aug 1977, Guo-Da Tao 16898 (HITBC, KUN) ; • ibid., Mengla County, Mengman Town, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.683334&amp;materialsCitation.latitude=21.616667" title="Search Plazi for locations around (long 101.683334/lat 21.616667)">Jingpiao Village</a>, 21°37'N, 101°41'E, Investigation Team 25230 (HITBC) ; • ibid., Yaoqu Town, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.6&amp;materialsCitation.latitude=21.666666" title="Search Plazi for locations around (long 101.6/lat 21.666666)">Nazuo Village</a>, in miscellaneous wood forest, 21°40'N, 101°36'E, 700–800 m a. s. l., 23 Nov 1974, Zeng-Hong Yang 012484 (HITBC, KUN) ; • ibid., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.99746&amp;materialsCitation.latitude=23.40204" title="Search Plazi for locations around (long 100.99746/lat 23.40204)">Puer City</a>, Jinggu County, 23°24'7.35"N, 100°59'50.87"E, 1224 m a. s. l., 12 Sept 2020, Yan-Xiong Gong G 38 (IBSC) ; • ibid., Puer County, Mohei Town, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.166664&amp;materialsCitation.latitude=23.266666" title="Search Plazi for locations around (long 101.166664/lat 23.266666)">Choushui Village</a>, 23°16'N, 101°10'E, 1300 m a. s. l., 27 Aug 1990, Guo-Da Tao 46403 (HITBC) ; • ibid., 09 Nov 2019, Ming-Deng Yuan &amp; Jiang-Ping Shu YS 5 (IBSC); • ibid., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.95&amp;materialsCitation.latitude=23.35" title="Search Plazi for locations around (long 100.95/lat 23.35)">the road along Puer County to Jinggu County</a>, in the dank of the valley, 23°21'N, 100°57'E, 950 m a. s. l., Hong Wang 4805 (HITBC) .</p><p>Conservation status.</p><p>So far, four subpopulations with more than 500 mature individuals have been found in the field (AOO 40 km 2, EOO 8500 km 2). According to the criteria D 1 of the IUCN Red List Categories and Criteria (IUCN 2024), the species can be assessed as “ Vulnerable ”. However, many other subpopulations of this species may be found nearby in southern Yunnan or northern Laos. Considering that this species has no economic uses, we recommend evaluating it as Least Concern (LC).</p></div>	https://treatment.plazi.org/id/5616C4CDFAFD5A2A956B2163E809C8AD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Xu, Yi-Da;Yuan, Ming-Deng;Wang, Rui-Jiang	Xu, Yi-Da, Yuan, Ming-Deng, Wang, Rui-Jiang (2025): Two new species of Hedyotis L. (Rubiaceae) from Yunnan and Guangxi, China. PhytoKeys 263: 135-149, DOI: 10.3897/phytokeys.263.163043
