taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
35B9C03B44245392A6F69A46B4149A49.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1296554	https://doi.org/10.3897/zookeys.1233.142976.figure1	Figure 1. Collection sites of Dugesia in Hainan Island and Leizhou Peninsula.	Figure 1. Collection sites of Dugesia in Hainan Island and Leizhou Peninsula.	2025-04-01	Wang, Lei;Chang, Yi-Fang;Sun, Xin-Xin;Sluys, Ronald;Liu, De-Zeng;Dong, Zi-Mei;Chen, Guang-Wen		Zenodo	biologists	Wang, Lei;Chang, Yi-Fang;Sun, Xin-Xin;Sluys, Ronald;Liu, De-Zeng;Dong, Zi-Mei;Chen, Guang-Wen			
35B9C03B44245392A6F69A46B4149A49.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1296555	https://doi.org/10.3897/zookeys.1233.142976.figure2	Figure 2. Topology of the molecular phylogenetic tree obtained from Bayesian analysis of the concatenated dataset (dataset III). Numbers at nodes indicate support values (pp / bs). New species indicated by asterisks. Scale bar: substitutions per site.	Figure 2. Topology of the molecular phylogenetic tree obtained from Bayesian analysis of the concatenated dataset (dataset III). Numbers at nodes indicate support values (pp / bs). New species indicated by asterisks. Scale bar: substitutions per site.	2025-04-01	Wang, Lei;Chang, Yi-Fang;Sun, Xin-Xin;Sluys, Ronald;Liu, De-Zeng;Dong, Zi-Mei;Chen, Guang-Wen		Zenodo	biologists	Wang, Lei;Chang, Yi-Fang;Sun, Xin-Xin;Sluys, Ronald;Liu, De-Zeng;Dong, Zi-Mei;Chen, Guang-Wen			
35B9C03B44245392A6F69A46B4149A49.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1296560	https://doi.org/10.3897/zookeys.1233.142976.figure7	Figure 7. Habitat and external appearance of Dugesia aconcinna A, B sampling site and habitat C sexually mature, live individual. Abbreviations: coa: copulatory apparatus; e: eye; ph: pharynx. Scale bar: 2 mm.	Figure 7. Habitat and external appearance of Dugesia aconcinna A, B sampling site and habitat C sexually mature, live individual. Abbreviations: coa: copulatory apparatus; e: eye; ph: pharynx. Scale bar: 2 mm.	2025-04-01	Wang, Lei;Chang, Yi-Fang;Sun, Xin-Xin;Sluys, Ronald;Liu, De-Zeng;Dong, Zi-Mei;Chen, Guang-Wen		Zenodo	biologists	Wang, Lei;Chang, Yi-Fang;Sun, Xin-Xin;Sluys, Ronald;Liu, De-Zeng;Dong, Zi-Mei;Chen, Guang-Wen			
35B9C03B44245392A6F69A46B4149A49.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1296561	https://doi.org/10.3897/zookeys.1233.142976.figure8	Figure 8. Dugesia aconcinna. Photomicrographs of sagittal (B – D), transverse (A, E) and horizontal (F) sections. A Paratype ZMHNU - TPYC 8, showing penis papilla, common atrium, left oviduct and right oviduct B holotype ZMHNU - TPYC 5, showing copulatory bursa, seminal vesicle, vasa deferentia, penis papilla, penis fold, male atrium, common atrium, and genital pore C holotype ZMHNU - TPYC 5, showing copulatory bursa, seminal vesicle, diaphragm, penis papilla, penial fold, male atrium, and common atrium D holotype ZMHNU - TPYC 5, showing copulatory bursa, seminal vesicle, diaphragm, ejaculatory duct, penial fold, male atrium, and common atrium E paratype ZMHNU - TPYC 8, showing penis papilla, ejaculatory duct, penial fold, male atrium, and bursal canal F paratype ZMHNU - TPYC 9, showing copulatory bursa, bursal canal, and penis fold. Abbreviations: bc: bursal canal; ca: common atrium; cb: copulatory bursa; d: diaphragm; ed: ejaculatory duct; go: gonopore; lod: left oviduct; ma: male atrium; pf: penial fold; pp: penis papilla; rod: right oviduct; sv: seminal vesicle; vd: vas deferens. Scale bars: 100 μm.	Figure 8. Dugesia aconcinna. Photomicrographs of sagittal (B – D), transverse (A, E) and horizontal (F) sections. A Paratype ZMHNU - TPYC 8, showing penis papilla, common atrium, left oviduct and right oviduct B holotype ZMHNU - TPYC 5, showing copulatory bursa, seminal vesicle, vasa deferentia, penis papilla, penis fold, male atrium, common atrium, and genital pore C holotype ZMHNU - TPYC 5, showing copulatory bursa, seminal vesicle, diaphragm, penis papilla, penial fold, male atrium, and common atrium D holotype ZMHNU - TPYC 5, showing copulatory bursa, seminal vesicle, diaphragm, ejaculatory duct, penial fold, male atrium, and common atrium E paratype ZMHNU - TPYC 8, showing penis papilla, ejaculatory duct, penial fold, male atrium, and bursal canal F paratype ZMHNU - TPYC 9, showing copulatory bursa, bursal canal, and penis fold. Abbreviations: bc: bursal canal; ca: common atrium; cb: copulatory bursa; d: diaphragm; ed: ejaculatory duct; go: gonopore; lod: left oviduct; ma: male atrium; pf: penial fold; pp: penis papilla; rod: right oviduct; sv: seminal vesicle; vd: vas deferens. Scale bars: 100 μm.	2025-04-01	Wang, Lei;Chang, Yi-Fang;Sun, Xin-Xin;Sluys, Ronald;Liu, De-Zeng;Dong, Zi-Mei;Chen, Guang-Wen		Zenodo	biologists	Wang, Lei;Chang, Yi-Fang;Sun, Xin-Xin;Sluys, Ronald;Liu, De-Zeng;Dong, Zi-Mei;Chen, Guang-Wen			
35B9C03B44245392A6F69A46B4149A49.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1296562	https://doi.org/10.3897/zookeys.1233.142976.figure9	Figure 9. Dugesia aconcinna. Sagittal reconstruction of the copulatory apparatus of holotype TPYC 5. A Female copulatory apparatus B penial fold C male copulatory apparatus. Abbreviations: bc: bursal canal; ca: common atrium; cb: copulatory bursa; cg: cement glands; cpg: cyanophil penial glands; d: diaphragm; ed: ejaculatory duct; epg: extrabulbar penial glands; go: gonopore; lod: left oviduct; ma: male atrium; mo: mouth; pf: penial fold; pp: penis papilla; rod: right oviduct; spv: spermiducal vesicles; sg: shell glands; sv: seminal vesicle; vd: vas deferens. Scale bars: 100 μm.	Figure 9. Dugesia aconcinna. Sagittal reconstruction of the copulatory apparatus of holotype TPYC 5. A Female copulatory apparatus B penial fold C male copulatory apparatus. Abbreviations: bc: bursal canal; ca: common atrium; cb: copulatory bursa; cg: cement glands; cpg: cyanophil penial glands; d: diaphragm; ed: ejaculatory duct; epg: extrabulbar penial glands; go: gonopore; lod: left oviduct; ma: male atrium; mo: mouth; pf: penial fold; pp: penis papilla; rod: right oviduct; spv: spermiducal vesicles; sg: shell glands; sv: seminal vesicle; vd: vas deferens. Scale bars: 100 μm.	2025-04-01	Wang, Lei;Chang, Yi-Fang;Sun, Xin-Xin;Sluys, Ronald;Liu, De-Zeng;Dong, Zi-Mei;Chen, Guang-Wen		Zenodo	biologists	Wang, Lei;Chang, Yi-Fang;Sun, Xin-Xin;Sluys, Ronald;Liu, De-Zeng;Dong, Zi-Mei;Chen, Guang-Wen			
6A0099F70FD45834BE0B3762F9816AF2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1296554	https://doi.org/10.3897/zookeys.1233.142976.figure1	Figure 1. Collection sites of Dugesia in Hainan Island and Leizhou Peninsula.	Figure 1. Collection sites of Dugesia in Hainan Island and Leizhou Peninsula.	2025-04-01	Wang, Lei;Chang, Yi-Fang;Sun, Xin-Xin;Sluys, Ronald;Liu, De-Zeng;Dong, Zi-Mei;Chen, Guang-Wen		Zenodo	biologists	Wang, Lei;Chang, Yi-Fang;Sun, Xin-Xin;Sluys, Ronald;Liu, De-Zeng;Dong, Zi-Mei;Chen, Guang-Wen			
6A0099F70FD45834BE0B3762F9816AF2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1296555	https://doi.org/10.3897/zookeys.1233.142976.figure2	Figure 2. Topology of the molecular phylogenetic tree obtained from Bayesian analysis of the concatenated dataset (dataset III). Numbers at nodes indicate support values (pp / bs). New species indicated by asterisks. Scale bar: substitutions per site.	Figure 2. Topology of the molecular phylogenetic tree obtained from Bayesian analysis of the concatenated dataset (dataset III). Numbers at nodes indicate support values (pp / bs). New species indicated by asterisks. Scale bar: substitutions per site.	2025-04-01	Wang, Lei;Chang, Yi-Fang;Sun, Xin-Xin;Sluys, Ronald;Liu, De-Zeng;Dong, Zi-Mei;Chen, Guang-Wen		Zenodo	biologists	Wang, Lei;Chang, Yi-Fang;Sun, Xin-Xin;Sluys, Ronald;Liu, De-Zeng;Dong, Zi-Mei;Chen, Guang-Wen			
6A0099F70FD45834BE0B3762F9816AF2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1296556	https://doi.org/10.3897/zookeys.1233.142976.figure3	Figure 3. Habitat and external appearance of Dugesia saccata A, B sampling site and habitat C sexually mature, live individual. Abbreviations: coa: copulatory apparatus; e: eye; ho: hyperplasic ovary; ph: pharynx. Scale bar: 2 mm.	Figure 3. Habitat and external appearance of Dugesia saccata A, B sampling site and habitat C sexually mature, live individual. Abbreviations: coa: copulatory apparatus; e: eye; ho: hyperplasic ovary; ph: pharynx. Scale bar: 2 mm.	2025-04-01	Wang, Lei;Chang, Yi-Fang;Sun, Xin-Xin;Sluys, Ronald;Liu, De-Zeng;Dong, Zi-Mei;Chen, Guang-Wen		Zenodo	biologists	Wang, Lei;Chang, Yi-Fang;Sun, Xin-Xin;Sluys, Ronald;Liu, De-Zeng;Dong, Zi-Mei;Chen, Guang-Wen			
6A0099F70FD45834BE0B3762F9816AF2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1296557	https://doi.org/10.3897/zookeys.1233.142976.figure4	Figure 4. Dugesia saccata A 1, B 1 metaphase plate and karyogram of triploid complement A 2, B 2 metaphase plate and karyogram of diploid complement C idiogram. Scale bars: 5 μm.	Figure 4. Dugesia saccata A 1, B 1 metaphase plate and karyogram of triploid complement A 2, B 2 metaphase plate and karyogram of diploid complement C idiogram. Scale bars: 5 μm.	2025-04-01	Wang, Lei;Chang, Yi-Fang;Sun, Xin-Xin;Sluys, Ronald;Liu, De-Zeng;Dong, Zi-Mei;Chen, Guang-Wen		Zenodo	biologists	Wang, Lei;Chang, Yi-Fang;Sun, Xin-Xin;Sluys, Ronald;Liu, De-Zeng;Dong, Zi-Mei;Chen, Guang-Wen			
6A0099F70FD45834BE0B3762F9816AF2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1296558	https://doi.org/10.3897/zookeys.1233.142976.figure5	Figure 5. Dugesia saccata. Photomicrographs of transverse (A, B) and sagittal (C, D) sections. A Paratype ZMHNU - YZCB 8, showing symmetrical openings of oviducts into the bursal canal B paratype ZMHNU - YZCB 8, showing symmetrical openings of vasa deferentia into the seminal vesicle C holotype ZMHNU - YZCB 1, showing copulatory bursa and bursal canal D holotype ZMHNU - YZCB 1, showing penis papilla, vasa deferentia, seminal vesicle, diaphragm, and ejaculatory duct. Abbreviations: bc: bursal canal; cb: copulatory bursa; d: diaphragm; ed: ejaculatory duct; lod: left oviduct; ma: male atrium; od: oviduct; pp: penis papilla; rod: right oviduct; se, sac-shaped expansion of bursal canal; sv: seminal vesicle; vd: vas deferens. Scale bars: 100 μm.	Figure 5. Dugesia saccata. Photomicrographs of transverse (A, B) and sagittal (C, D) sections. A Paratype ZMHNU - YZCB 8, showing symmetrical openings of oviducts into the bursal canal B paratype ZMHNU - YZCB 8, showing symmetrical openings of vasa deferentia into the seminal vesicle C holotype ZMHNU - YZCB 1, showing copulatory bursa and bursal canal D holotype ZMHNU - YZCB 1, showing penis papilla, vasa deferentia, seminal vesicle, diaphragm, and ejaculatory duct. Abbreviations: bc: bursal canal; cb: copulatory bursa; d: diaphragm; ed: ejaculatory duct; lod: left oviduct; ma: male atrium; od: oviduct; pp: penis papilla; rod: right oviduct; se, sac-shaped expansion of bursal canal; sv: seminal vesicle; vd: vas deferens. Scale bars: 100 μm.	2025-04-01	Wang, Lei;Chang, Yi-Fang;Sun, Xin-Xin;Sluys, Ronald;Liu, De-Zeng;Dong, Zi-Mei;Chen, Guang-Wen		Zenodo	biologists	Wang, Lei;Chang, Yi-Fang;Sun, Xin-Xin;Sluys, Ronald;Liu, De-Zeng;Dong, Zi-Mei;Chen, Guang-Wen			
6A0099F70FD45834BE0B3762F9816AF2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1296559	https://doi.org/10.3897/zookeys.1233.142976.figure6	Figure 6. Dugesia saccata. Sagittal reconstruction of the copulatory apparatus of holotype YZCB 1 A female copulatory apparatus B male copulatory apparatus. Abbreviations: bc: bursal canal; ca: common atrium; cb: copulatory bursa; cg: cement glands; cpg: cyanophil penial glands; d: diaphragm; ed: ejaculatory duct; epg: extrabulbar penial glands; go: gonopore; lod: left oviduct; ma: male atrium; mo: mouth; pb: penis bulb; pp: penis papilla; rod: right oviduct; se, sac-shaped expansion of bursal canal; spv: spermiducal vesicles; sg: shell glands; sv: seminal vesicle; vd: vas deferens. Scale bars: 100 μm.	Figure 6. Dugesia saccata. Sagittal reconstruction of the copulatory apparatus of holotype YZCB 1 A female copulatory apparatus B male copulatory apparatus. Abbreviations: bc: bursal canal; ca: common atrium; cb: copulatory bursa; cg: cement glands; cpg: cyanophil penial glands; d: diaphragm; ed: ejaculatory duct; epg: extrabulbar penial glands; go: gonopore; lod: left oviduct; ma: male atrium; mo: mouth; pb: penis bulb; pp: penis papilla; rod: right oviduct; se, sac-shaped expansion of bursal canal; spv: spermiducal vesicles; sg: shell glands; sv: seminal vesicle; vd: vas deferens. Scale bars: 100 μm.	2025-04-01	Wang, Lei;Chang, Yi-Fang;Sun, Xin-Xin;Sluys, Ronald;Liu, De-Zeng;Dong, Zi-Mei;Chen, Guang-Wen		Zenodo	biologists	Wang, Lei;Chang, Yi-Fang;Sun, Xin-Xin;Sluys, Ronald;Liu, De-Zeng;Dong, Zi-Mei;Chen, Guang-Wen			
