taxonID	type	description	language	source
3D6F384EFFD6FF9685E6A0C2FDBAFBBD.taxon	description	(Figs 1, 3, 5, 20)	en	Romantsov, Pavel V., Bezděk, Jan (2025): Two new species of Clytrini from South-east Asia (Coleoptera: Chrysomelidae: Cryptocephalinae). Zootaxa 5689 (2): 379-391, DOI: 10.11646/zootaxa.5689.2.8, URL: https://doi.org/10.11646/zootaxa.5689.2.8
3D6F384EFFD6FF9685E6A0C2FDBAFBBD.taxon	materials_examined	Type locality. Thailand, Chiang Mai Prov., 100 km N from Chaing Mai, Chiang Dao Hill Resort, 19 ° 33 ' 28.9 " N, 99 ° 04 ' 33.3 " E. Type specimens. Holotype: ♂ (ZIN), “ Thailand, Chiang Mai Prov., / 100 km N from Chaing Mai, Chiang / Dao Hill Resort, h = 494 m, / 19 ° 33 ' 28.9 " N, 99 ° 04 ' 33.3 " E / 10. VI. 2017, A. S. Prosvirov leg. [w, p] ”. Paratypes: 1 ♀ (PRCS), same data as in holotype, but “ 17. VI. 2017 ”; 1 ♀ (PRCS), “ Thailand, Chiang Mai Prov. 100 / km N from Chaing Mai, Chiang / Dao Hill Resort evn. h = 550 m, at light / 19 ° 33 ' 30.31 " N, 99 ° 4 ' 38.69 " E / 10. VI. 2017, A. S. Prosvirov [w, p] ”; 1 ♂ 1 ♀ (♂ PRCS, ♀ JBCB), “ Thailand, Chiang Mai Prov. 100 / km N from Chaing Mai, Chiang / Dao Hill Resort. 19 ° 33 ' 30.31 " N, / 99 ° 4 ' 38.69 " E, h = 550 m., at light / 17. VI. 2017, A. S. Prosvirov [w, p] ”; 1 ♀ (ZIN), “ Laos / Vientiane [w, p] // Coll. R. I. Sc. N. B. / Laos [y, p] ”. The type specimens are provided with one additional printed red label: “ Holotypus [or Paratypus] / Aetheomorpha prosvirovi / Romantsov & Bezděk, sp. n. / det Romantsov & Bezděk, 2025 ”.	en	Romantsov, Pavel V., Bezděk, Jan (2025): Two new species of Clytrini from South-east Asia (Coleoptera: Chrysomelidae: Cryptocephalinae). Zootaxa 5689 (2): 379-391, DOI: 10.11646/zootaxa.5689.2.8, URL: https://doi.org/10.11646/zootaxa.5689.2.8
3D6F384EFFD6FF9685E6A0C2FDBAFBBD.taxon	description	Description. Measurements. Males: 4.2 – 4.9 mm (holotype 4.9 mm), females: 4.3 – 4.9 mm. Body elongate. Head brown with area along inner edges of eyes connecting to vertex black, palpi light brown, apices of mandibles black. Pronotum light brown with four black spots in transverse row at basal third. Scutellum black with brownish small spot on apex. Elytra light brown, each elytron with black pattern consisting of five black spots (humeral spot, spot at basal part between scutellum and humerus obliquely connected with humeral one, spot in middle near lateral margin, spot at two thirds near suture, and last on apex) and black suture, lateral and apical margins; variable lateral black stripe connecting with humeral spot as well as with third and apical spots. Underside brown, prosternum with black elongate spot on each side, mesosternum and metasternum in middle and partly laterally black, abdomen including pygidium brown. Antennae brown, antennomeres V – VIII brownish ventrally and laterally, dorsally darkened, IX – XI black. Legs brown with femora (except bases) and tarsi black. Male (holotype, Fig. 1). Head large, impunctate, 1.42 times narrower than pronotum. Frontoclypeus with slightly rugose clypeal part, anterior margin with wide incision in middle and with triangular anterior angles. Frons very slightly convex with small groove in middle, surface near eyes covered by fine microsculpture and adpressed silver setae. Frontal tubercles absent. Eyes rather large, slightly bean-shaped with small notch on inner margin, 1.4 times as long as wide. Interocular space twice as wide as transverse diameter of eye and 1.7 times narrower than head with eyes. Vertex slightly convex, lustrous and impunctate. Antennae short and robust, about 3.8 times shorter than body, just extending to posterior third of pronotum. Antennomere I large, stout; II short, almost round; III shortest; antennomere IV inverse trapezoid; V – X strongly serrate, XI elongate with triangular apex. Antennomeres I – IV lustrous with sparse, short, erected setae on edges, V – XI dull with slightly rugose surface covered with short adpressed setae and with longer erected setae on edges. Length ratio of antennomeres I – XI as 9: 4: 3: 4: 5: 4.5: 4.5: 4: 4: 4.5: 7, width ratio as 6.5: 4.5: 4: 5: 8: 8: 7: 6: 5.5: 4: 3. Pronotum convex, transverse, 2.13 as wide as long, about as wide as elytra on base, widest in middle. Anterior margin slightly convex in middle, lateral margins evenly rounded, posterior margin almost straight near lateral angles, in middle of pronotal base distinctly protruded opposite scutellum. Surface lustrous, sparsely covered with very small punctures (visible only at 20 x magnification) and with denser microscopic punctures in interstices between them. Anterior angles rectangular with rounded tip, posterior ones almost imperceptible, widely rounded. Anterior angles bearing setigerous pore with rather long, curved pale seta. Anterior margin bordered only near angles then unbordered; posterior margin thinly bordered and lateral margins widely bordered. Scutellum triangular with sharply rounded apex, about 1.7 times as wide as long; surface covered with fine punctures, scutellar apex very weakly elevated upon level of elytra. Elytra subcylindrical, 1.27 times as long as wide. Basal margin bordered only near shoulders, lateral margin widely bordered in anterior two third and thinly bordered in posterior third. Elytral surface covered with confused punctures in basal part but arranged in more or less distinct rows in middle and lateral parts. Punctures at basal third rather large, then gradually diminishing and disappearing in apical slope. Epipleura glabrous, basally moderately wide, then gradually narrowed, disappearing in apical third of elytra. In lateral view, lateral margin of elytra distinctly lobed below shoulders. Pygidium slightly exposed with rounded apex. Legs robust. All tibiae almost straight, noticeably widened towards apex. Tarsi short. Protarsomere I enlarged, slightly wider than III. Length ratio of protarsomeres I – III and V as 10: 8: 10: 15; width ratio of protarsomeres I – III as 10: 8: 9. Mesotarsomere I almost as wide as III; metatarsomere I narrower than III. Claws simple. Aedeagus (Fig. 3) rather short and wide with triangular apex. Underside with wide and low ridge medially and ridges at lateral margins, area between ridges rather deeply depressed. In lateral view apex strongly bent ventrally. Length of aedeagus 1.25 mm, width 0.4 mm. Females. Tarsi are narrower than in males. Pygidium with rounded apex, without any notches (Fig. 5). Spermatheca U-shaped with sharp apex (Fig. 20). Variability. One male and two females have tibiae completely black. Female from Laos has posterior part of head with more developed black pattern (frons and vertex black with transverse brown spot in middle), pygidium with spot near apex black and blurred blackish strokes on each sides of abdominal ventrites.	en	Romantsov, Pavel V., Bezděk, Jan (2025): Two new species of Clytrini from South-east Asia (Coleoptera: Chrysomelidae: Cryptocephalinae). Zootaxa 5689 (2): 379-391, DOI: 10.11646/zootaxa.5689.2.8, URL: https://doi.org/10.11646/zootaxa.5689.2.8
3D6F384EFFD6FF9685E6A0C2FDBAFBBD.taxon	diagnosis	Differential diagnosis. This new species belongs to a group of species from Indochina and India characterized by a pronotum with four black spots and elytra with black suture, lateral margins, and four black spots. This group includes Aetheomorpha weigeli Medvedev, 2010, A. bilobata Wang & Zhou, 2012 from China (Yunnan), A. quatuordecimpunctata (Jacoby, 1895) from South India, and possibly also A. decemnotata (Jacoby, 1892), which is distributed across China (Yunnan), Myanmar, Vietnam, Laos and Thailand. With its large head and triangular apex of the aedeagus, Aetheomorpha prosvirovi sp. nov. is most similar to A. weigeli. These two species can be distinguished as follows. Aetheomorpha prosvirovi sp. nov. (Fig. 1): body smaller (4.2 – 4.9 mm); black pattern on head more developed; spots on pronotum larger, intermediate ones not connected; third spot on elytra usually connecting with lateral black edging; middle antennomeres more transverse (antennomeres V – VIII 1.5 – 1.8 times as wide as long); underside of aedeagus with a low, wide longitudinal ridge, in lateral view apex strongly bent ventrally; female pygidium not incised at the apex. Distribution: Norhern Thailand (Chiang Mai), Laos (Vientiane). Aetheomorpha weigeli (Fig. 2): body larger (5.1 – 5.9 mm); black pattern on head less developed; spots on pronotum smaller, intermediate two sometimes connected, forming a large oblong spot; third spot on elytra not connected to the lateral black edge; middle antennomeres less transverse (antennomeres V – VIII approximately as wide as long); underside of aedeagus with a high, sharp ridge medially, in lateral view apex slightly bent ventrally (Fig. 4); female pygidium slightly incised at the apex (Fig. 6); spermathecal with basal and apical parts almost parallel (Fig. 21). Distribution: China (Yunnan), Northern Thailand (Mae Hong Son).	en	Romantsov, Pavel V., Bezděk, Jan (2025): Two new species of Clytrini from South-east Asia (Coleoptera: Chrysomelidae: Cryptocephalinae). Zootaxa 5689 (2): 379-391, DOI: 10.11646/zootaxa.5689.2.8, URL: https://doi.org/10.11646/zootaxa.5689.2.8
3D6F384EFFD6FF9685E6A0C2FDBAFBBD.taxon	description	Aetheomorpha bilobata can be easily distinguished from other species in this group by its smaller head and slightly elongated, broad, and feebly bilobed apex of aedeagus. Some specimens of A. decemnotata have four obscure spots on the pronotum, but this species differs in shape of aedeagus with a narrower apex and an underside without ridge. Aetheomorpha quatuordecimpunctata (Jacoby, 1895) from South India can be distinguished by its strongly transverse middle antennomeres, completely black head, and elytra lacking black lateral margins and a black suture. Its aedeagus has a narrower triangular apex, and the underside is evenly convex without any depression or ridge. In lateral view, the apex of the aedeagus is only very slightly bent ventrally.	en	Romantsov, Pavel V., Bezděk, Jan (2025): Two new species of Clytrini from South-east Asia (Coleoptera: Chrysomelidae: Cryptocephalinae). Zootaxa 5689 (2): 379-391, DOI: 10.11646/zootaxa.5689.2.8, URL: https://doi.org/10.11646/zootaxa.5689.2.8
3D6F384EFFD6FF9685E6A0C2FDBAFBBD.taxon	etymology	Etymology. The new species is named in honor of A. S. Prosvirov, a specialist in Elateridae, who collected the majority of the type series.	en	Romantsov, Pavel V., Bezděk, Jan (2025): Two new species of Clytrini from South-east Asia (Coleoptera: Chrysomelidae: Cryptocephalinae). Zootaxa 5689 (2): 379-391, DOI: 10.11646/zootaxa.5689.2.8, URL: https://doi.org/10.11646/zootaxa.5689.2.8
3D6F384EFFD6FF9685E6A0C2FDBAFBBD.taxon	distribution	Distribution. Thailand, Laos.	en	Romantsov, Pavel V., Bezděk, Jan (2025): Two new species of Clytrini from South-east Asia (Coleoptera: Chrysomelidae: Cryptocephalinae). Zootaxa 5689 (2): 379-391, DOI: 10.11646/zootaxa.5689.2.8, URL: https://doi.org/10.11646/zootaxa.5689.2.8
3D6F384EFFD3FF9685E6A2B7FCA0F84F.taxon	description	(Figs 2, 4, 6, 21)	en	Romantsov, Pavel V., Bezděk, Jan (2025): Two new species of Clytrini from South-east Asia (Coleoptera: Chrysomelidae: Cryptocephalinae). Zootaxa 5689 (2): 379-391, DOI: 10.11646/zootaxa.5689.2.8, URL: https://doi.org/10.11646/zootaxa.5689.2.8
3D6F384EFFD3FF9685E6A2B7FCA0F84F.taxon	materials_examined	Type locality. “ China, S. Yunnan, Prov. Xishuangbanna, 23 km NW Jinghong, way Na Ban to Man Fei ”. Type specimens examined. Holotype (Fig. 2): ♂ (NMEG), “ CHINA: S-YUNNAN / (Xishuangbanna) / 23 km NW Jinghong / way Na Ban to Man Fei / (NNNR), 06. vi. 2008 LF / leg. A. Weigel 630 m [w, p] // HOLOTYPUS [p] / Aetheomorpha / weigeli [h] / L. Medvedev [r, p] ”; Paratype: ♀ (ZIN), “ CHINA, S. Yunnan / Prov. Xishuangbanna 23 km NW Junghong / vic. Na Ban (NNNR) [w, p] // 730 m NN 22 ° 09.49 ' N / 100 ° 39.92 ' E, 06. VI. 2008 / leg. A. Weigel [w, p] // PARATYPUS [p] / Aetheomorpha / weigeli [h] / L. Medvedev [r, p] // Zoological Institute / St. Petersburg / INS _ COL _ 0005251 [w, p] ”. Additional material examined. 17 specimens — CHINA: Yunnan, Xishuangbanna, 23 km NW Junghong, Na Ban village, 22 ° 10 ' 04 " N 100 ° 39 ' 52 " E, 680 m, 18. v. 2008, 1 ♀, A. Weigel leg. (ZIN). THAILAND: Mae Hong Son Prov., Kiwlom pass near Soppong, 19 ° 26 ' N 98 ° 19 ' E, 1400 m, 23. vi. - 2. vii. 2002, 1 ♂ 11 ♀♀, R. & H. Fouqué leg. (JBCB); Doi Phu Ka Nat. Park, 50 km NNE of Nan, 28. vi. - 1. vii. 1997, 1 ♂, J. Kaláb leg. (JBCB); Chiang Mai Prov., Kaeng Pan Tao, 19 ° 16.825 ' N 98 ° 58.017 ' E, 418 m, 22. vii. 2006, 1 ♀, M. V. L. Barclay & H. Mendel leg. (NHMUK). MYANMAR: Shan State, Pindaya env., 20.92033 ° N 96.64942 ° E, 1330 m, 15. - 16. vi. 2015, 1 ♀, J. Žák leg. (JBCB), same data, 1 ♀, W. Grosser leg. (JBCB).	en	Romantsov, Pavel V., Bezděk, Jan (2025): Two new species of Clytrini from South-east Asia (Coleoptera: Chrysomelidae: Cryptocephalinae). Zootaxa 5689 (2): 379-391, DOI: 10.11646/zootaxa.5689.2.8, URL: https://doi.org/10.11646/zootaxa.5689.2.8
3D6F384EFFD3FF9685E6A2B7FCA0F84F.taxon	distribution	Distribution. China (Yunnan), Myanmar, Thailand.	en	Romantsov, Pavel V., Bezděk, Jan (2025): Two new species of Clytrini from South-east Asia (Coleoptera: Chrysomelidae: Cryptocephalinae). Zootaxa 5689 (2): 379-391, DOI: 10.11646/zootaxa.5689.2.8, URL: https://doi.org/10.11646/zootaxa.5689.2.8
3D6F384EFFD1FF9A85E6A786FE61FB99.taxon	description	(Figs 7, 16, 22)	en	Romantsov, Pavel V., Bezděk, Jan (2025): Two new species of Clytrini from South-east Asia (Coleoptera: Chrysomelidae: Cryptocephalinae). Zootaxa 5689 (2): 379-391, DOI: 10.11646/zootaxa.5689.2.8, URL: https://doi.org/10.11646/zootaxa.5689.2.8
3D6F384EFFD1FF9A85E6A786FE61FB99.taxon	materials_examined	Type locality. Vietnam, Hòa Bình Prov., Mai Chau Vill., Eco Homestay, 20 ° 38 ' 56.6 " N, 105 ° 03 ' 59.2 " E. Type specimens. Holotype (Fig. 7): ♂ (PRCS), “ N Vietnam, Hòa Bình Prov., Mai / Chau Vill., Eco Homestay / N 20 ° 38 ' 56.6 ", E 105 ° 3 ' 59.2 " / 170 m, at light, / 20. IV. 2019 P. Romantsov leg. [w, p] ”. Paratype: 1 ♀ (ZIN), same data as in holotype. The type specimens are provided with one additional printed red label: “ Holotypus [or Paratypus] / Smaragdina bienkowskii / Romantsov & Bezděk, sp. n. / det Romantsov & Bezděk, 2025 ”.	en	Romantsov, Pavel V., Bezděk, Jan (2025): Two new species of Clytrini from South-east Asia (Coleoptera: Chrysomelidae: Cryptocephalinae). Zootaxa 5689 (2): 379-391, DOI: 10.11646/zootaxa.5689.2.8, URL: https://doi.org/10.11646/zootaxa.5689.2.8
3D6F384EFFD1FF9A85E6A786FE61FB99.taxon	description	Description. Measurements. Male: 6.3 mm (holotype), female: 6.5 mm. Body elongate. Head brown. Pronotum brown, but slightly lighter than head. Scutellum brown. Elytra light brown. Underside and legs brown. Antennae brown, antennomere IV with inner dorsal half darkened, V – VI brownish ventrally and laterally, dorsally black, VII – XI black. Male (holotype, Fig. 7). Head large, impunctate, 1.38 times narrower than pronotum. Labrum with slightly rounded apical margin, surface slightly convex, lustrous, impunctate with several setae at anterior third. Frontoclypeus with lustrous and slightly convex clypeal part, anterior margin slightly concave. Frons slightly convex with depression between eyes, surface near eyes covered by very fine microsculpture without setae. Frontal tubercles absent. Eyes large, slightly bean-shaped with small notch on inner margin, 1.33 times as long as wide. Interocular space narrow, 0.77 times as wide as transverse diameter of eye and about 3.3 times narrower than head with eyes. Vertex slightly convex, lustrous and impunctate. Antennae moderately short, about 2.7 times shorter than body. Antennomere I large, stout, II pentagonal, III shortest, slightly widened at apex, IV – X strongly flattened and serrate, last antennomere spear-shaped with elongate sharp apex. Antennomeres I – III lustrous with sparse, short, erected setae on inner edge, IV – XI dull with slightly rugose surface covered with short adpressed setae and with two or three longer erected setae on anterior angles. Length ratio of antennomeres I – XI as 11: 5: 4: 9: 9: 9: 9: 9: 8.5: 8: 13, width ratio as 6: 5: 3.5: 8: 9: 9: 8: 7: 8: 7: 6.5. Penultimate maxillary palpomere slightly wider and about 1.4 times longer than next one, apical palpomere with truncated apex. Pronotum convex, transverse, 1.75 as wide as long, about as wide as elytra on base, widest at anterior third. Anterior margin slightly convex in middle, lateral margins rounded, posterior margin almost straight near lateral angles, in middle of pronotal base distinctly protruded opposite scutellum. Surface lustrous, very sparsely covered with small punctures. Anterior angles obtusangulate with rounded tip, posterior ones almost imperceptible, widely rounded. Anterior angles bearing setigerous pore with rather long, curved pale seta. Anterior margin bordered near angles, in middle part unbordered; posterior margin thinly bordered and lateral margins widely bordered. Scutellum triangular, approximately as wide as long, surface lustrous and impunctate, scutellar apex very weakly elevated upon level of elytra. Elytra subcylindrical, 1.57 times as long as wide. Basal margin bordered only near shoulders, lateral margins widely bordered in anterior half and thinly bordered in posterior half. Elytral surface covered with confused punctures in basal part but arranged in more or less distinct rows in middle and lateral parts. Punctures at basal third rather large, then gradually diminishing and disappearing in apical slope. Epipleura glabrous, moderately wide in anterior third, then rather suddenly narrow and disappearing at level of metacoxae. In lateral view, lateral margin of elytra moderately lobed below shoulders. Pygidium with rounded apex, not exposed. Legs robust. All tibiae almost straight, noticeably widened towards apex. Tarsomeres I of all legs slightly narrower than III. Length ratio of protarsomeres I – III and V as 15: 12: 14: 22; width ratio of protarsomeres I – III as 12: 11: 14. Claws simple but slightly thickened at base. Aedeagus (Fig. 16) rather short and wide with finger-shaped emarginated apex. Underside of aedeagus slightly convex without distinct depressions or ridges. In lateral view apex strongly bent ventrally. Length of aedeagus 1.7 mm, width 0.5 mm. Female. Antennae less serrated. Eyes smaller and less convex than in male, interocular space somewhat wider (1.05 times as wide as transverse diameter of eye and about 2.57 times narrower than head with eyes). Pygidium of female with slightly truncated apex, so its posterior margin looks almost straight with very faint, almost imperceptible notch in middle. Spermatheca with long frizzy ductus ending by rather large spindle (Fig. 22). Length of spermatheca (excluding ductus) 0.4 mm. Variability. Coloration of female similar as in holotype but metepisterna slightly darkened.	en	Romantsov, Pavel V., Bezděk, Jan (2025): Two new species of Clytrini from South-east Asia (Coleoptera: Chrysomelidae: Cryptocephalinae). Zootaxa 5689 (2): 379-391, DOI: 10.11646/zootaxa.5689.2.8, URL: https://doi.org/10.11646/zootaxa.5689.2.8
3D6F384EFFD1FF9A85E6A786FE61FB99.taxon	diagnosis	Differential diagnosis. Smaragdina bienkowskii sp. nov. belongs to the S. sikkimia species group (which also includes S. constrictifrons, S. sikkimia and S. vietnamica). This group is characterized by a yellowish-brown to brown dorsum, large eyes, and a narrow frons, noticeably narrower than the transverse diameter of the eye in males and approximately equal in females, as well as a finger-shaped apical process of the aedeagus. Smaragdina sikkimia has an aedeagus very similar to that of S. bienkowskii sp. nov., but it differs in having a black or darkened ventral side and tarsi. Additionally, males of S. sikkimia have a narrower frons (0.60 - 0.65 as wide as transverse diameter of eye, compared to 0.77 in S. bienkowskii sp. nov.). The aedeagus of S. sikkimia is depressed below the gonopore, therefore, in lateral view, it appears thin and sinuous in its preapical part (Fig. 18). Females of S. sikkimia differ in the shape of spermatheca (Fig. 23). In the identification keys for Oriental Smaragdina by Medvedev (2010) and Warchałowski (2012), S. bienkowskii sp. nov. would be placed near S. constrictifrons. However, S. constrictifrons differs from S. bienkowskii sp. nov. by having an apical process of the aedeagus that is toothed on each side and not emarginated on the apex. All species in the S. sikkimia species group can be identified using the key provided below. Smaragdina diversiceps (Pic, 1941) and S. fulvitarsis Medvedev, 1992 belong to another large Smaragdina species complex, characterized by a with wide frons (significantly wider than transverse diameter of the eye, even in males, and especially in female), but they have somewhat similar shapes of aedeagus (see the drawings in Medvedev 2010 a). However, both these species are smaller in size (S. bienkowskii sp. nov.: 6.3 – 6.5 mm, S. diversiceps: 4.1 – 6.1 mm, S. fulvitarsis: 4.0 – 4.7) and also differ from S. bienkowskii sp. nov. in couloration: S. diversiceps has a black head, underside and humeral spot on elytra, and S. fulvitarsis has a darkened metasternum. Due to their narrow frons, Smaragdina oculata Medvedev, 1988 and S. megalayana Medvedev & Kantner, 2002, are also externally similar to S. bienkowskii sp. nov. However, they differ in the shape of the aedeagus, which has a triangular apex with a pointed tip. Smaragdina tamdaoana Medvedev, 2010, known only from females and also having brown upperside, is easily distinguished from S. bienkowskii sp. nov. by its very wide frons, which is more than twice as wide as the transverse diameter of the eye.	en	Romantsov, Pavel V., Bezděk, Jan (2025): Two new species of Clytrini from South-east Asia (Coleoptera: Chrysomelidae: Cryptocephalinae). Zootaxa 5689 (2): 379-391, DOI: 10.11646/zootaxa.5689.2.8, URL: https://doi.org/10.11646/zootaxa.5689.2.8
3D6F384EFFD1FF9A85E6A786FE61FB99.taxon	etymology	Etymology. The new species is dedicated to Andrzej O. Bieńkowski, well known specialist in Chrysomelidae.	en	Romantsov, Pavel V., Bezděk, Jan (2025): Two new species of Clytrini from South-east Asia (Coleoptera: Chrysomelidae: Cryptocephalinae). Zootaxa 5689 (2): 379-391, DOI: 10.11646/zootaxa.5689.2.8, URL: https://doi.org/10.11646/zootaxa.5689.2.8
3D6F384EFFD1FF9A85E6A786FE61FB99.taxon	distribution	Distribution. Vietnam.	en	Romantsov, Pavel V., Bezděk, Jan (2025): Two new species of Clytrini from South-east Asia (Coleoptera: Chrysomelidae: Cryptocephalinae). Zootaxa 5689 (2): 379-391, DOI: 10.11646/zootaxa.5689.2.8, URL: https://doi.org/10.11646/zootaxa.5689.2.8
3D6F384EFFD1FF9485E6A6A6FD8AFE6D.taxon	diagnosis	Diagnosis. Dorsum yellowish brown to brown; eyes large, frons narrow, noticeably narrower than the transverse diameter of the eye in males and approximately equal in females; aedeagus with finger-shaped apical process. Species included. Smaragdina bienkowskii sp. nov., S. constrictifrons Medvedev, 2010; S. sikkimia (Jacoby, 1903) and S. vietnamica Medvedev, 2014.	en	Romantsov, Pavel V., Bezděk, Jan (2025): Two new species of Clytrini from South-east Asia (Coleoptera: Chrysomelidae: Cryptocephalinae). Zootaxa 5689 (2): 379-391, DOI: 10.11646/zootaxa.5689.2.8, URL: https://doi.org/10.11646/zootaxa.5689.2.8
3D6F384EFFDFFF9A85E6A2D3FE7EF9E7.taxon	description	(Figs 8, 17)	en	Romantsov, Pavel V., Bezděk, Jan (2025): Two new species of Clytrini from South-east Asia (Coleoptera: Chrysomelidae: Cryptocephalinae). Zootaxa 5689 (2): 379-391, DOI: 10.11646/zootaxa.5689.2.8, URL: https://doi.org/10.11646/zootaxa.5689.2.8
3D6F384EFFDFFF9A85E6A2D3FE7EF9E7.taxon	materials_examined	Type locality. “ NW Thailand, Mae Hong Son, Huai Sua Tao ”. Type specimens examined. Holotype: ♂ (ZIN), “ NW THAI 11 - 17. V. 1992 / MAE HONG SON / Huai Sua Tao / L. Dembicky leg. [w, p] // HOLOTYPUS [p] / Smaragdina / constrictifrons [h] / L. Medvedev det. 20 [p] 10 [r, h] ”. Paratypes: 1 ♂ 1 ♀ (ZIN), “ NW THAI 11 - 17. V. 1992 / MAE HONG SON / Huai Sua Tao / L. Dembicky leg. [w, p] // PARATYPUS [p] / Smaragdina / constrictifrons [h] / L. Medvedev det. 20 [p] 10 [r, h] ” [r].	en	Romantsov, Pavel V., Bezděk, Jan (2025): Two new species of Clytrini from South-east Asia (Coleoptera: Chrysomelidae: Cryptocephalinae). Zootaxa 5689 (2): 379-391, DOI: 10.11646/zootaxa.5689.2.8, URL: https://doi.org/10.11646/zootaxa.5689.2.8
3D6F384EFFDFFF9A85E6A2D3FE7EF9E7.taxon	distribution	Distribution. Thailand.	en	Romantsov, Pavel V., Bezděk, Jan (2025): Two new species of Clytrini from South-east Asia (Coleoptera: Chrysomelidae: Cryptocephalinae). Zootaxa 5689 (2): 379-391, DOI: 10.11646/zootaxa.5689.2.8, URL: https://doi.org/10.11646/zootaxa.5689.2.8
3D6F384EFFDFFF9B85E6A000FF4AFDD9.taxon	description	(Figs 9 – 12, 18, 23)	en	Romantsov, Pavel V., Bezděk, Jan (2025): Two new species of Clytrini from South-east Asia (Coleoptera: Chrysomelidae: Cryptocephalinae). Zootaxa 5689 (2): 379-391, DOI: 10.11646/zootaxa.5689.2.8, URL: https://doi.org/10.11646/zootaxa.5689.2.8
3D6F384EFFDFFF9B85E6A000FF4AFDD9.taxon	materials_examined	Type locality. “ Sikkim ”. Type specimens examined. Syntypes: 1 ♀ (NHMUK), “ Type / H. T. [w, p, round label with red collar] // Sikkim [w, p] // Jacoby Coll. / 1909 - 28 a. [w, p] // SYN- / TYPE [w, p, round label with blue collar] // Gynandrophth. / sikkima Jac. [b, h] // NHMUK 014436765 [w, p, with QR code] ”; 1 ♀ (NHMUK), “ Sikkim [w, p] // Jacoby Coll. / 1909 - 28 a. [w, p] // SYN- / TYPE [w, p, round label with blue collar] // NHMUK 014436766 [w, p, with QR code] ”. Additional material examined. 44 specimens — INDIA: Sikkim, Rangeli River, 900 m 15. iv. 1977, 1 ♂, B. Bhakta leg. (ZIN); Sikkim, Mungphu, 1 ♀ (NHMUK); Sikkim, Gopaldhara, Rungbong valley, 26 ♀♀, H. Stevens leg. (NHMUK). West Bengal Prov., Darjeeling Distr., Lodhama-Jhepl, 1100 - 1350 m, 21. v. 1975, 1 ♂, W. Wittmer leg. (ZIN); West Bengal Prov., Darjeeling Distr., Mirik, 900 m, 24. iv. 1979, 2 ♂♂, B. Bhakta leg. (ZIN); West Bengal Prov., Darjeeling Distr., Tashi Dhing, 27. iv. 1979, 1 ♂, B. Bhakta leg. (ZIN); West Bengal Prov., Darjeeling Distr., Kalimpong env., 25. iv. 1979, 1 ♂, B. Bhakta leg. (ZIN); West Bengal Prov., Darjeeling Distr., Kalimpong, Gangae Thora, 1000 m, 25. iv. 1979, 1 ♀, B. Bhakta leg. (ZIN). NEPAL: Gandaki Distr., Gorkha, 800 - 1100 m, 23. - 28. v. 2001, 1 ♂ 1 ♀, P. Kresl leg. (NMPC); same data but 600 - 1500 m, 29. v. - 3. vi. 1999, 7 ♀♀ (NMPC); same data but 1150 - 1500 m, 21. - 29. v. 2001, 1 ♀ (NMPC).	en	Romantsov, Pavel V., Bezděk, Jan (2025): Two new species of Clytrini from South-east Asia (Coleoptera: Chrysomelidae: Cryptocephalinae). Zootaxa 5689 (2): 379-391, DOI: 10.11646/zootaxa.5689.2.8, URL: https://doi.org/10.11646/zootaxa.5689.2.8
3D6F384EFFDFFF9B85E6A000FF4AFDD9.taxon	distribution	Distribution. Bhutan, India (Assam, Sikkim, West Bengal), Nepal (Jacoby 1903, Medvedev 1988 a, present paper).	en	Romantsov, Pavel V., Bezděk, Jan (2025): Two new species of Clytrini from South-east Asia (Coleoptera: Chrysomelidae: Cryptocephalinae). Zootaxa 5689 (2): 379-391, DOI: 10.11646/zootaxa.5689.2.8, URL: https://doi.org/10.11646/zootaxa.5689.2.8
3D6F384EFFDDFF9885E6A6EEFDBAFC98.taxon	description	(Figs 13 – 15, 19, 24)	en	Romantsov, Pavel V., Bezděk, Jan (2025): Two new species of Clytrini from South-east Asia (Coleoptera: Chrysomelidae: Cryptocephalinae). Zootaxa 5689 (2): 379-391, DOI: 10.11646/zootaxa.5689.2.8, URL: https://doi.org/10.11646/zootaxa.5689.2.8
3D6F384EFFDDFF9885E6A6EEFDBAFC98.taxon	materials_examined	Type locality. “ Vietnam, Thai Nguyen Prov., vic. Ngoc Thanh, vic. Me Linh (IEBR station), 21 ° 23 ' 3 " N, 105 ° 42 ' 44 " E ”. Type specimens examined. Holotype: ♂ (NMEG), “ N-VIETNAM Thai Nguyen Pr. / vic. Ngoc Thanh, Le Minh / (IEBR station), 12. V. 2012 / 21 ° 23 ' 3 " N, 105 ° 42 ' 44 " E / 60 m, leg. A. Weigel by light [w, p] // collection / NATURKUNDE / MUSEUM ERFURT [y, p] // HOLOTYPUS [p] / Smaragdina / vietnamica [h] / L. Medvedev [r, p] ”. Paratypes: 2 ♀♀ (NMEG), “ N-VIETNAM Thai Nguyen Pr. / vic. Ngoc Thanh, Le Minh / (IEBR station), 12. V. 2012 / 21 ° 23 ' 3 " N, 105 ° 42 ' 44 " E / 60 - 80 m, leg. A. Weigel [w, p] // collection / NATURKUNDE / MUSEUM ERFURT [y, p] // PARATYPUS [p] / Smaragdina / vietnamica [h] / L. Medvedev [r, p] ”; 1 ♀ (ZIN), “ N-Vietnam Thai Nguyen Pr. / vic. Ngoc Thanh, Le Minh / (IEBR station), 12. V. 2012 / 21 ° 23 ' 3 " N, 105 ° 42 ' 44 " E / 60 m, leg. A. Weigel by light [w, p] // PARATYPUS [p] / Smaragdina / vietnamica [h] / L. Medvedev [r, p] // collection / NATURKUNDE / MUSEUM ERFURT [y, p] ”. Additional material examined. 6 specimens — VIETNAM: Thai Nguyen Prov., vic. Ngoc Thanh, Le Minh, IEBR station, 21 ° 23 ' 3 " N, 105 ° 42 ' 44 " E, 12. v. 2012, 1 ♂, A. Weigel leg. (ZIN); Vinh Phuc Prov., Melinh Biodiversity Station, 21 ° 23 ' 04 " N 105 ° 42 ' 44 " E, 80 - 200 m, 9. - 12. vi. 2011, 2 ♂♂ 3 ♀♀, E. Jendek leg. (NMPC).	en	Romantsov, Pavel V., Bezděk, Jan (2025): Two new species of Clytrini from South-east Asia (Coleoptera: Chrysomelidae: Cryptocephalinae). Zootaxa 5689 (2): 379-391, DOI: 10.11646/zootaxa.5689.2.8, URL: https://doi.org/10.11646/zootaxa.5689.2.8
3D6F384EFFDDFF9885E6A6EEFDBAFC98.taxon	distribution	Distribution. North Vietnam.	en	Romantsov, Pavel V., Bezděk, Jan (2025): Two new species of Clytrini from South-east Asia (Coleoptera: Chrysomelidae: Cryptocephalinae). Zootaxa 5689 (2): 379-391, DOI: 10.11646/zootaxa.5689.2.8, URL: https://doi.org/10.11646/zootaxa.5689.2.8
