taxonID	type	description	language	source
63E9E7D332BE5983A0C696635E26470F.taxon	diagnosis	Diagnosis. Body small to medium-sized (~ 9 – 25 mm long, ~ 0.7 – 3.3 mm wide), with 20 rings. Paraterga poorly to moderately developed. Sternal lobe or cone (s) present between ♂ coxae 4. First pair of ♂ without femoral tubercles (= adenostyles). Transverse metatergal sulci distinct. Legs with neither modifications nor adenostyles. Gonopod with a long and subcylindrical coxite, slightly curved caudally; prefemoral part (= prefemorite) short, ~ 3 – 4 × shorter than acropodite; femoral part (= femorite) strongly reduced or very short compared to solenophore, solenomere mostly rod-shaped or subflagelliform, sheathed distally by solenophore; both lamina medialis and lamina lateralis well developed; femoral process (fp) present or absent.	en	Likhitrakarn, Natdanai, Golovatch, Sergei I., Inkhavilay, Khamla, Panha, Somsak, Sutcharit, Chirasak (2025): Taxonomic review of the millipede genus Touranella Attems, 1937, with a redescription of the type species, T. gracilis Attems, 1937, and descriptions of three new species from Laos (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 1238: 183-208, DOI: 10.3897/zookeys.1238.147550
63E9E7D332BE5983A0C696635E26470F.taxon	type_taxon	Type species. Touranella gracilis Attems, 1937, by original designation.	en	Likhitrakarn, Natdanai, Golovatch, Sergei I., Inkhavilay, Khamla, Panha, Somsak, Sutcharit, Chirasak (2025): Taxonomic review of the millipede genus Touranella Attems, 1937, with a redescription of the type species, T. gracilis Attems, 1937, and descriptions of three new species from Laos (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 1238: 183-208, DOI: 10.3897/zookeys.1238.147550
97B290B7A1395D3EBDE6D5AA21F7EFF7.taxon	description	Figs 4, 5, 6	en	Likhitrakarn, Natdanai, Golovatch, Sergei I., Inkhavilay, Khamla, Panha, Somsak, Sutcharit, Chirasak (2025): Taxonomic review of the millipede genus Touranella Attems, 1937, with a redescription of the type species, T. gracilis Attems, 1937, and descriptions of three new species from Laos (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 1238: 183-208, DOI: 10.3897/zookeys.1238.147550
97B290B7A1395D3EBDE6D5AA21F7EFF7.taxon	diagnosis	Diagnosis. The new species seems to be particularly similar to T. champasak from Bolaven Plateau, Laos, especially in gonopodal structure. Both species share a strongly reduced gonopodal femorite devoid of a femoral process and a distally slightly twisted solenophore (Figs 5, 6). However, the new species differs from T. champasak by having a long, acute process d located around the mid-length of a more clearly erect and thicker solenophore (Figs 5, 6). Additionally, pleurosternal carinae are complete crests with sharp caudal denticles extending beyond the rear tergal margin only on rings 2 and 3, reduced to a rounded caudal crest on rings 4 and 5, and absent thereafter (Fig. 4 C, E) (vs absent on rings 18 – 19). Moreover, ♂ tarsal brushes are present until ring 8 (vs until ring 16).	en	Likhitrakarn, Natdanai, Golovatch, Sergei I., Inkhavilay, Khamla, Panha, Somsak, Sutcharit, Chirasak (2025): Taxonomic review of the millipede genus Touranella Attems, 1937, with a redescription of the type species, T. gracilis Attems, 1937, and descriptions of three new species from Laos (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 1238: 183-208, DOI: 10.3897/zookeys.1238.147550
97B290B7A1395D3EBDE6D5AA21F7EFF7.taxon	description	Description. Length 15.2 mm, width of midbody pro- and metazona 1.1 and 1.3 mm, respectively (♂). Coloration of live animals dark brown (Fig. 4 A); legs pale orange, venter and a few basal podomeres pale brown to yellow brown. Coloration in alcohol after 11 years of preservation faded to pale brown; antennae and epiproct pale brown to pallid, venter and podomeres pallid (Fig. 4 B – J). Clypeolabral region and vertex sparsely setose, epicranial suture distinct. Antennae long (Fig. 4 A), reaching until ring 5 when stretched dorsally. In width, ring 4 <3 <2 <collum <ring 5 <6 <7 <head <8 – 17, thereafter body gently and gradually tapering towards telson. Collum with three transverse rows of setae: 3 + 3 anterior, 2 + 2 intermediate, and 3 + 3 posterior; caudal corner very narrowly rounded, slightly declined ventrally, not produced past rear tergal margin (Fig. 4 A – C). Tegument smooth and shining, prozona finely shagreened, metaterga nearly smooth, faintly rugulose and leathery (Fig. 4 B – F, H). Postcollum metaterga each with two transverse rows of setae: 2 + 2 anterior and 2 + 2 posterior, traceable at least as insertion points when setae broken off. Tergal setae simple, slender, short, ~ 1 / 3 metatergal length (Fig. 4 F). Axial line well visible on metazona, traceable also on prozona. Paraterga weak (Fig. 4 A – F, H), lying at ~ 1 / 2 midbody height, slightly upturned, anterior edge rounded and narrowly bordered; caudal corner very narrowly rounded on rings 2 – 4, not produced past rear tergal margin; almost missing on following rings; pore-bearing rings with evident lateral bulges (Fig. 4 B, D). Ozopores evident, lateral, each lying in an ovoid groove at ~ 1 / 3 metatergal length in front of posterior edge of metaterga (Fig. 4 C, E, H). Transverse metatergal sulci complete on rings 5 – 16, incomplete on ring 17, very narrow, shallow, not reaching the bases of paraterga, at most faintly beaded at bottom (Fig. 4 B – F, H). Stricture between pro- and metazona wide, deep, clearly ribbed at bottom down to base of paraterga (Fig. 4 B – F, H). Pleurosternal carinae complete crests with a very sharp median tooth on rings 2 and 3, reduced to a rounded caudal crest on rings 4 and 5, thereafter missing (Fig. 4 C, E, H). Epiproct (Fig. 4 F – H) conical, flattened dorsoventrally, with two small, rounded, apical papillae; tip subtruncate; lateral pre-apical papillae very small, lying close to tip. Hypoproct roundly subtriangular, setigerous knobs at caudal edge small and well-separated (Fig. 4 G). Sterna sparsely setose, without modifications; an entire, high, rounded, linguiform, setose sternal lobe between ♂ coxae 4 (Fig. 4 I, J). Legs long, midbody legs 1.2 – 1.4 × as long as body height, prefemora without modifications, ♂ tarsal brushes present until ring 8. Gonopods simple, slim and suberect (Figs 5, 6). Coxite slightly curved caudad, densely setose distodorsally (Figs 5 A, B, 6 A, B, D). Prefemoral part (= prefemorite, pfe) as usual, densely setose, ~ 3.5 × shorter than acropodite (femoral + postfemoral parts) (Figs 5 A, B, 6 A – C). Femorite (fe) short, with a strong, long, flagelliform solenomere (sl) twisted distad and with a strong, oblique, lateral sulcus demarcating a postfemoral part (Figs 5 A, B, 6 A – C). Solenophore (sph) long, slightly twisted distad, sheathing most of solenomere (sl), slightly curved caudad (Figs 5 A, B, 6 A – D). Solenophore consisting of a well-developed lamina lateralis (ll) and a smaller lamina medialis (lm) (Figs 5 B, 6 B, C). Tip of lamina medialis subtruncate (Figs 5 B, 6 B), tip of lamina lateralis subrectangular with a long, slender, curved, nearly pointed process d (d), this rising distal to lamina lateralis (Figs 5 A, B, 6 A – D). Both supporting a long and flagelliform solenomere.	en	Likhitrakarn, Natdanai, Golovatch, Sergei I., Inkhavilay, Khamla, Panha, Somsak, Sutcharit, Chirasak (2025): Taxonomic review of the millipede genus Touranella Attems, 1937, with a redescription of the type species, T. gracilis Attems, 1937, and descriptions of three new species from Laos (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 1238: 183-208, DOI: 10.3897/zookeys.1238.147550
97B290B7A1395D3EBDE6D5AA21F7EFF7.taxon	distribution	Distribution. Known only from the type locality, apparently endemic to the southern part of Laos.	en	Likhitrakarn, Natdanai, Golovatch, Sergei I., Inkhavilay, Khamla, Panha, Somsak, Sutcharit, Chirasak (2025): Taxonomic review of the millipede genus Touranella Attems, 1937, with a redescription of the type species, T. gracilis Attems, 1937, and descriptions of three new species from Laos (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 1238: 183-208, DOI: 10.3897/zookeys.1238.147550
97B290B7A1395D3EBDE6D5AA21F7EFF7.taxon	etymology	Etymology. The species epithet chenla refers to the ancient kingdom of Chenla, a powerful kingdom that thrived from approximately 550 to 802 AD across the present-day territories of Thailand, Laos, Cambodia, and Vietnam. The naming also reflects the historical connection between the Chenla era and the construction of the Vat Phou temple complex in Champasak Province, Laos, where the species was discovered (Wikipedia 2025 a, b). This new species so named pays tribute to the rich cultural and historical heritage of the region.	en	Likhitrakarn, Natdanai, Golovatch, Sergei I., Inkhavilay, Khamla, Panha, Somsak, Sutcharit, Chirasak (2025): Taxonomic review of the millipede genus Touranella Attems, 1937, with a redescription of the type species, T. gracilis Attems, 1937, and descriptions of three new species from Laos (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 1238: 183-208, DOI: 10.3897/zookeys.1238.147550
AF25E3555C7B5FBDAFF73435864ECA48.taxon	description	Figs 2, 3	en	Likhitrakarn, Natdanai, Golovatch, Sergei I., Inkhavilay, Khamla, Panha, Somsak, Sutcharit, Chirasak (2025): Taxonomic review of the millipede genus Touranella Attems, 1937, with a redescription of the type species, T. gracilis Attems, 1937, and descriptions of three new species from Laos (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 1238: 183-208, DOI: 10.3897/zookeys.1238.147550
AF25E3555C7B5FBDAFF73435864ECA48.taxon	distribution	Distribution. It is known only from the type locality and appears to be endemic to Vietnam.	en	Likhitrakarn, Natdanai, Golovatch, Sergei I., Inkhavilay, Khamla, Panha, Somsak, Sutcharit, Chirasak (2025): Taxonomic review of the millipede genus Touranella Attems, 1937, with a redescription of the type species, T. gracilis Attems, 1937, and descriptions of three new species from Laos (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 1238: 183-208, DOI: 10.3897/zookeys.1238.147550
2763A5D63EF95FD1A24F10D263463331.taxon	description	Figs 10, 11, 12	en	Likhitrakarn, Natdanai, Golovatch, Sergei I., Inkhavilay, Khamla, Panha, Somsak, Sutcharit, Chirasak (2025): Taxonomic review of the millipede genus Touranella Attems, 1937, with a redescription of the type species, T. gracilis Attems, 1937, and descriptions of three new species from Laos (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 1238: 183-208, DOI: 10.3897/zookeys.1238.147550
2763A5D63EF95FD1A24F10D263463331.taxon	diagnosis	Diagnosis. The new species closely resembles both T. chenla sp. nov. and T. champasak, especially in its moniliform body with significantly reduced paraterga (Fig. 4 A – H, 7 A – H). However, it differs from T. chenla sp. nov. by possessing a broader and better expanded apical process d, which arises proximal to the lamina lateralis (Figs 11, 12) (vs a slender, long, curved, apical process d that rises distal to the lamina lateralis (Figs 5, 6 )), coupled with a larger size, measuring 21.4 – 22.6 mm long (vs smaller, 15.2 mm long). In comparison to T. champasak, this species has pleurosternal carinae with complete crests and sharp caudal teeth on rings 2 – 4, and absent from ring 5 (vs present until ring 17 and absent thereafter). ♂ legs are 1.3 – 1.5 × as long as midbody height (vs longer, 1.7 – 1.8 ×), and ♂ tarsal brushes present until ring 10 (vs until ring 16).	en	Likhitrakarn, Natdanai, Golovatch, Sergei I., Inkhavilay, Khamla, Panha, Somsak, Sutcharit, Chirasak (2025): Taxonomic review of the millipede genus Touranella Attems, 1937, with a redescription of the type species, T. gracilis Attems, 1937, and descriptions of three new species from Laos (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 1238: 183-208, DOI: 10.3897/zookeys.1238.147550
2763A5D63EF95FD1A24F10D263463331.taxon	description	Description. Length 21.4 – 22.6 (♂), width of midbody pro- and metazona 2.6 – 2.8 and 3.2 – 3.3 mm (♂), respectively. Coloration of live animals dark brown-orange (Fig. 10 A), with a contrasting longitudinal pale yellow-orange stripe, this being narrow on prozona, but characteristically bead-shaped and broadened on metazona; paraterga, epiproct, and venter pale orange to pale yellow; head and antennae dark brown; legs contrasting pale orange, a few basal podomeres pale yellow to pallid. Coloration in alcohol after eight years of preservation faded to dark brown; antennae dark brown and increasingly faded distally; paraterga, venter, and epiproct pale brown to pallid; and podomeres pallid, basal podomeres of legs pallid, increasingly dark brown distally (Fig. 10 B – J). Clypeolabral region and vertex sparsely setose, epicranial suture distinct. Antennae long, reaching until ring 5 when stretched dorsally (Fig. 10 A). In width, ring 3 = 4 <collum <ring 2 <5 <6 – 17 (♂); thereafter body gently and gradually tapering towards telson. Collum with three transverse rows of setae: 3 + 3 anterior, 2 + 2 intermediate, and 4 + 4 posterior; caudal corner very narrowly rounded, paraterga slightly declined ventrally, not produced past rear tergal margin (Fig. 10 A, B). Tegument smooth and shining, prozona finely shagreened, metaterga nearly smooth, faintly rugulose and leathery (Fig. 10 B – F, H) Postcollum metaterga each with two transverse rows of setae: 2 + 2 in anterior and 3 + 3 in posterior row, the latter traceable as insertion points when setae broken off. Tergal setae simple, short, slender, ~ 1 / 5 metatergal length. Axial line visible on metazona, traceable also on prozona. Paraterga weak (Fig. 10 A – F, H), slightly upturned posteriorly, lying at ~ 1 / 2 midbody height, lateral edge without incisions. Paraterga 2 subhorizontal, broad in dorsal view, thin in lateral view, anterior edge well-developed, slightly rounded and oblique laterally, caudal tip rounded, slightly produced past rear tergal margin (Fig. 10 B, C). Paraterga 3 and 4 with evident lateral bulges, caudal corner very broadly rounded, not produced past rear tergal margin (Fig. 10 B, C). Following rings with small lateral bulges, not produced past rear tergal margin, broader on pore-bearing rings (Fig. 10 B, D, F). Calluses on paraterga delimited by a sulcus both dorsally and ventrally. Ozopores evident, lateral, each lying in an ovoid groove at ~ 1 / 3 metatergal length in front of posterior edge of metaterga (Fig. 10 E, H). Transverse metatergal sulci usually distinct (Fig. 10 B – F, H), incomplete on ring 17, complete on rings 5 – 16, shallow, not reaching the bases of paraterga, at most faintly beaded at bottom. Stricture between pro- and metazona wide and rather deep, ribbed at bottom down to base of paraterga (Fig. 10 B – E). Pleurosternal carinae complete crests with sharp caudal teeth on rings 2 – 4, thereafter missing (Fig. 10 C, E, H). Epiproct (Fig. 10 F – H) conical, flattened dorsoventrally, tip subtruncate, subapical lateral papillae small, but visible, lying close to tip. Hypoproct roundly subtrapeziform, small setigerous knobs at caudal edge small and well-separated (Fig. 10 G). Sterna sparsely setose, without modifications (Fig. 10 G); an entire, high, rounded, linguiform, setose, sternal lobe between ♂ coxae 4 (Fig. 10 I, J). A paramedian pair of evident tubercles in front of gonopodal aperture. Legs long and slender, midbody ones ~ 1.3 – 1.5 as long as body height; prefemora without modifications; ♂ tarsal brushes present until ring 10 (♂). Gonopods relatively simple and suberect (Figs 11, 12). Coxite slightly curved caudally, rather densely setose distodorsally (Fig. 11 A, B). Prefemoral part (pfe) densely setose, ~ 1 / 4 as long as acropodite (femoral + postfemoral parts) (Fig. 11 A – C). Femorite (fe) strongly reduced, without femoral processes, bearing a medial, strong, long, flagelliform solenomere (sl), twisted distally, and with an oblique lateral sulcus defining a postfemoral part (Fig. 11 A, B). Solenophore (sph) long, rather slender, slightly curved caudally, sheathing most of solenomere (sl), directed caudally, with a lateral shoulder (u) at base (Figs 11 C, D, 12 A, D, E). Solenophore consisting of a well-developed, strongly twisted lamina lateralis (ll) and a smaller, slightly twisted lamina medialis (lm) (Fig. 11 A – D). Tip of the lamina lateralis (ll) split into two broad, expanded apical laminae, a median lamina with a small denticle, and halfway bearing a broad, expanded apical lamina (Figs 11 A, D, 12 C – F), subtruncate tip process d rising proximal to lamina lateralis (Fig. 11 A, B).	en	Likhitrakarn, Natdanai, Golovatch, Sergei I., Inkhavilay, Khamla, Panha, Somsak, Sutcharit, Chirasak (2025): Taxonomic review of the millipede genus Touranella Attems, 1937, with a redescription of the type species, T. gracilis Attems, 1937, and descriptions of three new species from Laos (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 1238: 183-208, DOI: 10.3897/zookeys.1238.147550
2763A5D63EF95FD1A24F10D263463331.taxon	distribution	Distribution. Known only from the type locality, apparently endemic to the central part of Laos.	en	Likhitrakarn, Natdanai, Golovatch, Sergei I., Inkhavilay, Khamla, Panha, Somsak, Sutcharit, Chirasak (2025): Taxonomic review of the millipede genus Touranella Attems, 1937, with a redescription of the type species, T. gracilis Attems, 1937, and descriptions of three new species from Laos (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 1238: 183-208, DOI: 10.3897/zookeys.1238.147550
2763A5D63EF95FD1A24F10D263463331.taxon	etymology	Etymology. To honor Dr. Peter Jäger, a renowned arachnologist from the Senckenberg Research Institute in Frankfurt, Germany, who collected the type specimens of this species.	en	Likhitrakarn, Natdanai, Golovatch, Sergei I., Inkhavilay, Khamla, Panha, Somsak, Sutcharit, Chirasak (2025): Taxonomic review of the millipede genus Touranella Attems, 1937, with a redescription of the type species, T. gracilis Attems, 1937, and descriptions of three new species from Laos (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 1238: 183-208, DOI: 10.3897/zookeys.1238.147550
6075D95BDFBC598ABF27C5A3BAC7785F.taxon	description	Figs 7, 8, 9	en	Likhitrakarn, Natdanai, Golovatch, Sergei I., Inkhavilay, Khamla, Panha, Somsak, Sutcharit, Chirasak (2025): Taxonomic review of the millipede genus Touranella Attems, 1937, with a redescription of the type species, T. gracilis Attems, 1937, and descriptions of three new species from Laos (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 1238: 183-208, DOI: 10.3897/zookeys.1238.147550
6075D95BDFBC598ABF27C5A3BAC7785F.taxon	diagnosis	Diagnosis. Morphologically, in gonopodal structure this new species seems to be especially similar to both T. chenla sp. nov. and T. champasak, but it is distinguished by the uniform black coloration lacking a cingulate pattern (Fig. 7 A – H). It differs from T. champasak by having process d at the mid-length of the solenophore (Figs 8, 9) (vs absent); a more erect and thicker solenophore (Figs 8, 9) (vs slender and curved). T. srisonchaii sp. nov. differs from T. chenla sp. nov. in having ♂ tarsal brushes present until ring 8 (vs until ring 14) and a twisted solenophore with a laminate and subtruncate process d (Figs 8, 9) (vs a suberect gonopodal solenophore with a slender and acute process d, Figs 5, 6).	en	Likhitrakarn, Natdanai, Golovatch, Sergei I., Inkhavilay, Khamla, Panha, Somsak, Sutcharit, Chirasak (2025): Taxonomic review of the millipede genus Touranella Attems, 1937, with a redescription of the type species, T. gracilis Attems, 1937, and descriptions of three new species from Laos (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 1238: 183-208, DOI: 10.3897/zookeys.1238.147550
6075D95BDFBC598ABF27C5A3BAC7785F.taxon	description	Description. Length 14.8 – 21.0 (♂) or 14.6 – 19.8 mm (♀), width of midbody pro- and metazona 1.5 – 2.3 and 1.8 – 2.2 mm (♂) or 1.8 – 2.4 and 2.2 – 2.6 mm (♀), respectively. Coloration of live animals mostly dark or blackish (Fig. 7 A); head, antennae, paraterga, and epiproct slightly lighter; venter dark brown, legs pale yellowish. Coloration in alcohol after seven years of preservation faded to dark brown or blackish; head, antennae, paraterga, and epiproct dark brown to pale brown; venter pale brown; legs pale yellowish to pallid (Fig. 7 B – J). Clypeolabral region and vertex sparsely setose, epicranial suture distinct. Antennae moderately long (Fig. 7 A, B), reaching until body ring 4 (♂, ♀) when stretched dorsally. Head in width <collum <ring 3 <4 <2 <5 – 16 (♂, ♀); thereafter body gently and gradually tapering towards telson. Collum with three transverse rows of setae: 3 + 3 anterior, 2 + 2 intermediate, and 4 + 4 posterior; caudal corner very narrowly rounded, slightly declined ventrally, not produced past rear tergal margin (Fig. 7 B, C). Tegument smooth and shining, prozona finely shagreened, metaterga finely rugulose (Fig. 7 A – F, H); surface below paraterga finely microgranulate (Fig. 7 C, E, F). Postcollum metaterga each with two transverse rows of setae: 3 + 3 anterior and 4 + 4 posterior, nearly always abraded, but still traceable as insertion points. Tergal setae simple, slender, short, ~ 1 / 5 metatergal length. Axial line well visible on metazona, traceable also on prozona. Paraterga moderately developed (Fig. 7 A – F, H), slightly upturned, lying at ~ 1 / 2 midbody height. Paraterga 2 subhorizontal, broad in dorsal view, thin in lateral view; shoulders well-developed, slightly rounded and oblique laterally; caudal tip rounded, slightly produced past rear tergal margin (Fig. 7 B, C). Following rings with evident lateral bulges, not produced past rear tergal margin, broader on pore-bearing rings (Fig. 7 B, D, F). Ozopores evident, lateral, each lying in an ovoid groove at ~ 1 / 3 metatergal length in front of posterior edge of metaterga (Fig. 7 C, E, H). Transverse metatergal sulci usually distinct (Fig. 7 B – F, H), incomplete on rings 4 and 19, complete on metaterga 5 – 17 (♂, ♀), shallow, not reaching the bases of paraterga, at most faintly beaded at bottom. Stricture between pro- and metazona narrow, beaded at bottom down to base of paraterga (Fig. 7 B – E). Pleurosternal carinae complete crests with a sharp caudal tooth on ring 2, reduced to a rounded caudal crest on rings 3 and 4, thereafter missing (♂, ♀) (Fig. 7 C, E, H). Epiproct (Fig. 7 F – H) conical, flattened dorsoventrally, with two small, rounded, apical papillae; tip rounded; lateral pre-apical papillae very small, lying close to tip. Hypoproct roundly subtriangular, setigerous knobs at caudal edge small and well-separated (Fig. 7 G). Sterna sparsely setose, without modifications (Fig. 7 G); an entire, rather short, rounded, linguiform, setose, sternal lobe between ♂ coxae 4 (Fig. 7 I, J). A paramedian pair of evident tubercles in front of gonopodal aperture. Legs moderately long and slender, midbody ones ~ 1.1 – 1.3 (♂) or 0.9 – 1.1 × (♀) as long as body height, prefemora without modifications, ♂ tarsal brushes present until ring 14. Gonopods relatively simple and suberect (Figs 8, 9). Coxite slightly curved caudally, rather densely setose distodorsally (Figs 8 A, B, 9 A, B). Prefemoral part (pfe) densely setose, ~ 1 / 3 as long as acropodite (femoral + postfemoral parts) (Figs 8 A, B, 9 A, B). Femorite (fe) rather short, with a medial, strong, long, flagelliform solenomere (sl), strongly twisted distally, and with an oblique lateral sulcus defining a postfemoral part (Figs 8 A, B, 9 A, B). Solenophore (sph) long, stout, suberect, with a clear lateral shoulder, sheathing most of solenomere (sl) (Figs 8 A, B, 9 A – D). Lamina lateralis (ll) well-developed, strongly twisted, and lamina medialis (lm) suberect (Figs 8 B, 9 D, E). Tip of lamina lateralis a broad, expanded, apical lamina with three small denticles (Figs 8 A, B, 9 A – G), at halfway bearing a large, long, slightly curved, subtruncate tip process d, this rising distal to lamina lateralis (Figs 8 A, B, 9 A – G).	en	Likhitrakarn, Natdanai, Golovatch, Sergei I., Inkhavilay, Khamla, Panha, Somsak, Sutcharit, Chirasak (2025): Taxonomic review of the millipede genus Touranella Attems, 1937, with a redescription of the type species, T. gracilis Attems, 1937, and descriptions of three new species from Laos (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 1238: 183-208, DOI: 10.3897/zookeys.1238.147550
6075D95BDFBC598ABF27C5A3BAC7785F.taxon	distribution	Distribution. Known only form the type locality, apparently endemic to the southern part of Laos.	en	Likhitrakarn, Natdanai, Golovatch, Sergei I., Inkhavilay, Khamla, Panha, Somsak, Sutcharit, Chirasak (2025): Taxonomic review of the millipede genus Touranella Attems, 1937, with a redescription of the type species, T. gracilis Attems, 1937, and descriptions of three new species from Laos (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 1238: 183-208, DOI: 10.3897/zookeys.1238.147550
6075D95BDFBC598ABF27C5A3BAC7785F.taxon	etymology	Etymology. To honor Dr. Ruttapon Srisonchai, diplopodologist at the Faculty of Science of Khon Kaen University, who has not only contributed to the study of millipede taxonomy in Thailand, but also collected the type series of this new species.	en	Likhitrakarn, Natdanai, Golovatch, Sergei I., Inkhavilay, Khamla, Panha, Somsak, Sutcharit, Chirasak (2025): Taxonomic review of the millipede genus Touranella Attems, 1937, with a redescription of the type species, T. gracilis Attems, 1937, and descriptions of three new species from Laos (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 1238: 183-208, DOI: 10.3897/zookeys.1238.147550
