taxonID	type	description	language	source
0176194EFFB4C23EFF494ECBFA9E0213.taxon	type_taxon	Type species: Agoo xavieri Bahder & Bartlett, 2019	en	Barrantes Barrantes, Edwin A., Zumbado Echavarria, Marco A., Bartlett, Charles R., Helmick, Ericka E., Bahder, Brian W. (2025): Multi-local analysis supports the transfer of Omolicna fulva to the genus Agoo and establishment of a new species in the genus from Costa Rica. Zootaxa 5584 (1): 113-125, DOI: 10.11646/zootaxa.5584.1.7, URL: https://doi.org/10.11646/zootaxa.5584.1.7
0176194EFFB4C23EFF494ECBFA9E0213.taxon	diagnosis	Diagnosis. Elongate cenchreines with narrow forewings. Head in lateral view narrowly projected anteriorly, with vertex + frons profile smoothly rounded. Frons narrow (but lateral margins not in contact), vertex subtriangular, with sensorial pits along lateral margins of frons and vertex; transverse carina at fastigium lacking; median carina of vertex and frons absent. Antennae short. Paranotal expansions behind antennae quadrate to semiquadrate in frontal view. In caudoventral view, medioventral lobe of pygofer simple, broad, subtriangular, distally attenuating to rounded apex. Aedeagus and endosoma nearly bilaterally symmetrical, aedeagus with at least one pair of elongate processes at apex, extremely complex endosoma with at least two pairs of large sclerotized processes.	en	Barrantes Barrantes, Edwin A., Zumbado Echavarria, Marco A., Bartlett, Charles R., Helmick, Ericka E., Bahder, Brian W. (2025): Multi-local analysis supports the transfer of Omolicna fulva to the genus Agoo and establishment of a new species in the genus from Costa Rica. Zootaxa 5584 (1): 113-125, DOI: 10.11646/zootaxa.5584.1.7, URL: https://doi.org/10.11646/zootaxa.5584.1.7
0176194EFFB4C23BFF4948F4FE3B0683.taxon	description	(Figures 2 – 6)	en	Barrantes Barrantes, Edwin A., Zumbado Echavarria, Marco A., Bartlett, Charles R., Helmick, Ericka E., Bahder, Brian W. (2025): Multi-local analysis supports the transfer of Omolicna fulva to the genus Agoo and establishment of a new species in the genus from Costa Rica. Zootaxa 5584 (1): 113-125, DOI: 10.11646/zootaxa.5584.1.7, URL: https://doi.org/10.11646/zootaxa.5584.1.7
0176194EFFB4C23BFF4948F4FE3B0683.taxon	materials_examined	Type Locality. Costa Rica, Puntarenas Province, Osa Peninsula.	en	Barrantes Barrantes, Edwin A., Zumbado Echavarria, Marco A., Bartlett, Charles R., Helmick, Ericka E., Bahder, Brian W. (2025): Multi-local analysis supports the transfer of Omolicna fulva to the genus Agoo and establishment of a new species in the genus from Costa Rica. Zootaxa 5584 (1): 113-125, DOI: 10.11646/zootaxa.5584.1.7, URL: https://doi.org/10.11646/zootaxa.5584.1.7
0176194EFFB4C23BFF4948F4FE3B0683.taxon	diagnosis	Diagnosis. Moderate sized 6.7 – 7.6 mm, body pallid to yellow, wings pale translucent with darker yellow band along trailing margin. Medioventral process of pygofer broadly triangular, wider than tall. Processes on inner margin of gonostyli in ventral view roughly quadrate, arising in distal half. Aedeagus bearing large, subtriangular processes on dorsolateral margins of shaft. Endosoma with four pairs of processes, bearing triangular projection on dorsal margin near apex and large pair of processes curved cephalad from apex. Anal tube broadly triangular, apex caudally projected (not downcurved), apex strongly bilobed from dorsal view.	en	Barrantes Barrantes, Edwin A., Zumbado Echavarria, Marco A., Bartlett, Charles R., Helmick, Ericka E., Bahder, Brian W. (2025): Multi-local analysis supports the transfer of Omolicna fulva to the genus Agoo and establishment of a new species in the genus from Costa Rica. Zootaxa 5584 (1): 113-125, DOI: 10.11646/zootaxa.5584.1.7, URL: https://doi.org/10.11646/zootaxa.5584.1.7
0176194EFFB4C23BFF4948F4FE3B0683.taxon	description	Description. Color. Body pallid (nearly white) to yellow (Fig. 2), darker dorsally; mesonotum white to yellow (when yellow, pale on lateral margins, near tegula); legs pale or yellow and pale proximally (coxae to midlength of femur white). Forewings translucent, mostly clear to strongly washed with yellow (especially dorsally). Structure. Body length: males with wings 6.7 – 7.2 mm, body length (without wings) 2.8 – 3.2 mm; females with wings 7.2 – 7.6 mm, body length 3.2 – 3.6 mm (Table 3). Head. Head narrower than pronotum. Vertex in dorsal view triangular, widest posteriorly; lateral margins foliate, bearing single row of sensorial pits, disc depressed, median carina absent; anterior margin narrow, concave (apical transverse carina absent); posterior margin quadrately concave (Fig. 3 A); in lateral view, profile smoothly rounded from posterior margin of vertex to frontoclypeal sutures (Fig. 3 B). Frons in frontal view with lateral margins foliate and slightly sinuate, bearing single row of sensorial pits (Fig. 3 C), disc depressed median carina absent. Clypeus elongate-triangular. Lateral margins carinate (not foliate), bearing distinct median carina. Eye hemispherical, emarginated at antenna. Lateral ocelli distinct below eye near midline. Antennal scape short, hidden, pedicel ovate mearing many irregularly arranged sensory plaques. Thorax. Pronotum in dorsal view roughly trapezoidal (about equal in length as vertex at midline), anterior median portion (between eyes), quadrate with anterior margin linear, posterior margin concave, median carina obscure, lateral carinae appear concurrent with anterior lateral pronotal margin; paradiscal broadly foliate behind antennae forming large fossae, partially surrounding antennae. Mesonotum at midline exceeding length of pronotum and vertex combined, tricarinate, median carina becoming obsolete posteriorly, lateral carinae medially curved from anterior margin, approximately reaching hind margin (Fig. 3 A). Hind tibiae unarmed laterally, with apical spine ornamentation of 7 - 6 - 5. Forewing elongate oval; apex of clavus near wing midlength; MP fused with ScP + R to form short composite vein from apex of basal cell: Fork of RP from ScP + RA nearly at same level of fusion of Pcu + 1 A in clavus, these just proximad of CuA fork; branching pattern: RA 1 - branched, RP 2 - branched, MP 4 - branched, CuA 2 - branched. Male Terminalia. Pygofer in lateral view narrow and irregular in shape; narrowest dorsally, widest ventrally, anterior and posterior margin irregularly sinuate (Fig. 5 A); in ventral view, medioventral process subtriangular (Fig. 5 B), broader than tall with acutely pointed apex. Gonostyli in lateral view spatulate, apex broadly rounded (Fig. 5 A); dorsal process small, bilobed, distal lobe sclerotized; in ventral view (Fig. 5 B), median margin bearing large, quadrate process just distad midlength, distally narrowed with apices medially hooked to blunt apices. Aedeagus simple (Fig. 6), approximately bilaterally symmetrical, shaft upcurved bearing a pair of apical processes (A 1 & A 2), elongated and slender, gently curving ventrad, angled cephalad, approximately reaching midpoint; second pair large, subtriangular processes (A 3 & A 4) arising on dorsolateral margins in basal 1 / 3 of shaft (Fig. 6 A, B), ventral margin serrate along midlength, endosoma with multiple processes present, first pair (E 1 & E 2) arising subapically on lateral margins, gently curved dorsad, angled cephalad, reaching apex of A 3 and A 4; second pair (E 3 & E 4) arising on dorsal margin along inner margin, nearly straight, angled directly caudad, extending just beyond aedeagal apex; third pair (E 5 & E 6) arising on endosomal ventral margin, sinuate along basal region, angled dorsad and cephalad, curving directly dorsad at apex; and the fourth pair (E 7 & E 8) arising at endosoma apex, immediately curving dorsad, approximately reaching apex of E 5 and E 6. Anal tube in lateral view roughly triangular and relatively broad, posteriorly projecting (not downcurved), narrowed to blunt apices; in ventral view broad, strongly bilobed distad of epiproct and paraproct; paraproct short. Plant Associations. Sweeping undetermined palm.	en	Barrantes Barrantes, Edwin A., Zumbado Echavarria, Marco A., Bartlett, Charles R., Helmick, Ericka E., Bahder, Brian W. (2025): Multi-local analysis supports the transfer of Omolicna fulva to the genus Agoo and establishment of a new species in the genus from Costa Rica. Zootaxa 5584 (1): 113-125, DOI: 10.11646/zootaxa.5584.1.7, URL: https://doi.org/10.11646/zootaxa.5584.1.7
0176194EFFB4C23BFF4948F4FE3B0683.taxon	distribution	Distribution. Costa Rica (Puntarenas Province).	en	Barrantes Barrantes, Edwin A., Zumbado Echavarria, Marco A., Bartlett, Charles R., Helmick, Ericka E., Bahder, Brian W. (2025): Multi-local analysis supports the transfer of Omolicna fulva to the genus Agoo and establishment of a new species in the genus from Costa Rica. Zootaxa 5584 (1): 113-125, DOI: 10.11646/zootaxa.5584.1.7, URL: https://doi.org/10.11646/zootaxa.5584.1.7
0176194EFFB4C23BFF4948F4FE3B0683.taxon	etymology	Etymology. The specific name is given in honor of Dr. Clifford Keil (retired faculty from the University of Delaware Department of Entomology and retired faculty member and director of the Museum of Invertebrates in the School of Biological Sciences, Pontifical Catholic University of Ecuador in Quito).	en	Barrantes Barrantes, Edwin A., Zumbado Echavarria, Marco A., Bartlett, Charles R., Helmick, Ericka E., Bahder, Brian W. (2025): Multi-local analysis supports the transfer of Omolicna fulva to the genus Agoo and establishment of a new species in the genus from Costa Rica. Zootaxa 5584 (1): 113-125, DOI: 10.11646/zootaxa.5584.1.7, URL: https://doi.org/10.11646/zootaxa.5584.1.7
0176194EFFB4C23BFF4948F4FE3B0683.taxon	materials_examined	Material Examined. Holotype male, “ Costa Rica, Puntarenas Pr. / Hotel Keili, roadside / 09 - VI- 2021 / Coll.: B. W. Bahder / Sweeping palms / Agoo keili ♂ ” (FLREC). Paratypes same as holotype (11 males, 4 females; 2 males and 1 female were yellow morph and 9 males and 1 female were pale morph.	en	Barrantes Barrantes, Edwin A., Zumbado Echavarria, Marco A., Bartlett, Charles R., Helmick, Ericka E., Bahder, Brian W. (2025): Multi-local analysis supports the transfer of Omolicna fulva to the genus Agoo and establishment of a new species in the genus from Costa Rica. Zootaxa 5584 (1): 113-125, DOI: 10.11646/zootaxa.5584.1.7, URL: https://doi.org/10.11646/zootaxa.5584.1.7
0176194EFFBFC234FF494EC8FD9702E7.taxon	materials_examined	“ U. S. A., FL, Miami-Dade Co. / Montgomery Botanical Center / 11 - XI- 2022 / Coll.: B. W. Bahder / Sweeping palms / Agoo fulvus ♂ ” (FLREC). “ U. S. A., FL, Lee Co., Estero / 12. VI. 1980 / Van Duzee ” (Holotype male).	en	Barrantes Barrantes, Edwin A., Zumbado Echavarria, Marco A., Bartlett, Charles R., Helmick, Ericka E., Bahder, Brian W. (2025): Multi-local analysis supports the transfer of Omolicna fulva to the genus Agoo and establishment of a new species in the genus from Costa Rica. Zootaxa 5584 (1): 113-125, DOI: 10.11646/zootaxa.5584.1.7, URL: https://doi.org/10.11646/zootaxa.5584.1.7
0176194EFFBFC234FF494EC8FD9702E7.taxon	description	Sequence Data. For A. keili sp. nov. molecular data for the COI, 18 S and D 9 - D 10 expansion regions of 28 S were generated and accessioned (Table 2). The molecular phylogeny generated based on 18 S, 28 S and COI show strong bootstrap support (100, 100 and 93, respectively) for the monophyly of Agoo, with A. keili sp. nov. resolving within the clade for all markers (Fig. 7). In addition, the consensus tree generated based on concatenated 18 S, 28 S and COI data also demonstrated strong bootstrap support for the monophyly of Agoo with A. keili sp. nov. resolving within Agoo adjacent to A. luzdenia with strong bootstrap support (98).	en	Barrantes Barrantes, Edwin A., Zumbado Echavarria, Marco A., Bartlett, Charles R., Helmick, Ericka E., Bahder, Brian W. (2025): Multi-local analysis supports the transfer of Omolicna fulva to the genus Agoo and establishment of a new species in the genus from Costa Rica. Zootaxa 5584 (1): 113-125, DOI: 10.11646/zootaxa.5584.1.7, URL: https://doi.org/10.11646/zootaxa.5584.1.7
0176194EFFBFC234FF494EC8FD9702E7.taxon	discussion	Remarks. The morphological characters observed (rounded / circular head shape in lateral, general wing venation and spinulation of aedeagus with large, apical processes of endosoma and basal retrose processes) support the placement of the novel taxon in the genus Agoo. Further supported by the strong molecular evidence for three separate loci. In addition, the similar morphological characters observed in A. keili sp. nov. along with strong molecular support indicate that the new species be placed in Agoo. Initially, the pale form and yellow form were sorted as two different species due to general color and pattern, thus far, being reliable for species diagnosis. However, after examination of the genitalia it was evident that they were the same. In addition, molecular data was generated for both forms and were 100 % identical for all three loci (data not shown). It is unknown if this is due to the presence of true color morphs or is potentially age related, but not observing this degree of color variance in the large number of specimens examined for other species of Agoo in previous studies make it likely the variance observed in A. keili sp. nov. is the presence of true color morphs. Agoo keili sp. nov. can be recognized as a pale species, lacking strong markings. Pygofer of male terminalia bearing a large broadly triangular medioventral process. Gonostyli in ventral view bearing roughly quadrate process just distad of midlength. Aedeagus bearing large, subtriangular processes on dorsolateral margins of the shaft. Endosoma bearing 5 pairs of processes, with triangular projection on dorsal margin near apex. Anal segment tube broadly triangular, with apex caudally projected (not downcurved), lobed, not downcurved apex strongly bilobed from dorsal view. The most similar species might be the Jamaican species A. beani, which is roughly similar in color and shares some male genitalic structures but the dorsolateral margins of the shaft bear elongated processes (not subtriangular), the endosoma bears fewer processes and lacks the triangular projection on dorsal margin near apex; also the median processes on the gonostyli (ventral view) are roughly triangular (not quadrate). In addition, it also superficially resembles A. xavieri in general body color, however, A. xavieri possesses a fuscous stripe along the length of the forewings. While the general scheme of the pygofer, gonostyli and anal segment resembles A. xavieri, the aedeagus is significantly different. Interestingly, based on the molecular evidence at hand, A. keili sp. nov. appears mostly closely allied to A. luzdenia, however is easily distinguished from it based solely on wing pattern (A. keili sp. nov. lacking patterns and A. luzdenia possessing a half fuscous, half red stripe along length of forewing with a black spot near the apex).	en	Barrantes Barrantes, Edwin A., Zumbado Echavarria, Marco A., Bartlett, Charles R., Helmick, Ericka E., Bahder, Brian W. (2025): Multi-local analysis supports the transfer of Omolicna fulva to the genus Agoo and establishment of a new species in the genus from Costa Rica. Zootaxa 5584 (1): 113-125, DOI: 10.11646/zootaxa.5584.1.7, URL: https://doi.org/10.11646/zootaxa.5584.1.7
0176194EFFBEC237FF494CEFFB2C012A.taxon	description	(Fig. 8)	en	Barrantes Barrantes, Edwin A., Zumbado Echavarria, Marco A., Bartlett, Charles R., Helmick, Ericka E., Bahder, Brian W. (2025): Multi-local analysis supports the transfer of Omolicna fulva to the genus Agoo and establishment of a new species in the genus from Costa Rica. Zootaxa 5584 (1): 113-125, DOI: 10.11646/zootaxa.5584.1.7, URL: https://doi.org/10.11646/zootaxa.5584.1.7
0176194EFFBEC237FF494CEFFB2C012A.taxon	discussion	Sequence data. For O. fulva, molecular data for the COI, 18 S and D 9 - D 10 expansion regions of 28 S were generated and accessioned (Table 2). The molecular phylogeny generated based on 18 S, 28 S and COI demonstrated strong bootstrap support (100, 100 and 93, respectively) for the monophyly of Agoo, with O. fulva resolving within Agoo the clade for all markers (Fig. 7), indicating that this species is misplaced to genus. Remarks. Morphological features (Fig. 8) appear consistent with the hypothesis that O. fulva belongs in Agoo rather than Omolicna Fennah. Here we move O. fulva to Agoo as A. fulvus, new combination.	en	Barrantes Barrantes, Edwin A., Zumbado Echavarria, Marco A., Bartlett, Charles R., Helmick, Ericka E., Bahder, Brian W. (2025): Multi-local analysis supports the transfer of Omolicna fulva to the genus Agoo and establishment of a new species in the genus from Costa Rica. Zootaxa 5584 (1): 113-125, DOI: 10.11646/zootaxa.5584.1.7, URL: https://doi.org/10.11646/zootaxa.5584.1.7
