taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
0E4A87D6FFD88078FEC1FDDDFEA11F8C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/13799411/files/figure.png	https://doi.org/10.5281/zenodo.13799411	Fig. 1. Female syntypes of Coenobita longitarsis De Man, 1902 (A–G, SL 13.1 mm; H, SL 6.2 mm; SMF 8170), Maluku, Indonesia. A, dorsal view of carapace and pereiopods; B, dorsal view of shield; C, dorsal view of anterior shield and ocular acicles; D, E, outer and inner surfaces of left chela; F, outer surface of right chela; G, outer view of left P2 and P3; H. lateral view showing right P3. Scale bars: A, B, E–H = 10 mm; C, D = 5 mm.	Fig. 1. Female syntypes of Coenobita longitarsis De Man, 1902 (A–G, SL 13.1 mm; H, SL 6.2 mm; SMF 8170), Maluku, Indonesia. A, dorsal view of carapace and pereiopods; B, dorsal view of shield; C, dorsal view of anterior shield and ocular acicles; D, E, outer and inner surfaces of left chela; F, outer surface of right chela; G, outer view of left P2 and P3; H. lateral view showing right P3. Scale bars: A, B, E–H = 10 mm; C, D = 5 mm.	2023-10-25	Shih, Hsi-Te;Chang, Kai;Pramono, Félix Adhi;Celia, Maria;Malay, D.		Zenodo	biologists	Shih, Hsi-Te;Chang, Kai;Pramono, Félix Adhi;Celia, Maria;Malay, D.			
0E4A87D6FFD88078FEC1FDDDFEA11F8C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/13799413/files/figure.png	https://doi.org/10.5281/zenodo.13799413	Fig. 2. Coenobita longitarsis De Man, 1902 (ò SL 10.1 mm, UF 11434). A, shield and cephalic appendages; B, sternite and coxae of male P5; C, basal segment of right antennule, lateral view; D, flagellum of right antennule, lateral view; E, left cheliped, outer view; F, right cheliped, outer view; G, left P3, outer view. Scale bars: A, C, E–G: 5 mm; B, D = 2 mm.	Fig. 2. Coenobita longitarsis De Man, 1902 (ò SL 10.1 mm, UF 11434). A, shield and cephalic appendages; B, sternite and coxae of male P5; C, basal segment of right antennule, lateral view; D, flagellum of right antennule, lateral view; E, left cheliped, outer view; F, right cheliped, outer view; G, left P3, outer view. Scale bars: A, C, E–G: 5 mm; B, D = 2 mm.	2023-10-25	Shih, Hsi-Te;Chang, Kai;Pramono, Félix Adhi;Celia, Maria;Malay, D.		Zenodo	biologists	Shih, Hsi-Te;Chang, Kai;Pramono, Félix Adhi;Celia, Maria;Malay, D.			
0E4A87D6FFD88078FEC1FDDDFEA11F8C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/13799415/files/figure.png	https://doi.org/10.5281/zenodo.13799415	Fig. 3. Coenobita longitarsis De Man, 1902 from New Britain, East New Britain Province, Papua New Guinea. A, B, ò (SL 10.1 mm, UF 11434); C, D, ñ (SL 10.3 mm, UF 11435). Scale bars: A–D = 30 mm.	Fig. 3. Coenobita longitarsis De Man, 1902 from New Britain, East New Britain Province, Papua New Guinea. A, B, ò (SL 10.1 mm, UF 11434); C, D, ñ (SL 10.3 mm, UF 11435). Scale bars: A–D = 30 mm.	2023-10-25	Shih, Hsi-Te;Chang, Kai;Pramono, Félix Adhi;Celia, Maria;Malay, D.		Zenodo	biologists	Shih, Hsi-Te;Chang, Kai;Pramono, Félix Adhi;Celia, Maria;Malay, D.			
0E4A87D6FFD88078FEC1FDDDFEA11F8C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/13799417/files/figure.png	https://doi.org/10.5281/zenodo.13799417	Fig. 4. Coenobita longitarsis De Man, 1902 and the shells used. A–C, specimen with its shell, Lamarckiella zeus (Jonas, 1843) (Chronidae), from Sibuyan Island, central Philippines (not collected). D, ñ (SL 11.2 mm, UF 11435) (and its shell partially broken artificially); E–G, shells of Discoconcha isis (Pfeiffer, 1860) (Camaenidae) used by ñ (SL 10.3 mm, UF 11435), from Papua New Guinea. Scale bars: D–G = 20 mm. A–C, courtesy of Emerson Sy.	Fig. 4. Coenobita longitarsis De Man, 1902 and the shells used. A–C, specimen with its shell, Lamarckiella zeus (Jonas, 1843) (Chronidae), from Sibuyan Island, central Philippines (not collected). D, ñ (SL 11.2 mm, UF 11435) (and its shell partially broken artificially); E–G, shells of Discoconcha isis (Pfeiffer, 1860) (Camaenidae) used by ñ (SL 10.3 mm, UF 11435), from Papua New Guinea. Scale bars: D–G = 20 mm. A–C, courtesy of Emerson Sy.	2023-10-25	Shih, Hsi-Te;Chang, Kai;Pramono, Félix Adhi;Celia, Maria;Malay, D.		Zenodo	biologists	Shih, Hsi-Te;Chang, Kai;Pramono, Félix Adhi;Celia, Maria;Malay, D.			
0E4A87D6FFD88078FEC1FDDDFEA11F8C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/13799419/files/figure.png	https://doi.org/10.5281/zenodo.13799419	Fig. 5. Coenobita longitarsis De Man, 1902 (A, C, E, ò SL 10.1 mm, UF 11434) and C. spinosus H. Milne Edwards, 1837 (B, D, F, ò SL 21.8 mm, UF 15602). A, B, outer view of left cheliped; C, D, outer view of dactyl and propodus of left P3; E, F, coxae of male P5. Scale bars: A–D = 10 mm; E, F = 3 mm.	Fig. 5. Coenobita longitarsis De Man, 1902 (A, C, E, ò SL 10.1 mm, UF 11434) and C. spinosus H. Milne Edwards, 1837 (B, D, F, ò SL 21.8 mm, UF 15602). A, B, outer view of left cheliped; C, D, outer view of dactyl and propodus of left P3; E, F, coxae of male P5. Scale bars: A–D = 10 mm; E, F = 3 mm.	2023-10-25	Shih, Hsi-Te;Chang, Kai;Pramono, Félix Adhi;Celia, Maria;Malay, D.		Zenodo	biologists	Shih, Hsi-Te;Chang, Kai;Pramono, Félix Adhi;Celia, Maria;Malay, D.			
0E4A87D6FFD88078FEC1FDDDFEA11F8C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/13799421/files/figure.png	https://doi.org/10.5281/zenodo.13799421	Fig. 6. Coenobita pseudorugosus Nakasone, 1988 (ò SL 14.2 mm, NCHUZOOL 17199). A, shield and cephalic appendages; B, sternite and coxae of male P5; C, basal segment of right antennule, lateral view; D, flagellum of right antennule, lateral view; E, left cheliped, outer view; F, right cheliped, outer view; G, left P3, outer view. Scale bars: A, E, F = 10 mm; B, C, G = 5 mm; D = 2 mm.	Fig. 6. Coenobita pseudorugosus Nakasone, 1988 (ò SL 14.2 mm, NCHUZOOL 17199). A, shield and cephalic appendages; B, sternite and coxae of male P5; C, basal segment of right antennule, lateral view; D, flagellum of right antennule, lateral view; E, left cheliped, outer view; F, right cheliped, outer view; G, left P3, outer view. Scale bars: A, E, F = 10 mm; B, C, G = 5 mm; D = 2 mm.	2023-10-25	Shih, Hsi-Te;Chang, Kai;Pramono, Félix Adhi;Celia, Maria;Malay, D.		Zenodo	biologists	Shih, Hsi-Te;Chang, Kai;Pramono, Félix Adhi;Celia, Maria;Malay, D.			
0E4A87D6FFD88078FEC1FDDDFEA11F8C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/13799425/files/figure.png	https://doi.org/10.5281/zenodo.13799425	Fig. 7. Coenobita pseudorugosus Nakasone, 1988. A, B, ò (SL 14.2 mm, NCHUZOOL 17199, preserved specimen). C–H, color in life (specimens from SW Sumatra; not kept).	Fig. 7. Coenobita pseudorugosus Nakasone, 1988. A, B, ò (SL 14.2 mm, NCHUZOOL 17199, preserved specimen). C–H, color in life (specimens from SW Sumatra; not kept).	2023-10-25	Shih, Hsi-Te;Chang, Kai;Pramono, Félix Adhi;Celia, Maria;Malay, D.		Zenodo	biologists	Shih, Hsi-Te;Chang, Kai;Pramono, Félix Adhi;Celia, Maria;Malay, D.			
0E4A87D6FFD88078FEC1FDDDFEA11F8C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/13799427/files/figure.png	https://doi.org/10.5281/zenodo.13799427	Fig. 8. Outer view of left cheliped in Coenobita pseudorugosus Nakasone, 1988 (A, C, E, G) and C. rugosus H. Milne Edwards, 1837 (B, D, F, H). A, ò (SL = 4.7 mm, UF 13164); B, ò (SL = 5.1 mm, ZRC); C, ò (SL = 5.0 mm, UF 13164); D, ñ (SL = 6.8 mm, NCHUZOOL 17198); E, ò (SL = 12.2 mm, NCHUZOOL 17201); F, ò (SL = 14.1 mm, NCHUZOOL 17205); G, ò (SL = 14.2 mm, NCHUZOOL 17199); H, ò SL = 17.8 mm, NCHUZOOL 17202). Scale bars: A–D = 3 mm; E–F = 10 mm.	Fig. 8. Outer view of left cheliped in Coenobita pseudorugosus Nakasone, 1988 (A, C, E, G) and C. rugosus H. Milne Edwards, 1837 (B, D, F, H). A, ò (SL = 4.7 mm, UF 13164); B, ò (SL = 5.1 mm, ZRC); C, ò (SL = 5.0 mm, UF 13164); D, ñ (SL = 6.8 mm, NCHUZOOL 17198); E, ò (SL = 12.2 mm, NCHUZOOL 17201); F, ò (SL = 14.1 mm, NCHUZOOL 17205); G, ò (SL = 14.2 mm, NCHUZOOL 17199); H, ò SL = 17.8 mm, NCHUZOOL 17202). Scale bars: A–D = 3 mm; E–F = 10 mm.	2023-10-25	Shih, Hsi-Te;Chang, Kai;Pramono, Félix Adhi;Celia, Maria;Malay, D.		Zenodo	biologists	Shih, Hsi-Te;Chang, Kai;Pramono, Félix Adhi;Celia, Maria;Malay, D.			
0E4A87D6FFD88078FEC1FDDDFEA11F8C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/13799429/files/figure.png	https://doi.org/10.5281/zenodo.13799429	Fig. 9. Sternite and coxae of male P5 in Coenobita pseudorugosus Nakasone, 1988 (A, C, E, F) and C. rugosus H. Milne Edwards, 1837 (B, D). A, ò (SL 5.0 mm, UF 13164); B, ò (SL 5.1 mm, ZRC); C, ò (SL 14.2 mm, NCHUZOOL 17199); D, ò (SL 14.1 mm, NCHUZOOL 17205); E, ò (SL 4.7 mm, UF 13164); F, ò (SL 9.9 mm, NCHUZOOL 17200). Scale bars = 2 mm.	Fig. 9. Sternite and coxae of male P5 in Coenobita pseudorugosus Nakasone, 1988 (A, C, E, F) and C. rugosus H. Milne Edwards, 1837 (B, D). A, ò (SL 5.0 mm, UF 13164); B, ò (SL 5.1 mm, ZRC); C, ò (SL 14.2 mm, NCHUZOOL 17199); D, ò (SL 14.1 mm, NCHUZOOL 17205); E, ò (SL 4.7 mm, UF 13164); F, ò (SL 9.9 mm, NCHUZOOL 17200). Scale bars = 2 mm.	2023-10-25	Shih, Hsi-Te;Chang, Kai;Pramono, Félix Adhi;Celia, Maria;Malay, D.		Zenodo	biologists	Shih, Hsi-Te;Chang, Kai;Pramono, Félix Adhi;Celia, Maria;Malay, D.			
0E4A87D6FFD88078FEC1FDDDFEA11F8C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/13799431/files/figure.png	https://doi.org/10.5281/zenodo.13799431	Fig. 10. Outer view of left P3 in Coenobita pseudorugosus Nakasone, 1988 (A, C, E; SL 5.0–14.2 mm) and C. rugosus H. Milne Edwards, 1837 (B, D, F–H; SL 5.1–17.8 mm). A, ò (SL 5.0 mm, UF 13164); B, ò (SL 5.1 mm, ZRC); C, ñ (SL 6.9 mm, UF 7382); D, ñ (SL 6.8 mm, NCHUZOOL 17198); E, ò (SL 14.2 mm, NCHUZOOL 17199); F, ò (SL 14.1 mm, NCHUZOOL 17205); G, ñ (SL 16.4 mm, NCHUZOOL 17206); H, ò (SL 17.8 mm, NCHUZOOL 17202). Scale bars: A–D = 5 mm; E–H = 10 mm.	Fig. 10. Outer view of left P3 in Coenobita pseudorugosus Nakasone, 1988 (A, C, E; SL 5.0–14.2 mm) and C. rugosus H. Milne Edwards, 1837 (B, D, F–H; SL 5.1–17.8 mm). A, ò (SL 5.0 mm, UF 13164); B, ò (SL 5.1 mm, ZRC); C, ñ (SL 6.9 mm, UF 7382); D, ñ (SL 6.8 mm, NCHUZOOL 17198); E, ò (SL 14.2 mm, NCHUZOOL 17199); F, ò (SL 14.1 mm, NCHUZOOL 17205); G, ñ (SL 16.4 mm, NCHUZOOL 17206); H, ò (SL 17.8 mm, NCHUZOOL 17202). Scale bars: A–D = 5 mm; E–H = 10 mm.	2023-10-25	Shih, Hsi-Te;Chang, Kai;Pramono, Félix Adhi;Celia, Maria;Malay, D.		Zenodo	biologists	Shih, Hsi-Te;Chang, Kai;Pramono, Félix Adhi;Celia, Maria;Malay, D.			
0E4A87D6FFD88078FEC1FDDDFEA11F8C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/13799433/files/figure.png	https://doi.org/10.5281/zenodo.13799433	Fig. 11. Dorsal view of left P3 propodus in Coenobita pseudorugosus Nakasone, 1988 (A, C, E, G, I, K; SL 5.0–14.2 mm) and C. rugosus H. Milne Edwards, 1837 (B, D, F, H, J, L; SL 5.1–17.8 mm), showing the degree of swelling of lateral surface of left P3 propodus. A, ò (SL 5.0 mm, UF 13164); B, ò (SL 5.1 mm, ZRC); C, ñ (SL 6.9 mm, UF 7382); D, ñ (SL 6.8 mm, NCHUZOOL 17198); E, ò (SL 11.7 mm, NCHUZOOL 17201); F, ò (SL 11.2 mm, QM W17540); G, ò (SL 12.6 mm, NCHUZOOL 17200); H, ñ (SL 12.9 mm, NCHUZOOL 17203); I, ò (SL 13.0 mm, NCHUZOOL 17199); J, ñ (SL 13.1 mm, NCHUZOOL 17204); K, ò (SL 14.2 mm, NCHUZOOL 17199); L, ò (SL 17.8 mm, NCHUZOOL 17202). Scale bars: A–D = 3 mm; E–H = 5 mm.	Fig. 11. Dorsal view of left P3 propodus in Coenobita pseudorugosus Nakasone, 1988 (A, C, E, G, I, K; SL 5.0–14.2 mm) and C. rugosus H. Milne Edwards, 1837 (B, D, F, H, J, L; SL 5.1–17.8 mm), showing the degree of swelling of lateral surface of left P3 propodus. A, ò (SL 5.0 mm, UF 13164); B, ò (SL 5.1 mm, ZRC); C, ñ (SL 6.9 mm, UF 7382); D, ñ (SL 6.8 mm, NCHUZOOL 17198); E, ò (SL 11.7 mm, NCHUZOOL 17201); F, ò (SL 11.2 mm, QM W17540); G, ò (SL 12.6 mm, NCHUZOOL 17200); H, ñ (SL 12.9 mm, NCHUZOOL 17203); I, ò (SL 13.0 mm, NCHUZOOL 17199); J, ñ (SL 13.1 mm, NCHUZOOL 17204); K, ò (SL 14.2 mm, NCHUZOOL 17199); L, ò (SL 17.8 mm, NCHUZOOL 17202). Scale bars: A–D = 3 mm; E–H = 5 mm.	2023-10-25	Shih, Hsi-Te;Chang, Kai;Pramono, Félix Adhi;Celia, Maria;Malay, D.		Zenodo	biologists	Shih, Hsi-Te;Chang, Kai;Pramono, Félix Adhi;Celia, Maria;Malay, D.			
0E4A87D6FFD88078FEC1FDDDFEA11F8C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/13799435/files/figure.png	https://doi.org/10.5281/zenodo.13799435	Fig. 12. A neighbor-joining tree for 10 species of Coenobita from the Indo-West Pacific, based on the cytochrome c oxidase subunit I (COI) gene. Probability values at the nodes represent support values. Only values> 50% are shown. For haplotype names, see table 1. Table 2. Distinguishing characters of Coenobita pseudorugosus and C. rugosus	Fig. 12. A neighbor-joining tree for 10 species of Coenobita from the Indo-West Pacific, based on the cytochrome c oxidase subunit I (COI) gene. Probability values at the nodes represent support values. Only values> 50% are shown. For haplotype names, see table 1. Table 2. Distinguishing characters of Coenobita pseudorugosus and C. rugosus	2023-10-25	Shih, Hsi-Te;Chang, Kai;Pramono, Félix Adhi;Celia, Maria;Malay, D.		Zenodo	biologists	Shih, Hsi-Te;Chang, Kai;Pramono, Félix Adhi;Celia, Maria;Malay, D.			
