taxonID	type	description	language	source
0B4787FCFF8E1A27FF29FAD73B4AF837.taxon	diagnosis	Diagnosis. Taxicerini is composed of three genera: Discerota Mulsant & Rey, 1873; Halobrecta Thomson, 1858; Taxicera Mulsant & Rey, 1873. Taxicera differs from Discerota Mulsant & Rey, 1873 by the shorter legs (Kapp, 2005). Taxicera differs from Halobrecta Thomson, 1858 by the smaller or lacking umbilicus of spermatheca and shorter apical lobe of paramere (modified after Gusarov, 2004). Taxicera can be easily distinguished from Dalotia by the following characteristics: ligula not divided or divided apical 1 / 3 at most (figs. 34 & 35 in Brundin, 1943; 4, 24, 33, 41, 60, 67, 79 & 82 in Kapp, 2005) (Brundin, 1943); spermatheca not coiled, with circular head (figs. 39 in Brundin, 1943; 10 – 14, 25, 30, 41, 54 – 58, 72 & 83 in Kapp, 2005); median lobe of male aedeagus without athetine bridge (figs. 40, 42 & 43 in Brundin, 1943; 46 in Kapp, 2005) (Osswald et al., 2013).	en	Tasaku, Yuto, Maruyama, Munetoshi (2025): Resolving the taxonomic confusion of Taxicera Mulsant & Rey, 1873 and Dalotia Casey, 1910 (Coleoptera, Staphylinidae, Aleocharinae), with the rediscovery of Dalotia pseudorphana (Cameron, 1933), comb. nov., from bird nest material. Zootaxa 5609 (1): 70-82, DOI: 10.11646/zootaxa.5609.1.4, URL: https://doi.org/10.11646/zootaxa.5609.1.4
0B4787FCFF8D1A24FF29FF1E3D58FC16.taxon	diagnosis	Diagnosis. Although the general appearance is very similar to other genera of Athetini, it differs by the following characteristics: ligula with two long setae; long hypopharyngeal setae; spermatheca without umbilicus (modified after Gusarov, 2003 and Muona, 2023). Compared with Taxicera, this genus can be distinguished by following characteristics: ligula divided at apical half; median lobe with athetine bridge; spermatheca once coiled (see also Discussion).	en	Tasaku, Yuto, Maruyama, Munetoshi (2025): Resolving the taxonomic confusion of Taxicera Mulsant & Rey, 1873 and Dalotia Casey, 1910 (Coleoptera, Staphylinidae, Aleocharinae), with the rediscovery of Dalotia pseudorphana (Cameron, 1933), comb. nov., from bird nest material. Zootaxa 5609 (1): 70-82, DOI: 10.11646/zootaxa.5609.1.4, URL: https://doi.org/10.11646/zootaxa.5609.1.4
0B4787FCFF8D1A24FF29FF1E3D58FC16.taxon	discussion	Remarks. Muona (2023) described the highly complicated internal structure of the median lobe of male aedeagus for the diagnostic characteristic of Dalotia. It would be meant internal structure with large suspensoria and long sclerites described in Yosii & Sawada (1976). However, the large suspensoria are also observed in some athetine genera, such as Neodimetrota Muona, 2023, Notothecta Thomson, 1858. Furthermore, in Dalotia pseudorphana comb. nov., described later, the suspensoria is relatively small and long sclerites are not observed (Fig. 6 B in this paper and fig. 19 P in Sawada, 1977) but, it is clearly closely similar to D. coriaria and belong to Dalotia by the characteristics of ligula, median lobe, and spermatheca (see also Discussion). Thus, this characteristic should be deleted from the diagnosis of Dalotia.	en	Tasaku, Yuto, Maruyama, Munetoshi (2025): Resolving the taxonomic confusion of Taxicera Mulsant & Rey, 1873 and Dalotia Casey, 1910 (Coleoptera, Staphylinidae, Aleocharinae), with the rediscovery of Dalotia pseudorphana (Cameron, 1933), comb. nov., from bird nest material. Zootaxa 5609 (1): 70-82, DOI: 10.11646/zootaxa.5609.1.4, URL: https://doi.org/10.11646/zootaxa.5609.1.4
0B4787FCFF8D1A22FF29FBC13EA1FE14.taxon	description	(Figs. 1 A – B, 3)	en	Tasaku, Yuto, Maruyama, Munetoshi (2025): Resolving the taxonomic confusion of Taxicera Mulsant & Rey, 1873 and Dalotia Casey, 1910 (Coleoptera, Staphylinidae, Aleocharinae), with the rediscovery of Dalotia pseudorphana (Cameron, 1933), comb. nov., from bird nest material. Zootaxa 5609 (1): 70-82, DOI: 10.11646/zootaxa.5609.1.4, URL: https://doi.org/10.11646/zootaxa.5609.1.4
0B4787FCFF8D1A22FF29FBC13EA1FE14.taxon	materials_examined	Materials examined. Four males and six females, Kamimaki, Ina city, Nagano Pref., Japan, 26. VII. 2024, S. Hashizume (KUM); two males and one female, Daitô-jinja, Minamidaitô-jima Island, Okinawa Pref., Japan, 27. VI. 2023, T. Kinoda (KUM).	en	Tasaku, Yuto, Maruyama, Munetoshi (2025): Resolving the taxonomic confusion of Taxicera Mulsant & Rey, 1873 and Dalotia Casey, 1910 (Coleoptera, Staphylinidae, Aleocharinae), with the rediscovery of Dalotia pseudorphana (Cameron, 1933), comb. nov., from bird nest material. Zootaxa 5609 (1): 70-82, DOI: 10.11646/zootaxa.5609.1.4, URL: https://doi.org/10.11646/zootaxa.5609.1.4
0B4787FCFF8D1A22FF29FBC13EA1FE14.taxon	diagnosis	Diagnosis. Dalotia coriaria differs from other species of Dalotia by the following characteristics: apical margin of male tergite VIII dentate (fig. 55 G in Yosii & Sawada, 1976; 5 A in Lee et al., 2024); median lobe of male aedeagus with large suspensoria and long sclerites (figs. 55 K in Yosii & Sawada, 1976; 6 A & B in Muona, 2023; vagina with two pairs of sclerites (Fig. 3 D). Additional description. Male. Tergites IX, X, and sternite IX shown in Figs. 3 A & B. Female. Tergites IX, X, and sternite IX shown in Figs. 3 C & D. Vagina weakly sclerotized, semicircular, with two pairs of sclerite (Fig. 3 D). Spermatheca: head elongate, without umbilicus; stem short (about twice as long as spermathecal head), helically coiled once around base (Fig. 3 D).	en	Tasaku, Yuto, Maruyama, Munetoshi (2025): Resolving the taxonomic confusion of Taxicera Mulsant & Rey, 1873 and Dalotia Casey, 1910 (Coleoptera, Staphylinidae, Aleocharinae), with the rediscovery of Dalotia pseudorphana (Cameron, 1933), comb. nov., from bird nest material. Zootaxa 5609 (1): 70-82, DOI: 10.11646/zootaxa.5609.1.4, URL: https://doi.org/10.11646/zootaxa.5609.1.4
0B4787FCFF8D1A22FF29FBC13EA1FE14.taxon	distribution	Distribution. Japan: Honshu, Minamidaitô-jima Island (new record); cosmopolitan. Habitat. This species is mainly collected from piled grasses.	en	Tasaku, Yuto, Maruyama, Munetoshi (2025): Resolving the taxonomic confusion of Taxicera Mulsant & Rey, 1873 and Dalotia Casey, 1910 (Coleoptera, Staphylinidae, Aleocharinae), with the rediscovery of Dalotia pseudorphana (Cameron, 1933), comb. nov., from bird nest material. Zootaxa 5609 (1): 70-82, DOI: 10.11646/zootaxa.5609.1.4, URL: https://doi.org/10.11646/zootaxa.5609.1.4
0B4787FCFF8D1A22FF29FBC13EA1FE14.taxon	discussion	Remarks. This species is distributed worldwide (Newton, 2024). From Japan, it is only known from Honshu. Here, we record this species from Minamidaitô-jima Island, located approximately 400 km east of the Okinawajima Is., for the first time.	en	Tasaku, Yuto, Maruyama, Munetoshi (2025): Resolving the taxonomic confusion of Taxicera Mulsant & Rey, 1873 and Dalotia Casey, 1910 (Coleoptera, Staphylinidae, Aleocharinae), with the rediscovery of Dalotia pseudorphana (Cameron, 1933), comb. nov., from bird nest material. Zootaxa 5609 (1): 70-82, DOI: 10.11646/zootaxa.5609.1.4, URL: https://doi.org/10.11646/zootaxa.5609.1.4
0B4787FCFF8B1A2FFF29FE3F3ED3FD3C.taxon	description	(Figs. 1 C – D, 2, 4 – 6)	en	Tasaku, Yuto, Maruyama, Munetoshi (2025): Resolving the taxonomic confusion of Taxicera Mulsant & Rey, 1873 and Dalotia Casey, 1910 (Coleoptera, Staphylinidae, Aleocharinae), with the rediscovery of Dalotia pseudorphana (Cameron, 1933), comb. nov., from bird nest material. Zootaxa 5609 (1): 70-82, DOI: 10.11646/zootaxa.5609.1.4, URL: https://doi.org/10.11646/zootaxa.5609.1.4
0B4787FCFF8B1A2FFF29FE3F3ED3FD3C.taxon	materials_examined	Type material. Holotype. Male, data on labels: “ Kobe / Harad / 24. VI- 12 / A. / pseudorphana / TYPE Cam. // Cameron / Bequest / B. M. 1955 - 147 // Atheta / pseudorphana Cam. / P. M. Hammond / det. 1973 / HOLOTYPE ” (Fig. 2 dissected by K. Sawada; deposited in the Natural History Museum, London). Specimens examined. Two males, two females, and five unsexed, FPI (nest number K- 9, Parus varius), 13. VI. 2023 (SEHU); one male, one female, and four unsexed, TUS (nest number 22, Parus minor), 4. VII. 2023 (SEHU); one female and one unsexed, TUS (nest number 64, Parus minor), 19. VI. 2023 (KUM).	en	Tasaku, Yuto, Maruyama, Munetoshi (2025): Resolving the taxonomic confusion of Taxicera Mulsant & Rey, 1873 and Dalotia Casey, 1910 (Coleoptera, Staphylinidae, Aleocharinae), with the rediscovery of Dalotia pseudorphana (Cameron, 1933), comb. nov., from bird nest material. Zootaxa 5609 (1): 70-82, DOI: 10.11646/zootaxa.5609.1.4, URL: https://doi.org/10.11646/zootaxa.5609.1.4
0B4787FCFF8B1A2FFF29FE3F3ED3FD3C.taxon	diagnosis	Diagnosis. Dalotia pseudorphana comb. nov. differs from other species of Dalotia by the shape of male tergite VIII, and median lobe. From Dalotia coriaria, this species easily differs by the smaller body (Figs 1 C & D), truncated apical margin of male tergite VIII (Fig. 3 A), and vagina with spines (Fig. 5 D).	en	Tasaku, Yuto, Maruyama, Munetoshi (2025): Resolving the taxonomic confusion of Taxicera Mulsant & Rey, 1873 and Dalotia Casey, 1910 (Coleoptera, Staphylinidae, Aleocharinae), with the rediscovery of Dalotia pseudorphana (Cameron, 1933), comb. nov., from bird nest material. Zootaxa 5609 (1): 70-82, DOI: 10.11646/zootaxa.5609.1.4, URL: https://doi.org/10.11646/zootaxa.5609.1.4
0B4787FCFF8B1A2FFF29FE3F3ED3FD3C.taxon	description	Redescription. Body subparallel-sided, flat (Figs 1 C & D). Coloration. Head and abdomen black; antennomeres III – XI, pronotum, and elytra reddish brown; antennomeres I and II, legs, and mouth parts yellow. Head. Capsule circular, weakly shining; infraorbital carina absent; eyes large, longer than temporal length. Antennae. Antennomeres I – III elongate, II slightly shorter than I, III as long as II, IV – X transverse, XI oval, twice as long as width. Mouth parts. Labrum transverse, with slightly emarginated apical margin, with 8 macrosetae on each side. Epipharynx with setaceous α-sensillum, with very small β and γ-sensillum. Mandibles triangular, slender, almost symmetrical; light mandible with tiny tooth. Maxilla: maxillary palpus elongate, palpomere I very small, segment II about three times longer than I, III slightly longer than II, segment IV thin, as long as III. Mentum trapezoidal, with slightly emarginate apical margin. Labium (Fig. 6 E): palpus with three palpomeres, chaetotaxy shown in Fig. 6 F; prementum with narrow and smooth median area; ligula divided at apical half, with seta on each branch in ventral side; premental apodeme slender, as long as labial palpus. Thorax. Pronotum oblong, wider than long, weakly shining; hypomera almost invisible in lateral view. Elytra longer than wide, densely covered with setae. Hind wing fully developed. Tarsal formula 4 - 5 - 5. Abdomen strongly shining, sparsely covered with setae. Male. Tergite VIII quadrate, with straight posterior margin, with 4 + 4 macrosetae (Fig. 4 A). Sternite VIII semicircular, with 8 + 6 macrosetae (Fig. 4 B). Tergites IX, X, and sternite X as shown in Figs. 5 A & B. Median lobe of aedeagus (Figs. 6 A – C): apical lobe simple, slender, as long as basal capsule; basal capsule oval; athetine bridge thin, incomplete; internal structure simple. Parameres of aedeagus broad with slender apical lobe (Fig. 6 D). Female. Tergite VIII quadrate, with straight posterior margin, with 4 + 4 macrosetae (Fig. 4 C). Sternite VIII semicircular; marginal setae thick, 4 setae of center long (Fig. 4 D). Tergites IX, X, and sternite X shown in Figs. 5 C & D. Vagina weakly sclerotized, longer than wide, with about 40 spines on apical half (Fig. 5 D). Spermatheca (Fig. 5 E): head elongate, without umbilicus; stem short (about twice as long as spermathecal head), coiled once helically around base. Measurements. BL, 1.70 – 2.14, FBL 0.96 – 1.28, HW 0.34 – 0.39, PW 0.47 – 0.49, PL 0.32 – 0.37, EW 0.54 – 0.60.	en	Tasaku, Yuto, Maruyama, Munetoshi (2025): Resolving the taxonomic confusion of Taxicera Mulsant & Rey, 1873 and Dalotia Casey, 1910 (Coleoptera, Staphylinidae, Aleocharinae), with the rediscovery of Dalotia pseudorphana (Cameron, 1933), comb. nov., from bird nest material. Zootaxa 5609 (1): 70-82, DOI: 10.11646/zootaxa.5609.1.4, URL: https://doi.org/10.11646/zootaxa.5609.1.4
0B4787FCFF8B1A2FFF29FE3F3ED3FD3C.taxon	distribution	Distribution. Japan: Hokkaido (new record), Honshu. Habitat. Bird nest materials (Parus varius (Temminck & Schlegel, 1848) and Parus minor Temminck & Schlegel, 1848).	en	Tasaku, Yuto, Maruyama, Munetoshi (2025): Resolving the taxonomic confusion of Taxicera Mulsant & Rey, 1873 and Dalotia Casey, 1910 (Coleoptera, Staphylinidae, Aleocharinae), with the rediscovery of Dalotia pseudorphana (Cameron, 1933), comb. nov., from bird nest material. Zootaxa 5609 (1): 70-82, DOI: 10.11646/zootaxa.5609.1.4, URL: https://doi.org/10.11646/zootaxa.5609.1.4
0B4787FCFF8B1A2FFF29FE3F3ED3FD3C.taxon	discussion	Remarks. The holotype is in poor condition (whole body soaked with KOH and body color changed (Fig. 2 A); mouth parts, abdominal segments VIII – X, and aedeagus were dissected and glued by the previous researcher (Fig. 2 B )), so no further information is available. The identification was conducted based on illustrations and descriptions in Sawada (1977). Only the holotype has been known for this species for nearly a hundred years. It is probably because this species is nidicolous and difficult to collect. As surveys progress, this species will be found in other East Asian areas. We could not find this species from nest materials of Agropsar philippensis (J. R. Forster, 1781) and Passer montanus (Linnaeus, 1758), composed by plants matter, particularly grasses. Based on the morphological characteristics, this species should belong to Dalotia, not Taxicera Mulsant & Rey, 1873 (see also Discussion).	en	Tasaku, Yuto, Maruyama, Munetoshi (2025): Resolving the taxonomic confusion of Taxicera Mulsant & Rey, 1873 and Dalotia Casey, 1910 (Coleoptera, Staphylinidae, Aleocharinae), with the rediscovery of Dalotia pseudorphana (Cameron, 1933), comb. nov., from bird nest material. Zootaxa 5609 (1): 70-82, DOI: 10.11646/zootaxa.5609.1.4, URL: https://doi.org/10.11646/zootaxa.5609.1.4
0B4787FCFF861A2FFF29FB963E4EFA12.taxon	materials_examined	Type material. Not examined.	en	Tasaku, Yuto, Maruyama, Munetoshi (2025): Resolving the taxonomic confusion of Taxicera Mulsant & Rey, 1873 and Dalotia Casey, 1910 (Coleoptera, Staphylinidae, Aleocharinae), with the rediscovery of Dalotia pseudorphana (Cameron, 1933), comb. nov., from bird nest material. Zootaxa 5609 (1): 70-82, DOI: 10.11646/zootaxa.5609.1.4, URL: https://doi.org/10.11646/zootaxa.5609.1.4
0B4787FCFF861A2FFF29FB963E4EFA12.taxon	distribution	Distribution. Indonesia (Java).	en	Tasaku, Yuto, Maruyama, Munetoshi (2025): Resolving the taxonomic confusion of Taxicera Mulsant & Rey, 1873 and Dalotia Casey, 1910 (Coleoptera, Staphylinidae, Aleocharinae), with the rediscovery of Dalotia pseudorphana (Cameron, 1933), comb. nov., from bird nest material. Zootaxa 5609 (1): 70-82, DOI: 10.11646/zootaxa.5609.1.4, URL: https://doi.org/10.11646/zootaxa.5609.1.4
0B4787FCFF861A2FFF29FB963E4EFA12.taxon	discussion	Remarks. This species was transferred to Taxicera in Yosii & Sawada (1976), based on morphological similarities with T. academica (now junior synonym of D. coriaria). This species should also belong to Dalotia (see also Discussion).	en	Tasaku, Yuto, Maruyama, Munetoshi (2025): Resolving the taxonomic confusion of Taxicera Mulsant & Rey, 1873 and Dalotia Casey, 1910 (Coleoptera, Staphylinidae, Aleocharinae), with the rediscovery of Dalotia pseudorphana (Cameron, 1933), comb. nov., from bird nest material. Zootaxa 5609 (1): 70-82, DOI: 10.11646/zootaxa.5609.1.4, URL: https://doi.org/10.11646/zootaxa.5609.1.4
0B4787FCFF861A2FFF29FD173CDCFBBF.taxon	materials_examined	Type material. Not examined.	en	Tasaku, Yuto, Maruyama, Munetoshi (2025): Resolving the taxonomic confusion of Taxicera Mulsant & Rey, 1873 and Dalotia Casey, 1910 (Coleoptera, Staphylinidae, Aleocharinae), with the rediscovery of Dalotia pseudorphana (Cameron, 1933), comb. nov., from bird nest material. Zootaxa 5609 (1): 70-82, DOI: 10.11646/zootaxa.5609.1.4, URL: https://doi.org/10.11646/zootaxa.5609.1.4
0B4787FCFF861A2FFF29FD173CDCFBBF.taxon	distribution	Distribution. Malaysia: Sabah (Mt. Kinabalu).	en	Tasaku, Yuto, Maruyama, Munetoshi (2025): Resolving the taxonomic confusion of Taxicera Mulsant & Rey, 1873 and Dalotia Casey, 1910 (Coleoptera, Staphylinidae, Aleocharinae), with the rediscovery of Dalotia pseudorphana (Cameron, 1933), comb. nov., from bird nest material. Zootaxa 5609 (1): 70-82, DOI: 10.11646/zootaxa.5609.1.4, URL: https://doi.org/10.11646/zootaxa.5609.1.4
0B4787FCFF861A2FFF29FD173CDCFBBF.taxon	discussion	Remarks. This species was transferred to Taxicera in Sawada (1979). However, this species should be regarded as a species of Dalotia considering his description (see also Discussion).	en	Tasaku, Yuto, Maruyama, Munetoshi (2025): Resolving the taxonomic confusion of Taxicera Mulsant & Rey, 1873 and Dalotia Casey, 1910 (Coleoptera, Staphylinidae, Aleocharinae), with the rediscovery of Dalotia pseudorphana (Cameron, 1933), comb. nov., from bird nest material. Zootaxa 5609 (1): 70-82, DOI: 10.11646/zootaxa.5609.1.4, URL: https://doi.org/10.11646/zootaxa.5609.1.4
