taxonID	type	description	language	source
3F5B8791FFE1135BFF2BFC88FC853EA6.taxon	materials_examined	Type species: Symphytopria fulva Kieffer, designated by Kieffer (1912).	en	Chemyreva, Vasilisa G. (2024): Revision of the genus Symphytopria Kieffer (Hymenoptera, Diapriidae, Diapriinae) with notes on its taxonomy. Zootaxa 5541 (3): 375-382, DOI: 10.11646/zootaxa.5541.3.8, URL: http://dx.doi.org/10.11646/zootaxa.5541.3.8
3F5B8791FFE1135BFF2BFC88FC853EA6.taxon	diagnosis	Diagnosis. Body small to medium-sized (1.5 ‒ 2.5 mm), usually pale, mainly reddish or yellowish to partly dark brown, mostly smooth, with abundant setae. Setose cushions developed on postgenae, pronotum, base of T 2 and S 2; mesosoma ventrally, metapleuron, propodeum and petiole entirely pubescent. Head globular or subrectangular in dorsal view. Eye small, setose, circular and distinctly shorter than malar distance. Ocelli strongly reduced to absent. Antennal shelf distinctly projecting; toruli contiguous. Clypeus not separated dorsally from supraclypeal area by epistomal sulcus (Fig. 3 C). Malar sulcus and tentorial pits absent. Labrum not exposed. Maxillar palpi 5 - segmented and labial palpi 3 - segmented. Antenna of male 14 - and female 13 - segmented. Female antennae with abrupt 3 - segmented clava; A 1 as long as A 2 ‒ A 5 or A 2 ‒ A 6 measured together; A 4 ‒ A 9 small, as long as wide to transverse. Male antennae filiform with A 4 modified. Mesosoma elongate and slightly compressed. Pronotum almost invisible in dorsal view, mesonotum and pronotal collar almost perpendicular to each other; lateral side of pronotum smooth. Scutellum fused with mesoscutum and metanotum, usually without any sculpture, sometimes with two very shallow anterior scutellar pits and weak traces of posterior scutellar pits. Propodeum slightly transverse; in dorsal view propodeum 0.8 ‒ 1.0 times as long as projected part of petiole and 0.33 ‒ 0.43 times as long as mesonotum and metanotum combined (length of all structured measured medially). Mesopleuron narrow, totally bare and smooth; epicnemial pit absent. Tegula tiny, fore wing strongly reduced and, at most, reaching only middle of propodeum. All legs with trochantelli. In lateral view petiole strongly humped dorsally in its anterior part and usually elevated above T 2, while its posterior part situated lower than T 2 and covered by deep grooves (Figs 1 E, 4 D). Median and lateral grooves present on T 2 and S 2 (Fig. 4 D, E).	en	Chemyreva, Vasilisa G. (2024): Revision of the genus Symphytopria Kieffer (Hymenoptera, Diapriidae, Diapriinae) with notes on its taxonomy. Zootaxa 5541 (3): 375-382, DOI: 10.11646/zootaxa.5541.3.8, URL: http://dx.doi.org/10.11646/zootaxa.5541.3.8
3F5B8791FFE1135BFF2BFC88FC853EA6.taxon	discussion	Remarks. Although the tribal taxonomy of the subfamily Diapriinae is well treated (Masner & García 2002), the position of Symphytopria within the subfamily is doubtful. The antennal formula of the genus (male 14 - and female 13 - segmented antennae) agrees with the diagnosis of Diapriini given by Masner & García (2002), but the same antennal formula is only known for the species-poor genus Monelata Förster, which is morphologically distant from Symphytopria. Furthermore, the structure of the T 2 and S 2 bases in Symphytopria does not match the characters of the known Diapriini genera. I suggest that Symphytopria is closely related to the genus Idiotypa Förster and should be placed in the tribe Spilomicrini (Chemyreva et al. 2021). According to Masner & García (2002) the genera of the tribe Spilomicrini comprise diapirid wasps with “ antennal formula most frequently 13 – 13, rarely 12 – 13, exceptionally 14 – 14 ” and submarginal vein in fore wing “ distinctly to remarkably remote from fore margin, costal cell relatively wide ”. Unfortunately, the wing venation of Symphytopria species cannot be described, as all the known species of the genus have very tiny wings with reduced venation. However, 13 - segmented antennae in females and 14 - segmented in males are known in this tribe and only the combination of such characters in males and females of the same species is unique. This feature, together with the character of the T 2 and S 2 base structure and some other additional features: anterior scutellar pit, if present (like a trace), bifoveate or replaced by arc of minute crenulae; presence of trace of posterior scutellar pits in some specimens; dense cushion at the base of S 2, persuaded me to associate this genus with the genera of the tribe Spilomicrini. Species of this tribe most similar to Symphytopria are Idiotypa nitens (Szabó) and Spilomicrus antennatus (Jurine), which have much in common with Symphytopria species (Chemyreva 2021; Chemyreva at al. 2021).	en	Chemyreva, Vasilisa G. (2024): Revision of the genus Symphytopria Kieffer (Hymenoptera, Diapriidae, Diapriinae) with notes on its taxonomy. Zootaxa 5541 (3): 375-382, DOI: 10.11646/zootaxa.5541.3.8, URL: http://dx.doi.org/10.11646/zootaxa.5541.3.8
3F5B8791FFE3135DFF2BFF5CFE5F3A1E.taxon	description	(Figs 1, 2, 3)	en	Chemyreva, Vasilisa G. (2024): Revision of the genus Symphytopria Kieffer (Hymenoptera, Diapriidae, Diapriinae) with notes on its taxonomy. Zootaxa 5541 (3): 375-382, DOI: 10.11646/zootaxa.5541.3.8, URL: http://dx.doi.org/10.11646/zootaxa.5541.3.8
3F5B8791FFE3135DFF2BFF5CFE5F3A1E.taxon	materials_examined	Material examined. Type material. Holotype of Symphytopria fulva Kieffer: female, “ Castatbuono / Sieilia / V – 1906 / A. Dodero ” (Fig. 1 B). Holotype of Symphytopria fusciceps Kieffer: female, “ Aritzo / (Sardinia) / 30. V. 1902 / A. Dodero ” (Fig. 1 F). Holotype of Symphytopria nigroclavata Kieffer: female, “ Tempio / Sardegna / 21. IV. 903 / Dodero ” (Fig. 2 B). Additional material. Italy: M. Arqueri, presso Seui, Sardegna, iv. 1909, Salari [leg.], 2 ♀; Bannari, Sard. 9. v. [1] 909, A. Dodero [leg.], 1 ♀; Ficuzza, Sicilia, v. 1912, A. Dodero [leg.], 5 ♀; Aladei Sardi, Sard. v. [19] 07, T. Derosas [leg.], 1 ♀; Sardegna, Massiccio del Gennargentu (NU) Bruncu Spina, m 1550, 9. v. 2000, R. Poggi [leg.], 2 ♀; Sardegna, Cala Gonone (NU), 20. v. 1980, R. Poggi [leg.], 1 ♀; Sicilia sett., Madonie, Piano Zucchi (PA) sub Crataegus, 9. vii. 2011, m 1200, R. Poggi [leg.], 1 ♀; Sicilia, Ficuzza, Rocca Busambra (PA) 25. x. 1982, m 1200, R. Poggi [leg.], 2 ♀, 1 ♂; Sicilia, Ficuzza, Bosco d. Cappelliere (PA) 25. x. 1982, m 800, R. Poggi [leg.], 1 ♀; Sardegna, Aritzo, 1 – 10. v. 1910, Solari [leg.], 2 ♀; Aritzo, Sard., xi. 1911, A. Dodero [leg.], 1 ♀.	en	Chemyreva, Vasilisa G. (2024): Revision of the genus Symphytopria Kieffer (Hymenoptera, Diapriidae, Diapriinae) with notes on its taxonomy. Zootaxa 5541 (3): 375-382, DOI: 10.11646/zootaxa.5541.3.8, URL: http://dx.doi.org/10.11646/zootaxa.5541.3.8
3F5B8791FFE3135DFF2BFF5CFE5F3A1E.taxon	diagnosis	Diagnosis. Head concolorous with meso- and metasoma or slightly darker, with a row of four pits between eyes (Figs 1 D, 2 C); border between scutellum and mesonotum without any trace of sculpture (Figs 1, 2); propodeum without median carina and with distinctly arcuate posterior margin (Figs 1 C, 1 D, 3 A); petiole and base of T 2 usually densely pubescent.	en	Chemyreva, Vasilisa G. (2024): Revision of the genus Symphytopria Kieffer (Hymenoptera, Diapriidae, Diapriinae) with notes on its taxonomy. Zootaxa 5541 (3): 375-382, DOI: 10.11646/zootaxa.5541.3.8, URL: http://dx.doi.org/10.11646/zootaxa.5541.3.8
3F5B8791FFE3135DFF2BFF5CFE5F3A1E.taxon	description	Description. Male. Length of body about 2.0 mm. Body and appendages yellowish-brown, smooth. Top of head with four pits between eyes arranged in a row. Antennal shelf weakly sculptured along lower margin on frontal and lateral sides. The largest pleurostomal distance 0.8 times as long as the shortest distance between eyes. Eye as high as wide. Malar distance about 1.3 times as long as eye diameter. Antenna 14 - segmented, entirely yellowish-brown, filiform; A 4 the longest segment of flagellum, broadened apically and with straight carina developed from the base to top of this segment. Scutellum fused with mesonotum and metanotum, all smooth and scarcely setose. Mesopleuron narrow, smooth, bare laterally and pubescent ventrally. Petiole entirely pubescent, in lateral view swollen dorsally in its anterior part and distinctly elevated above margin of T 2; its posterior part situated lower than margin of T 2. Base of T 2 and S 2 with median and lateral grooves, densely pubescent while most of surface of T 2 and subsequent tergites and sternites smooth and scarcely setose; T 3 – T 5 short and equal in length; T 6 and T 7 short and weakly transverse, at rest situated almost perpendicularly to body axis; S 3 – S 6 short, almost equal in length; S 7 semicircular, finally punctured. Variation. Body length 1.8 – 2.5 mm. Metasoma dark brown to yellowish; clavomeres very weakly to distinctly darker than A 1 – A 10. A 4 – A 9 slightly to distinctly transverse. Antennal shelf with distinct (Fig. 2 A) to weak vertical grooves (Fig. 3 C) on frontal and lateral sides. Posterior half of plicae in lateral view curved upward or straight (Fig. 1 A, E). The hump of petiole from almost bare on top to completely densely pubescent; in lateral view the top of hump situated higher or at the same level as anterior margin of T 2.	en	Chemyreva, Vasilisa G. (2024): Revision of the genus Symphytopria Kieffer (Hymenoptera, Diapriidae, Diapriinae) with notes on its taxonomy. Zootaxa 5541 (3): 375-382, DOI: 10.11646/zootaxa.5541.3.8, URL: http://dx.doi.org/10.11646/zootaxa.5541.3.8
3F5B8791FFE3135DFF2BFF5CFE5F3A1E.taxon	distribution	Distribution. Italy.	en	Chemyreva, Vasilisa G. (2024): Revision of the genus Symphytopria Kieffer (Hymenoptera, Diapriidae, Diapriinae) with notes on its taxonomy. Zootaxa 5541 (3): 375-382, DOI: 10.11646/zootaxa.5541.3.8, URL: http://dx.doi.org/10.11646/zootaxa.5541.3.8
3F5B8791FFE4135CFF2BFA49FDDB38F9.taxon	description	(Fig. 4)	en	Chemyreva, Vasilisa G. (2024): Revision of the genus Symphytopria Kieffer (Hymenoptera, Diapriidae, Diapriinae) with notes on its taxonomy. Zootaxa 5541 (3): 375-382, DOI: 10.11646/zootaxa.5541.3.8, URL: http://dx.doi.org/10.11646/zootaxa.5541.3.8
3F5B8791FFE4135CFF2BFA49FDDB38F9.taxon	materials_examined	Material examined. Type material. Holotype of Symphytopria trisulcata Kieffer: female, “ Nava / VIII. 902 / Solari ” (Fig. 4 G). Holotype of Symphytopria facialis Kieffer: female, “ S. Bisso Ceraco / M. Sacro / prov. Salerno / Bensa 902 ” (Fig. 4 F). Additional material. Italy: Lippiano, Prov. Arezzo [Perugia Prov.], xii. 1917, A. Andreini [leg], 1 ♀; Prov. di Lucca, Bogni di L. [Lucca], 30. iii. 1910, A. Andreini [leg], 1 ♀; Gran Sasso, Prati di Tivo, faggeta [beech forest] (TE [Teramo Prov.]), 17. vi. 1988, R. Poggi [leg], m 1550, 1 ♀. France se, Dep. Var Dint Tanneron (NW di Cannes [NW from Cannes]), 31. iii. 2008, m 170, R. Poggi [leg], 1 ♀.	en	Chemyreva, Vasilisa G. (2024): Revision of the genus Symphytopria Kieffer (Hymenoptera, Diapriidae, Diapriinae) with notes on its taxonomy. Zootaxa 5541 (3): 375-382, DOI: 10.11646/zootaxa.5541.3.8, URL: http://dx.doi.org/10.11646/zootaxa.5541.3.8
3F5B8791FFE4135CFF2BFA49FDDB38F9.taxon	diagnosis	Diagnosis. Head distinctly darker than meso- and metasoma, without a row of pits between eyes; border between scutellum and mesonotum with more or less distinct trace of sculpture (Fig. 4 C, D); propodeum with more or less distinct median carina and with weakly arcuate posterior margin (Fig. 4 A, D); petiole and base of T 2 scanty pubescent.	en	Chemyreva, Vasilisa G. (2024): Revision of the genus Symphytopria Kieffer (Hymenoptera, Diapriidae, Diapriinae) with notes on its taxonomy. Zootaxa 5541 (3): 375-382, DOI: 10.11646/zootaxa.5541.3.8, URL: http://dx.doi.org/10.11646/zootaxa.5541.3.8
3F5B8791FFE4135CFF2BFA49FDDB38F9.taxon	description	Variation. Body length 2.1 – 2.4 mm. Metasoma dark brown to reddish, female clavomeres dark brown to brown. A 4 – A 9 slightly to distinctly transverse, A 11 and A 12 as long as wide to distinctly transverse (Fig. 4 A – C). Mesosoma in dorsal view 1.6 to 2.0 times as long as wide; shallow anterior scutellar pits visible as traces (Fig. 4 B, D) to almost absent (Fig. 4 A, C); median propodeal carina straight, distinct and darker than other parts of propodeum (Fig. 4 A) to slightly curved, poorly visible and same color as other parts of propodeum (Fig. 4 D).	en	Chemyreva, Vasilisa G. (2024): Revision of the genus Symphytopria Kieffer (Hymenoptera, Diapriidae, Diapriinae) with notes on its taxonomy. Zootaxa 5541 (3): 375-382, DOI: 10.11646/zootaxa.5541.3.8, URL: http://dx.doi.org/10.11646/zootaxa.5541.3.8
3F5B8791FFE4135CFF2BFA49FDDB38F9.taxon	distribution	Distribution. Italy and France.	en	Chemyreva, Vasilisa G. (2024): Revision of the genus Symphytopria Kieffer (Hymenoptera, Diapriidae, Diapriinae) with notes on its taxonomy. Zootaxa 5541 (3): 375-382, DOI: 10.11646/zootaxa.5541.3.8, URL: http://dx.doi.org/10.11646/zootaxa.5541.3.8
