identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
0B4787FCFF8E1A27FF29FAD73B4AF837.text	0B4787FCFF8E1A27FF29FAD73B4AF837.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Taxicera Mulsant & Rey 1873	<div><p>Genus  Taxicera Mulsant &amp; Rey, 1873</p><p>Taxicera Mulsant &amp; Rey, 1873: 188 (original description; type species:  Taxicera perfoliata Mulsant &amp; Rey, 1873 (now junior synonym of  Aleocara deplanata Gravenhorst, 1802)); Brundin, 1943 (description); Smetana, 2004: 419 (catalogue); Kapp, 2005: 1282 (description); Schülke &amp; Smetana, 2015: 728 (catalogue); Newton, 2024 (catalogue).</p><p>Homalota (Taxicera): Gozis, 1886: 12 (catalogue).</p><p>Atheta (Taxicera): Ganglbauer, 1895: 208; Bernhauer &amp; Scheerpeltz, 1926: 662 (catalogue).</p><p>Parataxicera Brundin, 1943 (original description, as subgenus of  Taxicera; type species:  Atheta moczarskii Bernhauer, 1914); Smetana, 2004: 419 (catalogue); Kapp, 2005: 1300 (as synonym of  Taxicera); Schülke &amp; Smetana, 2015: 728 (catalogue); Newton, 2024 (catalogue as synonym of  Taxicera).</p><p>Diagnosis.  Taxicerini is composed of three genera:  Discerota Mulsant &amp; Rey, 1873;  Halobrecta Thomson, 1858;  Taxicera Mulsant &amp; Rey, 1873 .  Taxicera differs from  Discerota Mulsant &amp; Rey, 1873 by the shorter legs (Kapp, 2005).  Taxicera differs from  Halobrecta Thomson, 1858 by the smaller or lacking umbilicus of spermatheca and shorter apical lobe of paramere (modified after Gusarov, 2004).  Taxicera can be easily distinguished from  Dalotia by the following characteristics: ligula not divided or divided apical 1/3 at most (figs. 34 &amp; 35 in Brundin, 1943; 4, 24, 33, 41, 60, 67, 79 &amp; 82 in Kapp, 2005) (Brundin, 1943); spermatheca not coiled, with circular head (figs. 39 in Brundin, 1943; 10–14, 25, 30, 41, 54–58, 72 &amp; 83 in Kapp, 2005); median lobe of male aedeagus without athetine bridge (figs. 40, 42 &amp; 43 in Brundin, 1943; 46 in Kapp, 2005) (Osswald et al., 2013).</p></div>	https://treatment.plazi.org/id/0B4787FCFF8E1A27FF29FAD73B4AF837	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Tasaku, Yuto;Maruyama, Munetoshi	Tasaku, Yuto, Maruyama, Munetoshi (2025): Resolving the taxonomic confusion of Taxicera Mulsant & Rey, 1873 and Dalotia Casey, 1910 (Coleoptera, Staphylinidae, Aleocharinae), with the rediscovery of Dalotia pseudorphana (Cameron, 1933), comb. nov., from bird nest material. Zootaxa 5609 (1): 70-82, DOI: 10.11646/zootaxa.5609.1.4, URL: https://doi.org/10.11646/zootaxa.5609.1.4
0B4787FCFF8D1A24FF29FF1E3D58FC16.text	0B4787FCFF8D1A24FF29FF1E3D58FC16.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dalotia Casey 1910	<div><p>Genus  Dalotia Casey, 1910</p><p>Dalotia Casey, 1910: 106 (original description, as a subgenus of  Dimetrota; type species:  Dimetrota pectorina Casey, 1910).</p><p>Atheta (Dalotia): Bernhauer &amp; Scheerpeltz, 1926: 662 (catalogue, as a synonym of the subgenus  Dimetrota); Smetana, 2004: 380 (catalogue); Schülke &amp; Smetana, 2015: 529 (catalogue).</p><p>Dalotia: Gusarov, 2003: 88 (raised to generic rank, diagnosis); Newton, 2024 (catalogue); Muona, 2023: 19 (diagnosis).</p><p>“  Taxicera ”: Yosii &amp; Sawada, 1976: 125 (diagnosis).</p><p>Diagnosis. Although the general appearance is very similar to other genera of  Athetini, it differs by the following characteristics: ligula with two long setae; long hypopharyngeal setae; spermatheca without umbilicus (modified after Gusarov, 2003 and Muona, 2023). Compared with  Taxicera, this genus can be distinguished by following characteristics: ligula divided at apical half; median lobe with athetine bridge; spermatheca once coiled (see also Discussion).</p><p>Remarks. Muona (2023) described the highly complicated internal structure of the median lobe of male aedeagus for the diagnostic characteristic of  Dalotia . It would be meant internal structure with large suspensoria and long sclerites described in Yosii &amp; Sawada (1976). However, the large suspensoria are also observed in some athetine genera, such as Neodimetrota Muona, 2023,  Notothecta Thomson, 1858 . Furthermore, in  Dalotia pseudorphana comb. nov., described later, the suspensoria is relatively small and long sclerites are not observed (Fig. 6B in this paper and fig. 19P in Sawada, 1977) but, it is clearly closely similar to  D. coriaria and belong to  Dalotia by the characteristics of ligula, median lobe, and spermatheca (see also Discussion). Thus, this characteristic should be deleted from the diagnosis of  Dalotia .</p></div>	https://treatment.plazi.org/id/0B4787FCFF8D1A24FF29FF1E3D58FC16	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Tasaku, Yuto;Maruyama, Munetoshi	Tasaku, Yuto, Maruyama, Munetoshi (2025): Resolving the taxonomic confusion of Taxicera Mulsant & Rey, 1873 and Dalotia Casey, 1910 (Coleoptera, Staphylinidae, Aleocharinae), with the rediscovery of Dalotia pseudorphana (Cameron, 1933), comb. nov., from bird nest material. Zootaxa 5609 (1): 70-82, DOI: 10.11646/zootaxa.5609.1.4, URL: https://doi.org/10.11646/zootaxa.5609.1.4
0B4787FCFF8D1A22FF29FBC13EA1FE14.text	0B4787FCFF8D1A22FF29FBC13EA1FE14.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dalotia coriaria (Kraatz 1856)	<div><p>Dalotia coriaria (Kraatz, 1856)</p><p>(Figs. 1A–B, 3)</p><p>Homalota coriaria Kraatz, 1856: 282 (original description); Wollaston, 1864: 543 (catalogue); Sharp, 1869: 204 (description).</p><p>Homalota subcoriaria Wollaston, 1864 (original description): 543; Bernhauer &amp; Scheerpeltz, 1926: 640 (catalogue, as synonym of  Atheta (Atheta) coriaria).</p><p>Homalota australis Jekel, 1873: 47 (original description); Bernhauer &amp; Scheerpeltz, 1926: 640 (catalogue, as a synonym of  Atheta (Atheta) coriaria).</p><p>Atheta (Atheta) coriaria: Bernhauer, 1907 (catalogue); Muona, 1979 (mentioned not belong to  Atheta): 48; Smetana, 2004: 374 (catalogue); Maruyama &amp; Kishimoto, 2013: 107 (catalogue); Schülke &amp; Smetana, 2015: 518 (catalogue).</p><p>Pseudota miscella Casey, 1910: 115 (original description); Gusarov, 2003 (synonymized with  Dalotia coriaria).</p><p>Dimetrota (Dalotia) pectorina Casey, 1910: 106 (original description); Gusarov, 2003 (synonymized with  Dalotia coriaria).</p><p>Dimetrota (Dalotia) crucialis Casey, 1910 (original description): 107; Gusarov, 2003 (synonymized with  Dalotia coriaria).</p><p>Atheta brachelytra Scheerpeltz, 1948: 194 (original description); Benick, 1973 (synonymized with  Atheta coriaria).</p><p>Atheta (Xenota) coriaria: Jarrige, 1957.</p><p>Atheta wynigeri Scheerpeltz, 1970: 49 (original description); Vogel, 2015 (synonymized with  Atheta (Atheta) coriaria).</p><p>Taxicera academica Yosii &amp; Sawada, 1976: 125 (original description); Muona, 1979: 48 (synonymized with  Atheta coriaria)).</p><p>Xenota coriaria: Seevers, 1978: 247 (catalogue).</p><p>Atheta coriaria: Ashe &amp; Watrous, 1984: 175 (description of larva).</p><p>Dalotia coriaria: Gusarov 2003: 89; Newton, 2024 (catalogue); Lee et al., 2024: 581 (redescription).</p><p>Materials examined.   Four males and six females,  Kamimaki, Ina city, Nagano Pref., Japan, 26.VII.2024, S. Hashizume (KUM) ;   two males and one female,  Daitô-jinja, Minamidaitô-jima Island, Okinawa Pref., Japan, 27. VI.2023, T.  Kinoda (KUM)  .</p><p>Diagnosis.  Dalotia coriaria differs from other species of  Dalotia by the following characteristics: apical margin of male tergite VIII dentate (fig. 55G in Yosii &amp; Sawada, 1976; 5A in Lee et al., 2024); median lobe of male aedeagus with large suspensoria and long sclerites (figs. 55K in Yosii &amp; Sawada, 1976; 6A &amp; B in Muona, 2023; vagina with two pairs of sclerites (Fig. 3D).</p><p>Additional description. Male. Tergites IX, X, and sternite IX shown in Figs. 3A &amp; B.</p><p>Female. Tergites IX, X, and sternite IX shown in Figs. 3C &amp; D. Vagina weakly sclerotized, semicircular, with two pairs of sclerite (Fig. 3D). Spermatheca: head elongate, without umbilicus; stem short (about twice as long as spermathecal head), helically coiled once around base (Fig. 3D).</p><p>Distribution. Japan: Honshu, Minamidaitô-jima Island (new record); cosmopolitan.</p><p>Habitat. This species is mainly collected from piled grasses.</p><p>Remarks. This species is distributed worldwide (Newton, 2024). From Japan, it is only known from Honshu. Here, we record this species from Minamidaitô-jima Island, located approximately 400 km east of the Okinawajima Is., for the first time.</p></div>	https://treatment.plazi.org/id/0B4787FCFF8D1A22FF29FBC13EA1FE14	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Tasaku, Yuto;Maruyama, Munetoshi	Tasaku, Yuto, Maruyama, Munetoshi (2025): Resolving the taxonomic confusion of Taxicera Mulsant & Rey, 1873 and Dalotia Casey, 1910 (Coleoptera, Staphylinidae, Aleocharinae), with the rediscovery of Dalotia pseudorphana (Cameron, 1933), comb. nov., from bird nest material. Zootaxa 5609 (1): 70-82, DOI: 10.11646/zootaxa.5609.1.4, URL: https://doi.org/10.11646/zootaxa.5609.1.4
0B4787FCFF8B1A2FFF29FE3F3ED3FD3C.text	0B4787FCFF8B1A2FFF29FE3F3ED3FD3C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dalotia pseudorphana (Cameron 1933) Tasaku & Maruyama 2025	<div><p>Dalotia pseudorphana (Cameron, 1933),  comb. nov.</p><p>(Figs. 1C–D, 2, 4–6)</p><p>Atheta (Acrotona) pseudorphana Cameron, 1933b: 216 (original description; type locality: Kobe, Harada).</p><p>Taxicera pseudorphana: Sawada, 1977: 214 (redescription); Newton, 2024 (catalogue).</p><p>Taxicera (Taxicera) pseudorphana: Smetana, 2004: 419 (catalogue); Schülke &amp; Smetana, 2015: 728 (catalogue); Maruyama &amp; Kishimoto, 2013 (catalogue).</p><p>Type material.   Holotype. Male, data on labels: “Kobe / Harad / 24.VI-12 / A. / pseudorphana / TYPE Cam. // Cameron / Bequest / B.M.1955-147 //  Atheta /  pseudorphana Cam. / P.M. Hammond / det. 1973 / HOLOTYPE ” (Fig. 2 dissected by K. Sawada; deposited in the Natural History Museum, London).</p><p>Specimens examined.   Two males, two females, and five unsexed, FPI (nest number K-9,  Parus varius), 13. VI.2023 (SEHU) ;   one male, one female, and four unsexed, TUS (nest number 22,  Parus minor), 4.VII.2023 (SEHU) ;   one female and one unsexed, TUS (nest number 64,  Parus minor), 19. VI  .2023 (KUM) .</p><p>Diagnosis.  Dalotia pseudorphana comb. nov. differs from other species of  Dalotia by the shape of male tergite VIII, and median lobe. From  Dalotia coriaria, this species easily differs by the smaller body (Figs 1C &amp; D), truncated apical margin of male tergite VIII (Fig. 3A), and vagina with spines (Fig. 5D).</p><p>Redescription. Body subparallel-sided, flat (Figs 1C &amp; D).</p><p>Coloration. Head and abdomen black; antennomeres III–XI, pronotum, and elytra reddish brown; antennomeres I and II, legs, and mouth parts yellow.</p><p>Head. Capsule circular, weakly shining; infraorbital carina absent; eyes large, longer than temporal length.</p><p>Antennae. Antennomeres I–III elongate, II slightly shorter than I, III as long as II, IV–X transverse, XI oval, twice as long as width.</p><p>Mouth parts. Labrum transverse, with slightly emarginated apical margin, with 8 macrosetae on each side. Epipharynx with setaceous α-sensillum, with very small β and γ-sensillum. Mandibles triangular, slender, almost symmetrical; light mandible with tiny tooth. Maxilla: maxillary palpus elongate, palpomere I very small, segment II about three times longer than I, III slightly longer than II, segment IV thin, as long as III. Mentum trapezoidal, with slightly emarginate apical margin. Labium (Fig. 6E): palpus with three palpomeres, chaetotaxy shown in Fig. 6F; prementum with narrow and smooth median area; ligula divided at apical half, with seta on each branch in ventral side; premental apodeme slender, as long as labial palpus.</p><p>Thorax. Pronotum oblong, wider than long, weakly shining; hypomera almost invisible in lateral view. Elytra longer than wide, densely covered with setae. Hind wing fully developed. Tarsal formula 4-5-5.</p><p>Abdomen strongly shining, sparsely covered with setae.</p><p>Male. Tergite VIII quadrate, with straight posterior margin, with 4+4 macrosetae (Fig. 4A). Sternite VIII semicircular, with 8+6 macrosetae (Fig. 4B). Tergites IX, X, and sternite X as shown in Figs. 5A &amp; B. Median lobe of aedeagus (Figs. 6A–C): apical lobe simple, slender, as long as basal capsule; basal capsule oval; athetine bridge thin, incomplete; internal structure simple. Parameres of aedeagus broad with slender apical lobe (Fig. 6D).</p><p>Female. Tergite VIII quadrate, with straight posterior margin, with 4+4 macrosetae (Fig. 4C). Sternite VIII semicircular; marginal setae thick, 4 setae of center long (Fig. 4D). Tergites IX, X, and sternite X shown in Figs. 5C &amp; D. Vagina weakly sclerotized, longer than wide, with about 40 spines on apical half (Fig. 5D). Spermatheca (Fig. 5E): head elongate, without umbilicus; stem short (about twice as long as spermathecal head), coiled once helically around base.</p><p>Measurements. BL, 1.70–2.14, FBL 0.96–1.28, HW 0.34–0.39, PW 0.47–0.49, PL 0.32–0.37, EW 0.54–0.60.</p><p>Distribution. Japan: Hokkaido (new record), Honshu.</p><p>Habitat. Bird nest materials ( Parus varius (Temminck &amp; Schlegel, 1848) and  Parus minor Temminck &amp; Schlegel, 1848).</p><p>Remarks. The holotype is in poor condition (whole body soaked with KOH and body color changed (Fig. 2A); mouth parts, abdominal segments VIII–X, and aedeagus were dissected and glued by the previous researcher (Fig. 2B)), so no further information is available. The identification was conducted based on illustrations and descriptions in Sawada (1977).</p><p>Only the holotype has been known for this species for nearly a hundred years. It is probably because this species is nidicolous and difficult to collect. As surveys progress, this species will be found in other East Asian areas. We could not find this species from nest materials of  Agropsar philippensis (J. R. Forster, 1781) and  Passer montanus (Linnaeus, 1758), composed by plants matter, particularly grasses.</p><p>Based on the morphological characteristics, this species should belong to  Dalotia, not  Taxicera Mulsant &amp; Rey, 1873 (see also Discussion).</p></div>	https://treatment.plazi.org/id/0B4787FCFF8B1A2FFF29FE3F3ED3FD3C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Tasaku, Yuto;Maruyama, Munetoshi	Tasaku, Yuto, Maruyama, Munetoshi (2025): Resolving the taxonomic confusion of Taxicera Mulsant & Rey, 1873 and Dalotia Casey, 1910 (Coleoptera, Staphylinidae, Aleocharinae), with the rediscovery of Dalotia pseudorphana (Cameron, 1933), comb. nov., from bird nest material. Zootaxa 5609 (1): 70-82, DOI: 10.11646/zootaxa.5609.1.4, URL: https://doi.org/10.11646/zootaxa.5609.1.4
0B4787FCFF861A2FFF29FB963E4EFA12.text	0B4787FCFF861A2FFF29FB963E4EFA12.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dalotia garuda (Sawada 1971) Tasaku & Maruyama 2025	<div><p>Dalotia garuda (Sawada, 1971),  comb. nov.</p><p>Ischnopoda (Oreostiba) garuda Sawada, 1971: 70 (original description; type locality: Mt. Panjerango (2800 m alt.), Java).</p><p>Taxicera garuda: Yosii &amp; Sawada, 1976: 127 (diagnosis); Newton, 2024 (catalogue).</p><p>Type material. Not examined.</p><p>Distribution. Indonesia (Java).</p><p>Remarks. This species was transferred to  Taxicera in Yosii &amp; Sawada (1976), based on morphological similarities with  T. academica (now junior synonym of  D. coriaria). This species should also belong to  Dalotia (see also Discussion).</p></div>	https://treatment.plazi.org/id/0B4787FCFF861A2FFF29FB963E4EFA12	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Tasaku, Yuto;Maruyama, Munetoshi	Tasaku, Yuto, Maruyama, Munetoshi (2025): Resolving the taxonomic confusion of Taxicera Mulsant & Rey, 1873 and Dalotia Casey, 1910 (Coleoptera, Staphylinidae, Aleocharinae), with the rediscovery of Dalotia pseudorphana (Cameron, 1933), comb. nov., from bird nest material. Zootaxa 5609 (1): 70-82, DOI: 10.11646/zootaxa.5609.1.4, URL: https://doi.org/10.11646/zootaxa.5609.1.4
0B4787FCFF861A2FFF29FD173CDCFBBF.text	0B4787FCFF861A2FFF29FD173CDCFBBF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dalotia pendleburyi (Cameron 1933) Tasaku & Maruyama 2025	<div><p>Dalotia pendleburyi (Cameron, 1933),  comb. nov.</p><p>Atheta (Atheta) pendleburyi Cameron, 1933a: 358 (original description; type locality: Lumu Lumu, 5500 feet).</p><p>Taxicera pendleburyi: Sawada, 1979: 341 (redescription; figures); Kapp, 2005: 1280 (catalogue); Newton, 2024 (catalogue).</p><p>Type material. Not examined.</p><p>Distribution. Malaysia: Sabah (Mt. Kinabalu).</p><p>Remarks. This species was transferred to  Taxicera in Sawada (1979). However, this species should be regarded as a species of  Dalotia considering his description (see also Discussion).</p></div>	https://treatment.plazi.org/id/0B4787FCFF861A2FFF29FD173CDCFBBF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Tasaku, Yuto;Maruyama, Munetoshi	Tasaku, Yuto, Maruyama, Munetoshi (2025): Resolving the taxonomic confusion of Taxicera Mulsant & Rey, 1873 and Dalotia Casey, 1910 (Coleoptera, Staphylinidae, Aleocharinae), with the rediscovery of Dalotia pseudorphana (Cameron, 1933), comb. nov., from bird nest material. Zootaxa 5609 (1): 70-82, DOI: 10.11646/zootaxa.5609.1.4, URL: https://doi.org/10.11646/zootaxa.5609.1.4
