identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
2445B91BC164FFEBE89CFAD765EB7497.text	2445B91BC164FFEBE89CFAD765EB7497.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chortastus camerunus Schaufuss 1905	<div><p>Chortastus camerunus Schaufuss, 1905</p><p>(Figs 1–4)</p><p>Chortastus camerunus Schaufuss, 1905: 15 .</p><p>Type material. Lost during second world war [Hamburg] .</p><p>Other material. [Democratic Republic of the Congo], Stanleyville [Kisangani], Yangambi, 1.IX.1956, J. Decelle, leg. [MRCA]; Yangambi, 17.VII.1952 K. E. Schedl, leg. [NHMW]; Ngowa-Kwango, 15.XII.1938, R. P. J. Mertens leg. [RBINS]. Congo-Brazzaville [Republic of Congo], Sibiti, in lamplight, leg. Endrödy-Younga, 28.11.1963 [NHMW], new country.</p><p>Diagnosis. Body length 5.1–5.3 mm, 2.1 × as long as wide; colour black. Male frons concave on lower half, with one tubercle on each side near upper eye margin, female with frons less impressed, lateral tubercles weakly developed and fine erect short setae over entire frons; elytra near glabrous, with very fine short setae; interstriae slightly elevated from disc to apex, interstriae 3 more strongly raised from posterior disc to apex having a distinct higher tubercle in the middle of declivity.</p><p>Distribution: Cameroon, Democratic Republic of the Congo, Republic of Congo, Gabon.</p><p>Host plant: Staudtia stipitata ( Myristicaceae).</p><p>Remarks. This species is easily recognized by its large size and tubercles on the declivity, and by the tuberculate impressed male frons. Species status was verified by material identified by Eggers and Schedl and compared to the original description. A neotype is therefore not needed.</p></div>	https://treatment.plazi.org/id/2445B91BC164FFEBE89CFAD765EB7497	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jordal, Bjarte H.	Jordal, Bjarte H. (2025): Taxonomic revision of the rare Afrotropical genus Chortastus Schaufuss (Coleoptera: Scolytinae). Zootaxa 5647 (1): 77-84, DOI: 10.11646/zootaxa.5647.1.5, URL: https://doi.org/10.11646/zootaxa.5647.1.5
2445B91BC164FFEBE89CFF7566F277FD.text	2445B91BC164FFEBE89CFF7566F277FD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chortastus Schaufuss 1905	<div><p>Chortastus Schaufuss, 1905</p><p>Afrochramesus Schedl, 1971, synonym by Wood, 1984</p><p>Type species: Chortastus camerunus Schaufuss, 1905</p><p>Diagnosis. Antennal club asymmetrically attached to the funicle, flattened, with an oblique partial septum, sutures faint or distinctly marked, asymmetrically procurved; funiculus 6-segmented; eyes deeply and broadly emarginated on upper third of its anterior margin; labial palp segment 1 as long as 2 and 3 combined, ligula not developed; maxillary palp segments of similar length, setae along the lacinial edge long and thin; pronotum smooth, distinctly constricted on its anterior third; basal margin of elytra crenulated, recurved in middle, scutellar shield not visible; tibiae with 3–5 short denticles at the apical margin, apex in meso- and and particularly the metatibiae forming a corbel-like structure; proventriculus with anterior plate as long as posterior part, with transverse rows of small teeth, median suture not present.</p><p>Phylogeny. Browne (1973) suggested that this genus is closely related to Phloeosinus Chapuis having fairly similar antennae, tibiae and proventriculus. He also remarked that it shares some similarities of the tibiae and antennae with Phrixosoma Blandford. Other authors place the genus in Polygraphini based on the invisible scutellar shield, smooth pronotum and flattened antennal club (e.g. Wood 1986). In recent molecular phylogenetic studies (Jordal &amp; Cognato 2012; Pistone et al. 2018), however, Chortastus verrucosus sp. nov. (named as Chortastus medius, PoCho 01) did not group closely with any of these taxa, or any other genera, and the status of the genus therefore remains enigmatic.</p><p>Biology. Sexual dimorphism is striking with females having a simple unmodified frons and males having either a tuft of long setae or concavities with lateral tubercles. Dissection of C. verrucosus sp. nov. revealed a female with spermatheca, which had a simple frons, and a tuft of setae attached to the gula on the underside of the head. The gular tuft is also seen in females of C. orientalis and is a character of uncertain function that appears in a handful of species across the entire subfamily.</p><p>Schedl (1959) described the reproductive biology in two species breeding under the bark of Staudtia trunks and large branches ( Myristicaceae). Monogamous pairs were formed in which the female made primarily two longitudinal egg tunnels away from a central mating niche. Based on illustrations the brood sizes could be more than 100 larvae in one family.</p><p>Key to species</p><p>1. Large species,&gt; 5 mm long; most striae distinctly impressed, interstriae 3 on declivity each with a raised ridge that midway contain a median pointed tubercle; elytra nearly glabrous, vestiture consisting of short, fine, recumbent setae (Congo Basin)................................................................................... Chortastus camerunus</p><p>- Body length &lt;3.5 mm; striae on disc not impressed; elytral vestiture of erect bristle-like setae........................ 2</p><p>2. Elytral declivity with a prominent ridge along interstriae 3 (expanding towards 4); strial punctures large and deep, broader than width of interstriae (Congo Basin)...................................................... Chortastus schenklingi</p><p>- Elytral interstriae 3 on the declivity only slightly if at all elevated more than other interstriae; strial punctures narrower than interstriae........................................................................................... 3</p><p>3. Elytral declivity with interstriae 2 impressed and interstriae 3 slightly elevated, appearing sulcate; body length &lt;2.8 mm ... 4</p><p>- Declivity with uniform interstriae; body length&gt; 3.2 mm (Tanzania)............................. Chortastus orientalis</p><p>4. Elytral interstriae on declivity with densely placed, sharp granules, all except interstriae 2 raised; female head on its underside with tuft of forward pointed setae (Cameroon)...................................... Chortastus verrucosus sp. nov.</p><p>- Elytra with interstriae smooth throughout, only interstriae 3 raised (Congo Basin)................... Chortastus minimus</p></div>	https://treatment.plazi.org/id/2445B91BC164FFEBE89CFF7566F277FD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jordal, Bjarte H.	Jordal, Bjarte H. (2025): Taxonomic revision of the rare Afrotropical genus Chortastus Schaufuss (Coleoptera: Scolytinae). Zootaxa 5647 (1): 77-84, DOI: 10.11646/zootaxa.5647.1.5, URL: https://doi.org/10.11646/zootaxa.5647.1.5
2445B91BC167FFE9E89CFA0C627373B4.text	2445B91BC167FFE9E89CFA0C627373B4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chortastus orientalis Schedl 1957	<div><p>Chortastus orientalis Schedl, 1957</p><p>(Figs 5–8)</p><p>Chortastus orientalis Schedl, 1957: 868 .</p><p>Type material. Holotype and paratypes: Tanzania, Lushoto, ex Ocotea usambarensis, 8.IX.1955 J. C. M. Gardner, leg. [NHMUK], additional paratypes: same data [NHMW], same data except 5.XII.1955 [NHMUK] .</p><p>Diagnosis. Body length 3.3–3.4 mm, 2.0–2.1 × as long as wide. Female frons simple, finely reticulate and dull from vertex to epistoma, with short erect setae evenly distributed; underside of head (gular area) with a tuft of forward pointing setae; antennal club with two clearly visible, asymmetrically procurved, almost angular, sutures; vestiture on pronotum and elytra similar; elytral interstriae on declivity increasingly granular towards posterior margin.</p><p>Distribution: Tanzania.</p><p>Host plant: Kuloa usambarensis ( Lauraceae).</p><p>Remarks. This species is only known from the type locality and is the only East-African representative of the genus. Specimens may be all females given the structure of the frons, and the presence of a gular tuft of setae.</p></div>	https://treatment.plazi.org/id/2445B91BC167FFE9E89CFA0C627373B4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jordal, Bjarte H.	Jordal, Bjarte H. (2025): Taxonomic revision of the rare Afrotropical genus Chortastus Schaufuss (Coleoptera: Scolytinae). Zootaxa 5647 (1): 77-84, DOI: 10.11646/zootaxa.5647.1.5, URL: https://doi.org/10.11646/zootaxa.5647.1.5
2445B91BC166FFE9E89CFEC466297638.text	2445B91BC166FFE9E89CFEC466297638.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chortastus minimus Hagedorn 1909	<div><p>Chortastus minimus Hagedorn, 1909</p><p>(Figs 9–18)</p><p>Chortastus minimus Hagedorn, 1909: 738 .</p><p>Chortastus medius Eggers, 1924: 100, syn. nov.</p><p>Chortastus agnatus Eggers, 1935: 301, synonym by Browne, 1973.</p><p>Chortastus sulcatus Eggers, 1935: 301, synonym by Browne, 1973</p><p>Chortastus brunneus Nunberg, 1967: 317, synonym of C. medius by Browne, 1973.</p><p>Afrochramesus baguenai Schedl, 1971: 197, synonym of C. medius by Beaver, 1998.</p><p>Type material. Syntypes: Cameroon [ZMHB]. Syntype of Chortastus medius: Congostaat [Democratic Republic of the Congo], Albertville [Kalemie] 1.XII.1918, R. Mayné, leg. [MRCA] . Lectotype of Chortastus agnatus: Cameroon, Conradt, leg. [USNM] . Holotype of Chortastus sulcatus: [Democratic Republic of the Congo], Sankuru, 1.IV.1925, ex Staudtia gabonensis, Lt. Ghesquière, leg. [MRCA] . Holoty,pe of Chortastus brunneus: [Democratic Republic of the Congo], Bas-Congo, Kimwenza, 1.IV.1956, R. P. Van Eyen, leg. [MRCA].</p><p>Other material. Congo-Brazzaville [Republic of Congo], Sibiti, leg. Endrödy-Younga, 27.11.1963 [NHMW]. Democratic Republic of the Congo, Mulongo, [Haut-Lomami], 20.5.1930, P. Gerard, leg. [NHMW]; Yangambi, 14.12.1951, K. E. Schedl, leg. [NHMW] and 17.VII.1952 [MRCA] .</p><p>Diagnosis. Body length 1.6–2.7 mm, 2.1–2.2 × as long as wide. Male frons with long golden setae from vertex and near eye margin reaching near epistoma, female frons with very short erect setae evenly distributed over entire frons; pronotum and elytra with similar type of setae, on elytral interstriae in a single row or in variably confused multiple rows; interstriae 2 on declivity slightly impressed, smooth and glabrous, interstriae 3 on lower half of declivity with a slightly elevated blunt ridge.</p><p>Distribution. Democratic Republic of the Congo, Republic of Congo, Cameroon, Angola.</p><p>Host plant: Staudtia stipitata ( Myristicaceae).</p><p>Remarks. Comparison of (holo-) types of C. minimus, C. medius, C. sulcatus, C. brunneus and C. agnatus, and a paratype of C. baguenai revealed a gradual variation from smallest ( minimus, agnatus and sulcatus) to medium ( baguenai) to largest ( medius), and from slightly more spaced setae ( minimus and agnatus) to a little denser ( sulcatus, brunneus and medius). Because all these species were described from nearby locations in the greater Congo basin, and that body size is known to vary a great deal in phloem feeding bark beetles, these intergrading types are inferred as conspecific.</p></div>	https://treatment.plazi.org/id/2445B91BC166FFE9E89CFEC466297638	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jordal, Bjarte H.	Jordal, Bjarte H. (2025): Taxonomic revision of the rare Afrotropical genus Chortastus Schaufuss (Coleoptera: Scolytinae). Zootaxa 5647 (1): 77-84, DOI: 10.11646/zootaxa.5647.1.5, URL: https://doi.org/10.11646/zootaxa.5647.1.5
2445B91BC166FFE9E89CFA48665A74D6.text	2445B91BC166FFE9E89CFA48665A74D6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chortastus schenklingi Hagedorn 1909	<div><p>Chortastus schenklingi Hagedorn, 1909</p><p>(Figs 19–22)</p><p>Chortastus schenklingi Hagedorn, 1909: 737 .</p><p>Type material. Syntypes: Kamerun [ZMHB]</p><p>Other material. Cameroon, Niombe, 1912, Herre leg. [NHMW]; Democratic Republic of the Congo, Sanzulu, 4.IV.1926, A. Collart, leg. [RBINS]; Binga, -Kondo [Zimbabwe border], 15.VI.1926 [RBINS], new country .</p><p>Diagnosis. Body length 2.7–3.1 mm, 2.0–2.1 × as long as wide. Male frons with a long tuft of golden setae from vertex and anterior margin of eyes to near epistoma; female frons flat with short erect setae; pronotum and elytra with similar type of short erect setae; interstriae 2 on declivity slightly impressed and smooth, interstriae 3 with an elevated ridge as high as the width of a femur.</p><p>Distribution: Nigeria, Cameroon, Gabon, Equatorial Guinea, Democratic Republic of the Congo.</p><p>Remarks. Despite the many records from five African countries, a host plant has not yet been noted. A first record for the Democratic Republic of the Congo is given. This species is unmistaken by the long ridges on the elytral declivity.</p></div>	https://treatment.plazi.org/id/2445B91BC166FFE9E89CFA48665A74D6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jordal, Bjarte H.	Jordal, Bjarte H. (2025): Taxonomic revision of the rare Afrotropical genus Chortastus Schaufuss (Coleoptera: Scolytinae). Zootaxa 5647 (1): 77-84, DOI: 10.11646/zootaxa.5647.1.5, URL: https://doi.org/10.11646/zootaxa.5647.1.5
2445B91BC160FFECE89CF92765D27074.text	2445B91BC160FFECE89CF92765D27074.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chortastus verrucosus Jordal 2025	<div><p>Chortastus verrucosus, sp. nov.</p><p>urn:lsid:zoobank.org:act: A0C654AC-BA37-4D05-A15D-045F469EDDC2</p><p>(Figs 23–25)</p><p>Type material. Holotype, female: Cameroon, SW Province, Limbe, Ekonjo (900 m), ex Caloncoba sp., 17. Nov. 2007, B. Jordal leg. [ZMUB].</p><p>Diagnosis, female. Antennal club with two asymmetrically procurved sutures; interstriae on elytral declivity slightly raised and strongly tuberculated; female with gular tuft of setae.</p><p>Description, female. Body length 2.6 mm, 2.2 × as long as wide, colour dark brown. Frons convex, shiny, finely punctured, with short erect setae evenly distributed; underside of head (gular area) with a tuft of forward pointing setae; antenna light yellow, club with two clearly visible, asymmetrically procurved sutures, suture 1 with a partial septum. Pronotum finely punctured, shiny; vestiture of mainly short spatulate setae. Elytra with weakly impressed striae becoming more strongly impressed near and at the declivity; interstriae increasingly raised posteriorly with distinct granules on the declivity; vestiture consisting of confused short and spatulate interstrial setae. Legs of the same colour as remaining body, tarsal segment 3 distinctly lobed; all tibiae with transversely set apical denticles only, protibiae with apical corner posteriorly curved, in mesotibiae anteriorly pointed; metatibial apex with a distinct false corbel.</p><p>Etymology. A Latin nominative adjective meaning warty or rough, referring to the granulated interstriae on the posterior half of the elytra.</p><p>Remarks. This species is closely related to C. minimus but differs clearly by the raised granular interstriae on the elytral declivity, and the gular tuft of pointed setae in the female. The only other species with a gular tuft is C. orientalis which also has two visible sutures in the antennal club, albeit more strongly angulate. The two species differ further in the structure of the elytral interstriae, body size, and distribution.</p><p>A single female specimen was collected from the bark of a dead branch of Caloncoba ( Achariaceae) in Cameroon, near Limbe. DNA sequence data are deposited in GenBank (as C. medius) with accession numbers present for ten different gene fragments (see Pistone et al. 2018).</p></div>	https://treatment.plazi.org/id/2445B91BC160FFECE89CF92765D27074	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jordal, Bjarte H.	Jordal, Bjarte H. (2025): Taxonomic revision of the rare Afrotropical genus Chortastus Schaufuss (Coleoptera: Scolytinae). Zootaxa 5647 (1): 77-84, DOI: 10.11646/zootaxa.5647.1.5, URL: https://doi.org/10.11646/zootaxa.5647.1.5
