taxonID	type	description	language	source
292987A43910C762FC0FF8291E474019.taxon	type_taxon	Type species. Onchidium peronii Cuvier, 1804, by monotypy. 1538 (AM C. 448363) Queensland 2870 (UMIZ 00169) Indonesia, Seram 1797 (UMIZ 00180) Indonesia, Sumatra P. verruculata unit # 1 5481 (MNHN-IM- 2013 - 62392) Vanuatu 1746 (UMIZ 00178) Indonesia, Sumatra P. verruculata unit # 2 990 (ZRC. MOL. 10496) Singapore 2546 (USMMC 00065) Peninsular Malaysia P. verruculata unit # 3 974 (USMMC 00064) Peninsular Malaysia 6164 (MNHN-IM- 2019 - 1384) Pakistan 6166 (MNHN-IM- 2019 - 1386) Pakistan P. verruculata unit # 4 6165 (MNHN-IM- 2019 - 1385) Pakistan 5510 3146 ((MNHN-IM- MNHN-IM- 2013 2019 - - 62398 1611)) Madagascar Mozambique P. verruculata unit # 5 3140 (MNHN-IM- 2019 - 1610) Madagascar 3753 (NSMT-Mo 78987) Japan, Wakayama 5383 (NSMT-Mo 78985) Japan, Wakayama Holotype P. setoensis 5384 (NSMT-Mo 78986) Japan, Wakayama 3157 (MNHN-IM- 2000 - 35265) Mauritius Holotype 6095 (MNHN-IM- 2013 - 53535) PNG, New Ireland P. griffithsi 2934 (UMIZ 00177) Indonesia, Kei 1620 (NTM P. 57626) Northern Territory 1655 (NTM P. 57626) Northern Territory 1623 (NTM P. 57627) Northern Territory 1667 (NTM P. 57627) Northern Territory P. willani 1628 (NTM P. 57625) Northern Territory Holotype 1626 (NTM P. 57627) Northern Territory 6388 NT species T 6386 NT species T 6390 NT species T P. watts New Species 6387 NT species T Holotype 6160 (BPBM 284527) Hawaii 706 (UF 303653) Hawaii P. platei 5412 (MNHN-IM- 2013 - 13351) PNG, Madang 735 (NHMUK 20060414) Mozambique 703 (UF 332088) Oman P. madagascariensis 5501 (MNHN-IM- 2009 - 16392) Madagascar 696 - 2 (UF 352288) Japan, Okinawa 696 - 3 (UF 352288) Japan, Okinawa P. okinawensis 696 - 4 (UF 352288) Japan, Okinawa Holotype 1553 (MNHN-IM- 2019 - 1607) Mauritius 5840 (MNHN-IM- 2019 - 1609) Guam P. peronii 5476 (MNHN-IM- 2013 - 16260) PNG, Madang 965 Onchidium typhae	en	Dayrat, Benoît, , Adam J. Bourke, Goulding,, Tricia C. (2025): A new species of Peronia Fleming, 1822 (Gastropoda, Euthyneura, Pulmonata, Onchidiidae) from the Northern Territory, Australia. Raffles Bulletin of Zoology 73: 110-124, DOI: 10.26107/RBZ-2025-0008
292987A43910C762FC0FF8291E474019.taxon	description	1001 Platevindex luteus 1183 Marmaronchis vaigiensis 1011 Melayonchis eloisae 960 Peronina tenera 2740 Wallaconchis sinanui 1732 Paromoionchis tumidus 5137 Alionchis jailoloensis 5013 Onchidella celtica 1688 Laspionchis boucheti 1523 Onchidina australis Fig. 2. Maximum parsimony consensus tree within Peronia based on ITS 2 sequences (identical sequences are represented as a single haplotype sequence). Numbers by the branches are the bootstrap values. Only the values> 50 % are shown. For the sake of simplicity, the colour used for each species here is the same as the colour used in the figures published in our recent revision of Peronia (Dayrat et al., 2020). Numbers for each individual correspond to unique identifiers for DNA extraction which can be found in our revision as well (Dayrat et al., 2020: table 2). 1538 (AM C. 448363) Queensland 2870 (UMIZ 00169) Indonesia, Seram 1797 (UMIZ 00180) Indonesia, Sumatra P. verruculata unit # 1 5481 (MNHN-IM- 2013 - 62392) Vanuatu 1746 (UMIZ 00178) Indonesia, Sumatra P. verruculata unit # 2 990 (ZRC. MOL. 10496) Singapore 2546 (USMMC 00065) Peninsular Malaysia P. verruculata unit # 3 974 (USMMC 00064) Peninsular Malaysia 6164 (MNHN-IM- 2019 - 1384) Pakistan 6166 (MNHN-IM- 2019 - 1386) Pakistan P. verruculata unit # 4 6165 (MNHN-IM- 2019 - 1385) Pakistan 3146 (MNHN-IM- 2019 - 1611) Madagascar 5510 (MNHN-IM- 2013 - 62398) Mozambique P. verruculata unit # 5 3140 (MNHN-IM- 2019 - 1610) Madagascar 3753 (NSMT-Mo 78987) Japan, Wakayama 5383 (NSMT-Mo 78985) Japan, Wakayama Holotype P. setoensis 5384 (NSMT-Mo 78986) Japan, Wakayama 3157 (MNHN-IM- 2000 - 35265) Mauritius Holotype 6095 (MNHN-IM- 2013 - 53535) PNG, New Ireland P. griffithsi 2934 (UMIZ 00177) Indonesia, Kei 1620 (NTM P. 57626) Northern Territory 1655 (NTM P. 57626) Northern Territory 1623 (NTM P. 57627) Northern Territory 1667 (NTM P. 57627) Northern Territory P. willani 1628 (NTM P. 57625) Northern Territory Holotype 1626 (NTM P. 57627) Northern Territory 6388 NT species T 6386 NT species T 6390 NT species T P. watts New Species 6387 NT species T Holotype 6160 (BPBM 284527) Hawaii 706 (UF 303653) Hawaii P. platei 5412 (MNHN-IM- 2013 - 13351) PNG, Madang 735 (NHMUK 20060414) Mozambique 703 (UF 332088) Oman P. madagascariensis 5501 (MNHN-IM- 2009 - 16392) Madagascar 696 - 4 (UF 352288) Japan, Okinawa Holotype 696 - 3 (UF 352288) Japan, Okinawa P. okinawensis 696 - 2 (UF 352288) Japan, Okinawa 1553 (MNHN-IM- 2019 - 1607) Mauritius 5840 (MNHN-IM- 2019 - 1609) Guam P. peronii 5476 (MNHN-IM- 2013 - 16260) PNG, Madang 965 Onchidium typhae 1001 Platevindex luteus 1183 Marmaronchis vaigiensis 1011 Melayonchis eloisae 960 Peronina tenera 2740 Wallaconchis sinanui 1732 Paromoionchis tumidus 5137 Alionchis jailoloensis 1688 Laspionchis boucheti 5013 Onchidella celtica 1523 Onchidina australis Fig. 3. Maximum parsimony consensus tree within Peronia based on concatenated nuclear ITS 2 and 28 S sequences. Numbers by the branches are the bootstrap values. Only the values> 50 % are shown. For the sake of simplicity, the colour used for each species here is the same as the colour used in the figures published in our recent revision of Peronia (Dayrat et al., 2020). Numbers for each individual correspond to unique identifiers for DNA extraction which can be found in our revision as well (Dayrat et al., 2020: table 2).	en	Dayrat, Benoît, , Adam J. Bourke, Goulding,, Tricia C. (2025): A new species of Peronia Fleming, 1822 (Gastropoda, Euthyneura, Pulmonata, Onchidiidae) from the Northern Territory, Australia. Raffles Bulletin of Zoology 73: 110-124, DOI: 10.26107/RBZ-2025-0008
292987A43910C762FC0FF8291E474019.taxon	diagnosis	Diagnosis. Body not flattened. Dorsal gills present. Dorsal eyes present. No retractable, central papilla present. Eyes at tip of short ocular tentacles. Male opening below right ocular tentacle and to its left. Foot wide. Pneumostome median, on ventral hyponotum. Intestinal loops of types I or V. Rectal gland absent. Accessory penial gland present, with muscular sac. Penis with hooks.	en	Dayrat, Benoît, , Adam J. Bourke, Goulding,, Tricia C. (2025): A new species of Peronia Fleming, 1822 (Gastropoda, Euthyneura, Pulmonata, Onchidiidae) from the Northern Territory, Australia. Raffles Bulletin of Zoology 73: 110-124, DOI: 10.26107/RBZ-2025-0008
292987A43914C76CFED7FAE1190743E9.taxon	description	(Figs. 4 – 9)	en	Dayrat, Benoît, , Adam J. Bourke, Goulding,, Tricia C. (2025): A new species of Peronia Fleming, 1822 (Gastropoda, Euthyneura, Pulmonata, Onchidiidae) from the Northern Territory, Australia. Raffles Bulletin of Zoology 73: 110-124, DOI: 10.26107/RBZ-2025-0008
292987A43914C76CFED7FAE1190743E9.taxon	materials_examined	Type locality. Fig. 4. Nightcliff foreshore, eastern side of Darwin Harbour, Darwin, Northern Territory, Australia, 12 ° 22.570 ′ S, 130 ° 50.867 ′ E, 12 June 2019, on large moist claystone boulders in a small rocky cove at low tide. Type material. Holotype, 45 / 30 mm [6387 H], designated here (NTM P. 59582). Additional material examined. Australia, same collection data as for the holotype, 1 specimen 38 / 26 mm [6388] (NTM P. 59583); Australia, same collection data as for the holotype, 1 specimen 38 / 24 mm [6389] (NTM P. 59584); Australia, same collection data as for the holotype, 1 specimen 45 / 25 mm [6390] (NTM P. 59585); Australia, same collection data as for the holotype, 1 specimen 33 / 22 mm [6386] (NTM P. 59586); Australia, same collection data as for the holotype, 1 specimen 40 / 25 mm [6391] (NTM P. 59587).	en	Dayrat, Benoît, , Adam J. Bourke, Goulding,, Tricia C. (2025): A new species of Peronia Fleming, 1822 (Gastropoda, Euthyneura, Pulmonata, Onchidiidae) from the Northern Territory, Australia. Raffles Bulletin of Zoology 73: 110-124, DOI: 10.26107/RBZ-2025-0008
292987A43914C76CFED7FAE1190743E9.taxon	distribution	Distribution. Endemic to Darwin, Northern Territory, Australia.	en	Dayrat, Benoît, , Adam J. Bourke, Goulding,, Tricia C. (2025): A new species of Peronia Fleming, 1822 (Gastropoda, Euthyneura, Pulmonata, Onchidiidae) from the Northern Territory, Australia. Raffles Bulletin of Zoology 73: 110-124, DOI: 10.26107/RBZ-2025-0008
292987A43914C76CFED7FAE1190743E9.taxon	etymology	Etymology. Peronia watts, new species, is named after Adam’s faithful dog that passed away during field work. The specific name ‘ watts ’ is a noun in apposition (ICZN, 1999: Article 31.1). Habitat. Fig. 4. The only specimens known were found crawling on large and moist claystone (porcellanite) boulders in a small rocky cove at low tide. It can be locally abundant in Darwin Harbour, but seems to be restricted to that specific habitat. Colour and morphology of live animals. Fig. 5. The dorsal colour of crawling animals is greyish-brown. After relaxation, the dorsal colour is light brown mottled with darker areas. Ventrally, the foot sole is light yellowish and the hyponotum is whitish. The dorsal notum is covered by dozens of papillae of various sizes. Some papillae bear up to four black dorsal eyes at their tip. The number of papillae with dorsal eyes is variable (10 – 13). The largest live animals were 45 mm long. Once preserved, all specimens are 30 mm long and 20 mm wide. Dorsal gills are few and restricted to the posterior end of the dorsal notum. They cannot be seen on live animals and can only be seen after relaxation. Live animals are very squishy, moist, and secrete an abundant sticky mucus when handled.	en	Dayrat, Benoît, , Adam J. Bourke, Goulding,, Tricia C. (2025): A new species of Peronia Fleming, 1822 (Gastropoda, Euthyneura, Pulmonata, Onchidiidae) from the Northern Territory, Australia. Raffles Bulletin of Zoology 73: 110-124, DOI: 10.26107/RBZ-2025-0008
292987A43914C76CFED7FAE1190743E9.taxon	description	Digestive system. Figs. 6, 7. Radular formulae are 82 × 145 – 1 – 145 ([6387 H] NTM P. 59582), 75 × 150 – 1 – 150 ([6388] NTM P. 59583), and 75 × 150 – 1 – 150 ([6389] NTM P. 59584). The median cusp of the rachidian teeth is up to 45 μm long. The hook of the lateral teeth is up to 70 μm long. The intestinal loops are of type I, with a transitional loop oriented between 3 to 6 o’clock. Reproductive system. Figs. 6, 8, 9. In the anterior (male) parts, the muscular sac of the accessory penial gland is between 5 and 7 mm long. The hollow spine of the accessory penial gland is narrow, elongated, and straight or only very slightly curved, and its shape (including at its tip) varies between individuals. Its length ranges from 0.9 to 1 mm. Its diameter at the conical base ranges from 100 to 130 μm. Its diameter at the tip ranges from 40 to 50 μm. The retractor muscle is thin. It can be very short, and much shorter than the penial sheath ([6387 H], [6389]), as long as the penial sheath ([6388]), or up to three times as long as the penial sheath ([6386], [6390], and [6391]). Whether the retractor muscle is short or long, it does not insert anywhere inside the visceral cavity. Inside the penial sheath, the penis is a narrow, elongated, soft, and hollow tube. Its distal end measures from 1.2 mm ([6386]) to 1.7 mm ([6389]) and bears conical hooks which are up to 70 μm long. Diagnostic features. Peronia watts, new species, is characterised by unique external and internal traits (see Table 3). It is the only Peronia species known to produce an abundant sticky mucus when handled. Internally, it is by far the Peronia species with the highest number of teeth per half row (up to 150), which is remarkable considering the relatively small size of individuals (up to 45 mm long) when compared to P. peronii (up to 115 teeth per half row for individuals up to 140 mm long). It also is the only species known in which the retractor muscle of the penis, whether long or short, does not seem to insert anywhere in the visceral cavity. The hooks of the penis are also long (up to 70 μm) considering the size of the individuals.	en	Dayrat, Benoît, , Adam J. Bourke, Goulding,, Tricia C. (2025): A new species of Peronia Fleming, 1822 (Gastropoda, Euthyneura, Pulmonata, Onchidiidae) from the Northern Territory, Australia. Raffles Bulletin of Zoology 73: 110-124, DOI: 10.26107/RBZ-2025-0008
