taxonID	type	description	language	source
8A245ADF197F5E3996445568C1814AF1.taxon	description	Figs 1 A, B, 3 A – D, 5 A, 7 A – D, 9 A, 11 A, 13 A, B, 14 A, B, 15 A Japanese name: Kushiromaru-gunbai	en	Souma, Jun (2025): A new species and two new records of the moss-feeding lace bug genus Acalypta (Hemiptera, Heteroptera, Tingidae) from Hokkaido, northern Japan, with an illustrated key to the Japanese species of the genus. ZooKeys 1229: 107-132, DOI: 10.3897/zookeys.1229.142344
8A245ADF197F5E3996445568C1814AF1.taxon	diagnosis	Diagnosis. Acalypta alutacea sp. nov. is recognized among the other species of Acalypta based on a combination of the following characteristics: brachypterous morph only known; hemelytron without dark spots (Fig. 1 A, B); antenniferous tubercle obtuse, curved inward (Figs 3 A – D, 13 A, B); basal part of antennal segment III not thickened; rostrum reaching anterior part of abdominal sternite II (Fig. 5 A); pronotum 3 / 4 times as long as maximum width across paranota; pronotal disc as long as calli; punctures on pronotal disc smaller than areolae of posterior process; posterior part of hood usually triangular; lateral carina of pronotum absent or reduced, shorter than hood, without or with a single minute areola; calli smooth; paranotum as wide as hood, not narrowed posteriorly, with 3 – 4 rows of areolae throughout its length; anterolateral angle of paranotum rounded, not or weakly protruding anteriorly, not reaching mid-level of compound eye; posterolateral angle of paranotum protruding posteriorly; posterior process strongly protruding posteriorly, as long as hood; costal area of hemelytron with 2 – 3 rows of areolae in basal part, a single row in middle part, and 1 – 2 rows in apical part (Fig. 7 A – D); discoidal area expanded beyond apical fourth of hemelytron, wider than subcostal area; Cu (cubitus) vein distinct throughout its length; abdomen pale brown in female (Fig. 11 A); and pygophore roundly inflated (Fig. 14 A).	en	Souma, Jun (2025): A new species and two new records of the moss-feeding lace bug genus Acalypta (Hemiptera, Heteroptera, Tingidae) from Hokkaido, northern Japan, with an illustrated key to the Japanese species of the genus. ZooKeys 1229: 107-132, DOI: 10.3897/zookeys.1229.142344
8A245ADF197F5E3996445568C1814AF1.taxon	description	Description. Brachypterous male. Frontal spine of head, pronotum except for calli, hemelytron, and sternal laminae pale brown; antenniferous tubercle, antennal segments I to III, ventral surface of thorax except for sternal laminae, and legs brown; most parts of head, antennal segment IV, calli, and abdomen dark brown; paranotum and hemelytron sometimes irregularly dark in very small sections, without dark spots; areolae of pronotum and hemelytron transparent; compound eyes dark red; pubescence on body yellowish (Figs 1 A, 3 A, B, 5 A, 7 A, B, 9 A). Body ovate; pubescence on most parts of body distinctly shorter than radius of compound eye (Figs 1 A, 3 A, B, 5 A, 9 A). Head covered with minute pubescence; pair of frontal spines separated at apices, reaching apex of clypeus; antenniferous tubercle obtuse, curved inward, shorter than frontal spine; vertex and clypeus smooth. Compound eye round in dorsal view. Antenna densely covered with minute pubescence in segments I to III and long pubescence in segment IV; pubescence on segment IV longer than pubescence on other parts of body; segment I cylindrical, shorter than segment IV; segment II conical, shortest amongst antennal segments; segment III linear, longest amongst antennal segments, not thickened in basal part; segment IV fusiform. Bucculae closed at anterior ends, with 3 rows of areolae at highest part. Rostrum reaching anterior part of abdominal sternite II. Pronotum (Figs 3 A, B, 13 A, B) glabrous, 3 / 4 times as long as maximum width across paranota. Pronotal disc coarsely punctate, as long as calli; punctures smaller than areolae of posterior process. Hood roof-shaped, shorter than median carina of pronotum, wider than vertex at widest part, not covering compound eye, with 5 rows of areolae at highest part; posterior part usually triangular. Median carina straight, extending to apex of posterior process, with a single row of areolae throughout its length. Lateral carina absent or reduced, shorter than hood, without or with a single minute areola. Calli smooth. Paranotum horizontally expanding outward, as wide as hood, not narrowed posteriorly, with 3 – 4 rows of areolae throughout its length; anterolateral angle rounded, not or weakly protruding anteriorly, not reaching mid-level of compound eye; posterolateral angle protruding posteriorly. Posterior process triangular, obtuse at apex, strongly protruding posteriorly, as long as hood. Hemelytron (Fig. 7 A, B) glabrous, extending beyond apex of abdomen; apex close to the other at rest; costal area with 2 – 3 rows of areolae in basal part, a single row in middle part, and 1 – 2 rows in apical part; subcostal area with 4 rows of areolae at widest part; discoidal area expanded beyond apical fourth of hemelytron, wider than subcostal area, with 4 – 5 rows of areolae at widest part; sutural area with 3 rows of areolae at widest part; hypocostal lamina with a single row of areolae throughout its length; Sc (subcosta), Hc (hypocosta), R + M (fused radius and media) and Cu (cubitus) veins distinct throughout their respective length. Thoracic pleura (Fig. 5 A) smooth in anterior part, coarsely punctate in posterior part. Ostiolar peritreme reduced. Mesosternum narrower than metasternum. Sternal laminae lower than buccula, open in both anterior and posterior ends; metasternal lamina as high as mesosternal lamina. Legs (Fig. 1 A) smooth, covered with minute pubescence; femora thickest at middle. Abdomen ovate in dorsal and ventral views, glabrous except for terminalia, smooth. Pygophore (Figs 9 A, 14 A) roundly inflated, semicircular in ventral view, covered with minute pubescence; anterior margin concave in middle part. Paramere (Fig. 14 B) slender, expanded in middle part, gently curved inward, covered with minute pubescence in middle part of outer and inner margins. Measurements (n = 3). Body length with hemelytra 2.30 – 2.60 mm; maximum width across hemelytra 1.30 – 1.50 mm; length of antennal segments I to IV 0.15 mm, 0.10 mm, 0.70 – 0.80 mm, and 0.20 – 0.25 mm, respectively; pronotal length 0.70 – 0.80 mm; pronotal width across paranota 1.10 – 1.20 mm; hemelytral length 1.65 – 1.85 mm; maximum width of hemelytron 0.70 – 0.80 mm. Brachypterous female. General habitus very similar to that of male (Figs 1 B, 3 C, D, 7 C, D, 11 A) except for the following characters: abdomen pale brown; body usually longer and wider than in male; antennal segment III shorter than in male; and apical part of abdomen pentagonal in ventral view. Measurements (n = 5). Body length with hemelytra 2.60 – 2.80 mm; maximum width across hemelytra 1.65 – 1.80 mm; length of antennal segments I to IV 0.15 mm, 0.10 mm, 0.60 mm, and 0.20 mm, respectively; pronotal length 0.80 – 0.90 mm; pronotal width across paranota 1.20 – 1.30 mm; hemelytral length 1.90 – 2.00 mm; maximum width of hemelytron 0.85 – 0.95 mm.	en	Souma, Jun (2025): A new species and two new records of the moss-feeding lace bug genus Acalypta (Hemiptera, Heteroptera, Tingidae) from Hokkaido, northern Japan, with an illustrated key to the Japanese species of the genus. ZooKeys 1229: 107-132, DOI: 10.3897/zookeys.1229.142344
8A245ADF197F5E3996445568C1814AF1.taxon	distribution	Distribution. Japan (Hokkaido) (Fig. 17) (Nakatani 2016, 2018).	en	Souma, Jun (2025): A new species and two new records of the moss-feeding lace bug genus Acalypta (Hemiptera, Heteroptera, Tingidae) from Hokkaido, northern Japan, with an illustrated key to the Japanese species of the genus. ZooKeys 1229: 107-132, DOI: 10.3897/zookeys.1229.142344
8A245ADF197F5E3996445568C1814AF1.taxon	etymology	Etymology. The species epithet is the Latin adjective “ alutaceus ”, referring to the pale brown body color.	en	Souma, Jun (2025): A new species and two new records of the moss-feeding lace bug genus Acalypta (Hemiptera, Heteroptera, Tingidae) from Hokkaido, northern Japan, with an illustrated key to the Japanese species of the genus. ZooKeys 1229: 107-132, DOI: 10.3897/zookeys.1229.142344
E3A239E19528595FAC3C9484B4E62C9B.taxon	description	Figs 1 C, D, 3 E – G, 5 B, 7 E – G, 9 B, 11 B, 13 C, D, 15 B – D Japanese name: Yachimaru-gunbai	en	Souma, Jun (2025): A new species and two new records of the moss-feeding lace bug genus Acalypta (Hemiptera, Heteroptera, Tingidae) from Hokkaido, northern Japan, with an illustrated key to the Japanese species of the genus. ZooKeys 1229: 107-132, DOI: 10.3897/zookeys.1229.142344
E3A239E19528595FAC3C9484B4E62C9B.taxon	distribution	Distribution. Austria, Belgium, Byelorussia, Croatia, Czech Republic, Denmark, Estonia, Finland, France, Great Britain, Germany, Hungary, Ireland, Italy, Japan (Hokkaido, Rishiri Island), Latvia, Luxembourg, Moldovia, Mongolia, Netherlands, Norway, Poland, Romania, Russia, Slovakia, Slovenia, Spain, Sweden, Switzerland (Fig. 17) (Péricart and Golub 1996; Aukema et al. 2013; Aukema 2025). Acalypta carinata is newly recorded from Japan.	en	Souma, Jun (2025): A new species and two new records of the moss-feeding lace bug genus Acalypta (Hemiptera, Heteroptera, Tingidae) from Hokkaido, northern Japan, with an illustrated key to the Japanese species of the genus. ZooKeys 1229: 107-132, DOI: 10.3897/zookeys.1229.142344
78A92F96FB6557BC83B4504429BB3B11.taxon	description	Figs 2 E, 4 E, 6 C, 8 E, 10 C, 12 C, 16 C, D Japanese name: Maru-gunbai	en	Souma, Jun (2025): A new species and two new records of the moss-feeding lace bug genus Acalypta (Hemiptera, Heteroptera, Tingidae) from Hokkaido, northern Japan, with an illustrated key to the Japanese species of the genus. ZooKeys 1229: 107-132, DOI: 10.3897/zookeys.1229.142344
78A92F96FB6557BC83B4504429BB3B11.taxon	distribution	Distribution. Japan (Hokkaido, Honshu, Awa Island, Sado Island, Awaji Island, Oki Islands (Dogo Island), Ikuchi Island, Shikoku, Shodo Island, Omi Island, Kyushu, Amakusa Islands (Shimoshima Island )) (Fig. 17) (Tomokuni 1983, 1985; Nozaki et al. 2016; Yamada and Tomokuni 2012; Yamada and Ishikawa 2016; Aukema 2025). Acalypta sauteri is newly recorded from Hokkaido and Awa Island. The discovery of A. sauteri from Hokkaido also represents the northernmost record of this species.	en	Souma, Jun (2025): A new species and two new records of the moss-feeding lace bug genus Acalypta (Hemiptera, Heteroptera, Tingidae) from Hokkaido, northern Japan, with an illustrated key to the Japanese species of the genus. ZooKeys 1229: 107-132, DOI: 10.3897/zookeys.1229.142344
