taxonID	type	description	language	source
03BDF65F0602CC63FF00432694AB0858.taxon	materials_examined	Material examined. Iran, Guilan province, Rezvanshahr, Paresar, Sandian, (37 ° 57 ′ 22.70 ″ N, 49 ° 12 ′ 82.95 ″ E), 02. ix. 2019, 1 ♀; on pupae of Hyphantria cunea, leg. A. Karami.	en	Karami, Amene, Talebi, Ali Asghar, Gilasian, Ebrahim, Fathipour, Yaghoub, Mehrabadi, Mohammad (2023): Native parasitoids of the fall webworm, Hyphantria cunea (Drury, 1773) (Lepidoptera, Erebidae), an invasive alien pest in northern Iran. Journal of Insect Biodiversity and Systematics 9 (1): 81-101, DOI: 10.52547/jibs.9.1.81, URL: https://doi.org/10.52547/jibs.9.1.81
03BDF65F0602CC63FF00432694AB0858.taxon	distribution	Distribution in Iran. Kerman (Lotfalizadeh et al., 2012) and Guilan (current study) provinces. General distribution. Australia (Queensland), Bangladesh, China (Beijing, Fujian, Guangxi, Guizhou, Hong Kong, Hubei, Hunan, Jiangsu, Shanxi, Sichuan, Zhejiang), Fiji, Guam, Hawaii, India (Andaman and Nicobar Islands, Arunachal Pradesh, Assam, Bihar, Delhi, Gujarat, Himachal Pradesh, Jammu and Kashmir, Karnataka, Kerala, Madhya Pradesh, Maharashtra, Meghalaya, Punjab, Tripura, Uttar Pradesh, West Bengal), Indochina, Indonesia (Java, West Papua), Iran, Japan ,, Malaysia (Sarawak), North Africa, Pakistan, Palau, Papua New Guinea, Philippines, South Korea, Taiwan, United States of America (Connecticut, Pennsylvania, Wisconsin), Vietnam (Devi & Singh, 2002; Konno et al., 2002; Gupta & Kalesh, 2012; Lotfalizadeh et al., 2012; Gupta et al., 2014; Narendran & van Achterberg, 2016; Noyes, 2019) (Fig. 4 A).	en	Karami, Amene, Talebi, Ali Asghar, Gilasian, Ebrahim, Fathipour, Yaghoub, Mehrabadi, Mohammad (2023): Native parasitoids of the fall webworm, Hyphantria cunea (Drury, 1773) (Lepidoptera, Erebidae), an invasive alien pest in northern Iran. Journal of Insect Biodiversity and Systematics 9 (1): 81-101, DOI: 10.52547/jibs.9.1.81, URL: https://doi.org/10.52547/jibs.9.1.81
03BDF65F0602CC63FF00432694AB0858.taxon	biology_ecology	Hosts. Lepidoptera: Hyphantria cunea (Drury, 1773), Lymantria dispar (Linnaeus, 1758) (Erebidae); Homona magnanima Diakonoff, 1948 (Tortricidae); Opisina arenosella Walker, 1864 (Oecophoridae) (Mao & Kunimi, 1994; Noyes, 2019).	en	Karami, Amene, Talebi, Ali Asghar, Gilasian, Ebrahim, Fathipour, Yaghoub, Mehrabadi, Mohammad (2023): Native parasitoids of the fall webworm, Hyphantria cunea (Drury, 1773) (Lepidoptera, Erebidae), an invasive alien pest in northern Iran. Journal of Insect Biodiversity and Systematics 9 (1): 81-101, DOI: 10.52547/jibs.9.1.81, URL: https://doi.org/10.52547/jibs.9.1.81
03BDF65F0603CC63FFF347F9944D0EB6.taxon	materials_examined	Material examined. Iran, Guilan province, Rezvanshahr, Paresar, Sandian, (37 ° 57 ′ 22.70 ″ N, 49 ° 12 ′ 82.95 ″ E), 02. ix. 2019, 18 ♂, 712 ♀; gregarious parasitoid on pupae of Hyphantria cunea, leg. A. Karami.	en	Karami, Amene, Talebi, Ali Asghar, Gilasian, Ebrahim, Fathipour, Yaghoub, Mehrabadi, Mohammad (2023): Native parasitoids of the fall webworm, Hyphantria cunea (Drury, 1773) (Lepidoptera, Erebidae), an invasive alien pest in northern Iran. Journal of Insect Biodiversity and Systematics 9 (1): 81-101, DOI: 10.52547/jibs.9.1.81, URL: https://doi.org/10.52547/jibs.9.1.81
03BDF65F0603CC63FFF347F9944D0EB6.taxon	distribution	Distribution in Iran. Guilan (Rezaei et al. 2003, current study) General distribution. China (Beijing, Shaanxi, Shandong, Zhejiang), Georgia, Iran, Italy, Japan, South Korea and Turkey (Yang, 1989; Boriani, 1994; Yang et al., 2006; Zhu & Huang, 2001; Konno et al., 2002; Rezaei et al., 2003; Japoshvili et al., 2006; Sullivan et al., 2011; Noyes, 2019) (Fig. 4 B).	en	Karami, Amene, Talebi, Ali Asghar, Gilasian, Ebrahim, Fathipour, Yaghoub, Mehrabadi, Mohammad (2023): Native parasitoids of the fall webworm, Hyphantria cunea (Drury, 1773) (Lepidoptera, Erebidae), an invasive alien pest in northern Iran. Journal of Insect Biodiversity and Systematics 9 (1): 81-101, DOI: 10.52547/jibs.9.1.81, URL: https://doi.org/10.52547/jibs.9.1.81
03BDF65F0603CC63FFF347F9944D0EB6.taxon	biology_ecology	Hosts. Lepidoptera: Hyphantria cunea (Drury, 1773), Lymantria dispar (Linnaeus, 1758), Stilpnotia salicis (Linnaeus, 1758), S. candida Staudinger, 1892, Ivela ochropoda (Eversmann, 1847) (Erebidae); Heliothis armigera (Hubner, 1808) (Noctuidae); Dendrolimus spectabilis Butler, 1877 (Lasiocampidae); Clostera anachoreta (Denis & Schiffermüller, 1775), Micromelalopha troglodyta Graeser, 1890 (Notodontidae); Clania variegata (Psychidae); Ostrinia furnacalis Guenée, 1854 (Crambidae); Antheraea pernyi (Guerin-Meneville, 1855) (Saturniidae) and Tachinidae (Diptera) attacking Hyphantria cunea (Yang, 1989; Boriani, 1991; Noyes, 2019).	en	Karami, Amene, Talebi, Ali Asghar, Gilasian, Ebrahim, Fathipour, Yaghoub, Mehrabadi, Mohammad (2023): Native parasitoids of the fall webworm, Hyphantria cunea (Drury, 1773) (Lepidoptera, Erebidae), an invasive alien pest in northern Iran. Journal of Insect Biodiversity and Systematics 9 (1): 81-101, DOI: 10.52547/jibs.9.1.81, URL: https://doi.org/10.52547/jibs.9.1.81
03BDF65F0603CC63FFF340D690C80D80.taxon	materials_examined	Material examined. Iran, Guilan province, Rezvanshahr, Paresar, Sandian, (37 ° 57 ′ 22.70 ″ N, 49 ° 12 ′ 82.95 ″ E), 02. ix. 2019, 203 ♂, 262 ♀; on pupae of Hyphantria cunea, leg. A. Karami.	en	Karami, Amene, Talebi, Ali Asghar, Gilasian, Ebrahim, Fathipour, Yaghoub, Mehrabadi, Mohammad (2023): Native parasitoids of the fall webworm, Hyphantria cunea (Drury, 1773) (Lepidoptera, Erebidae), an invasive alien pest in northern Iran. Journal of Insect Biodiversity and Systematics 9 (1): 81-101, DOI: 10.52547/jibs.9.1.81, URL: https://doi.org/10.52547/jibs.9.1.81
03BDF65F0603CC63FFF340D690C80D80.taxon	distribution	Distribution in Iran. Guilan (Rezaei et al., 2003; current study). General distribution. Austria, Azerbaijan, Belgium, Canada (Ontario, Quebec), Croatia, Czech Republic, Czechoslovakia, France, Germany, Hungary, Iran, Macedonia, Moldova, the Netherlands, North Africa, Poland, Romania, Russia (Adygey AO, Karachai-Cherkess AR, Kostroma Oblast, Rostov Oblast), Serbia, Slovakia, Spain, Sweden, Transcaucasus, European part of Turkey, Ukraine, United Kingdom, USA (Colorado, Connecticut, Delaware, Maine, Massachusetts, New Hampshire, New Jersey, New York, Ohio, Wisconsin), former Yugoslavia (Sharov & Izhevskiy, 1987; Rezaei et al., 2003; Noyes, 2019; Rahamani et al., 2022) (Fig. 4 C).	en	Karami, Amene, Talebi, Ali Asghar, Gilasian, Ebrahim, Fathipour, Yaghoub, Mehrabadi, Mohammad (2023): Native parasitoids of the fall webworm, Hyphantria cunea (Drury, 1773) (Lepidoptera, Erebidae), an invasive alien pest in northern Iran. Journal of Insect Biodiversity and Systematics 9 (1): 81-101, DOI: 10.52547/jibs.9.1.81, URL: https://doi.org/10.52547/jibs.9.1.81
03BDF65F0603CC63FFF340D690C80D80.taxon	biology_ecology	Hosts. Lepidoptera: Hyphantria cunea (Drury, 1773), Arctia caja (Linnaeus, 1758), Panaxia dominula Linnaeus, 1758 (Erebidae); Cerura Vinula (Linnaeus, 1758) (Notodontidae) (Askew, 1970; Noyes, 2019).	en	Karami, Amene, Talebi, Ali Asghar, Gilasian, Ebrahim, Fathipour, Yaghoub, Mehrabadi, Mohammad (2023): Native parasitoids of the fall webworm, Hyphantria cunea (Drury, 1773) (Lepidoptera, Erebidae), an invasive alien pest in northern Iran. Journal of Insect Biodiversity and Systematics 9 (1): 81-101, DOI: 10.52547/jibs.9.1.81, URL: https://doi.org/10.52547/jibs.9.1.81
03BDF65F0604CC64FF0044D190F30F99.taxon	materials_examined	Material examined. Iran, Guilan province, Rezvanshahr, Paresar, Sandian, (37 ° 57 ′ 22.70 ″ N, 49 ° 12 ′ 82.95 ″ E), 02. ix. 2019, 1 ♂; on pupae of Hyphantria cunea, leg. A. Karami.	en	Karami, Amene, Talebi, Ali Asghar, Gilasian, Ebrahim, Fathipour, Yaghoub, Mehrabadi, Mohammad (2023): Native parasitoids of the fall webworm, Hyphantria cunea (Drury, 1773) (Lepidoptera, Erebidae), an invasive alien pest in northern Iran. Journal of Insect Biodiversity and Systematics 9 (1): 81-101, DOI: 10.52547/jibs.9.1.81, URL: https://doi.org/10.52547/jibs.9.1.81
03BDF65F0604CC64FF0044D190F30F99.taxon	distribution	Distribution in Iran. East Azerbaijan (Kolarov & Ghahari, 2006; Masnadi-Yazdinejad & Jussila, 2008), Fars (Masnadi-Yazdinejad & Jussila, 2008), Golestan (Kolarov & Ghahari, 2006), Guilan (Mohammadi-Khoramabadi et al., 2013; current study), Kerman (Mohammadi-Khoramabadi et al., 2014), Mazandaran (Kolarov & Ghahari, 2006), Tehran (Masnadi-Yazdinejad & Jussila, 2008; Mohammadi-Khoramabadi et al., 2013; Ghahari & Gadallah, 2017) and West Azerbaijan (Masnadi-Yazdinejad & Jussila, 2008; Karimpour, 2018). General distribution. Afghanistan, Algeria, Argentina, Armenia, Austria, Azerbaizan, Brazil, Bulgaria, China, Cyprus, Egypt, Georgia, Germany, India, Iran, Italy, Japan, Kazakhstan, Korea, Kyrgyzstan, Libya, Mongolia, Montenegro, Morocco, Poland, Romania, Russia, Sri Lanka, Tajikistan, Tunisia, Turkey, Turkmenistan, Vietnam, Ukraine, United States of America, Uruguay, Uzbekistan, former Yugoslavia (Atanassov, 1978; Coulson et al., 1986; Horstmann, 2001; Çoruh & Özbek, 2008; Masnadi-Yazdinejad & Jussila, 2008; Sullivan et al., 2010; Özbek & Coruh, 2012; Çoruh et al., 2014; Di Giovanni et al., 2015; Choi et al., 2016; Yu et al., 2016; Piekarska-Boniecka et al., 2018) (Fig. 4 D).	en	Karami, Amene, Talebi, Ali Asghar, Gilasian, Ebrahim, Fathipour, Yaghoub, Mehrabadi, Mohammad (2023): Native parasitoids of the fall webworm, Hyphantria cunea (Drury, 1773) (Lepidoptera, Erebidae), an invasive alien pest in northern Iran. Journal of Insect Biodiversity and Systematics 9 (1): 81-101, DOI: 10.52547/jibs.9.1.81, URL: https://doi.org/10.52547/jibs.9.1.81
03BDF65F0604CC64FF0044D190F30F99.taxon	biology_ecology	Hosts. Lepidoptera: Hyphantria cunea (Drury, 1773), Lymantria dispar (Linnaeus, 1758), L. monacha (Linnaeus, 1758), Arctia caja (Linnaeus, 1758), Orgyia trigotephras Boisduval, 1829, Calliteara fascelina (Linnaeus, 1758) (Erebidae); Malacosoma castrensis (Linnaeus, 1758), M. franconica (Denis & Schiffermüller, 1775), M. neustria (Linnaeus, 1758) (Lasiocampidae); Cydia pomonella (Linnaeus, 1758), Archips rosana (Linnaeus, 1758) (Tortricidae); Pieris brassicae (Linnaeus, 1758), P. rapae (Linnaeus, 1758), P. napi (Linnaeus, 1758), Colias croceus (Fourcroy, 1785), (Pieridae); Papilio machaon Linnaeus, 1758 (Papilionidae); Lacanobia oleracea (Linnaeus, 1758), Acronicta rumicis (Linnaeus, 1758) (Noctuidae); Charaxes jasius (Linnaeus, 1767), Eurodryas aurinia Rottemburg, 1775, Vanessa atalanta (Linnaeus, 1758) (Nymphalidae); Lycaena dispar (Haworth, 1802), Polyommatus bellargus (Rottemburg, 1775) (Lycaenidae); Coleoptera: Lixus iridis Olivier, 1807 (Curculionidae) (Atanassov, 1978; Okyar & Yurtcan, 2007; Çoruh & Özbek, 2008; Masnadi-Yazdinejad & Jussila, 2008; Shaw et al., 2009; Stefanescu et al., 2009; Sullivan et al., 2010; Mohammadi-Khoramabadi et al., 2013; Yu et al., 2016; Karimpour, 2018; Mifsud et al., 2019).	en	Karami, Amene, Talebi, Ali Asghar, Gilasian, Ebrahim, Fathipour, Yaghoub, Mehrabadi, Mohammad (2023): Native parasitoids of the fall webworm, Hyphantria cunea (Drury, 1773) (Lepidoptera, Erebidae), an invasive alien pest in northern Iran. Journal of Insect Biodiversity and Systematics 9 (1): 81-101, DOI: 10.52547/jibs.9.1.81, URL: https://doi.org/10.52547/jibs.9.1.81
03BDF65F0604CC64FF00436F949F026D.taxon	materials_examined	Material examined. Iran, Guilan province, Rezvanshahr, Paresar, Sandian, (37 ° 57 ′ 22.70 ″ N, 49 ° 12 ′ 82.95 ″ E), 02. ix. 2019, 2 ♂, 3 ♀; on pupae of Hyphantria cunea, leg. A. Karami.	en	Karami, Amene, Talebi, Ali Asghar, Gilasian, Ebrahim, Fathipour, Yaghoub, Mehrabadi, Mohammad (2023): Native parasitoids of the fall webworm, Hyphantria cunea (Drury, 1773) (Lepidoptera, Erebidae), an invasive alien pest in northern Iran. Journal of Insect Biodiversity and Systematics 9 (1): 81-101, DOI: 10.52547/jibs.9.1.81, URL: https://doi.org/10.52547/jibs.9.1.81
03BDF65F0604CC64FF00436F949F026D.taxon	distribution	Distribution in Iran. Golestan (Riedel & Aghadokht, 2017), Guilan (Shirzadegan et al., 2021; current study), West Azarbaijan (Riedel & Aghadokht, 2017). General distribution. Austria, Azerbaijan, Belarus, Belgium, China (Fujian), Denmark, France, Germany, Hungary, Iran, Italy, Moldova, Netherlands, Poland, Romania, Russia (Astrakhanskaya Oblast, Gorno-Altay, Krasnodar Kray, Yaroslavl Oblast), Spain, Sweden, Turkey, United Kingdom (Thirion, 2005; Tereshkin, 2006; Horstmann, 2008; Sullivan et al., 2010; Riedel & Magnusson, 2014; Yu et al., 2016) (Fig. 4 E).	en	Karami, Amene, Talebi, Ali Asghar, Gilasian, Ebrahim, Fathipour, Yaghoub, Mehrabadi, Mohammad (2023): Native parasitoids of the fall webworm, Hyphantria cunea (Drury, 1773) (Lepidoptera, Erebidae), an invasive alien pest in northern Iran. Journal of Insect Biodiversity and Systematics 9 (1): 81-101, DOI: 10.52547/jibs.9.1.81, URL: https://doi.org/10.52547/jibs.9.1.81
03BDF65F0604CC64FF00436F949F026D.taxon	biology_ecology	Hosts. Lepidoptera: Hyphantria cunea (Drury, 1773) (Erebidae) (Sharov & Izhevskiy, 1987; Tolkanitz, 1990; Sullivan et al., 2010)	en	Karami, Amene, Talebi, Ali Asghar, Gilasian, Ebrahim, Fathipour, Yaghoub, Mehrabadi, Mohammad (2023): Native parasitoids of the fall webworm, Hyphantria cunea (Drury, 1773) (Lepidoptera, Erebidae), an invasive alien pest in northern Iran. Journal of Insect Biodiversity and Systematics 9 (1): 81-101, DOI: 10.52547/jibs.9.1.81, URL: https://doi.org/10.52547/jibs.9.1.81
03BDF65F0606CC66FF004480966D0F83.taxon	materials_examined	Material examined. Iran, Guilan province, Rezvanshahr, Paresar, Sandian, (37 ° 57 ′ 22.70 ″ N, 49 ° 12 ′ 82.95 ″ E), 02. ix. 2019, 190 ♂, 222 ♀; on larvae of Hyphantria cunea, leg. A. Karami.	en	Karami, Amene, Talebi, Ali Asghar, Gilasian, Ebrahim, Fathipour, Yaghoub, Mehrabadi, Mohammad (2023): Native parasitoids of the fall webworm, Hyphantria cunea (Drury, 1773) (Lepidoptera, Erebidae), an invasive alien pest in northern Iran. Journal of Insect Biodiversity and Systematics 9 (1): 81-101, DOI: 10.52547/jibs.9.1.81, URL: https://doi.org/10.52547/jibs.9.1.81
03BDF65F0606CC66FF004480966D0F83.taxon	distribution	Distribution in Iran. Guilan (Ghahari, 2017; current study), Mazandaran (Ghahari, 2017), Fars (Gheibi & Ostovan, 2010), Kohgiluyeh and Boyer-Ahmad (Saeidi, 2011) provinces. General distribution. Albania, Algeria, Andorra, Armenia, Austria, Australia (New South Wales, Queensland), Azerbaijan, Belgium, Canada (introduced) (British Columbia, East, Ontario, Prairies), China, Czech Republic, Denmark, Egypt, England, France, Georgia, Germany, Greece, Hungary, Iran, India, Indonesia, Israel, Italy, Japan, Kyrgyzstan, Kazakhstan, Lebanon, Lithuania, Malaysia, Moldova, Montenegro, Morocco, Netherlands, Nigeria, North Korea, Palestine, Papua New Guinea, Poland, Romania, Portugal, Russia (Eastern Siberia, Western Russia, Western Siberia), Serbia, Slovakia, South Africa, South Korea, Spain, Sweden, Switzerland, Taiwan, Tajikistan, Tunisia, Turkey, Ukraine, USA (introduced) (California, Great Plains, Northeast, Pacific Northwest), Uzbekistan, former Yugoslavia, (Cerretti & Ziegler, 2004; Tschorsnig et al., 2004; Vanhara & Tschorsnig, 2006; Stanković et al., 2014; O’Hara et al., 2020; Hammami et al., 2022) (Fig. 4 F).	en	Karami, Amene, Talebi, Ali Asghar, Gilasian, Ebrahim, Fathipour, Yaghoub, Mehrabadi, Mohammad (2023): Native parasitoids of the fall webworm, Hyphantria cunea (Drury, 1773) (Lepidoptera, Erebidae), an invasive alien pest in northern Iran. Journal of Insect Biodiversity and Systematics 9 (1): 81-101, DOI: 10.52547/jibs.9.1.81, URL: https://doi.org/10.52547/jibs.9.1.81
03BDF65F0606CC66FF004480966D0F83.taxon	biology_ecology	Hosts. Lepidoptera: Hyphantria cunea (Drury, 1773), Lymantria dispar (Linnaeus, 1758) Leucoma salicis (Linnaeus, 1758), Euproctis chrysorrhoea (Linnaeus, 1758); Orgyia trigotephras Boisduval, 1829, Ocneria terebinthina Stgr. (Erebidae); Cydalima perspectalis (Walker, 1859) (Crambidae); Hemileuca oliviae Cockerell, 1898 (Saturniidae); Thaumetopoea jordana (Staudinger, 1894, T. solitaria (Freyer, 1838), T. pityocampa (Denis & Schiffermüller, 1775), T. ispartaensis Doğanlar & Avcí (Notodontidae); Euphydryas aurinia (Rottemburg, 1775), Nymphalis antiopa (Linnaeus, 1758) (Nymphalidae); Thiacidas postica Walker, 1855, Mythimna unipuncta (Haworth, 1809); Dicycla oo (Linnaeus, 1758) (Noctuidae) (Halperin, 1990; Sabahi, 1997; Camerini & Groppali, 1999; Dehghani Zahedani et al., 2006; Farar, 2006; Al-e-Mansoor, 2008; Saeidi, 2011; Stanković et al., 2014; Ghahari, 2017; Tschorsnig, 2017; Farahani et al., 2018; Martini et al., 2019; O’Hara et al., 2020; Hammami et al., 2022).	en	Karami, Amene, Talebi, Ali Asghar, Gilasian, Ebrahim, Fathipour, Yaghoub, Mehrabadi, Mohammad (2023): Native parasitoids of the fall webworm, Hyphantria cunea (Drury, 1773) (Lepidoptera, Erebidae), an invasive alien pest in northern Iran. Journal of Insect Biodiversity and Systematics 9 (1): 81-101, DOI: 10.52547/jibs.9.1.81, URL: https://doi.org/10.52547/jibs.9.1.81
03BDF65F0606CC67FF00436A96F2097D.taxon	materials_examined	Material examined. Iran, Guilan province, Rezvanshahr, Paresar, Sandian, (37 ° 57 ′ 22.70 ″ N, 49 ° 12 ′ 82.95 ″ E), 02. ix. 2019, 181 ♂, 196 ♀; on larvae of Hyphantria cunea, leg. A. Karami.	en	Karami, Amene, Talebi, Ali Asghar, Gilasian, Ebrahim, Fathipour, Yaghoub, Mehrabadi, Mohammad (2023): Native parasitoids of the fall webworm, Hyphantria cunea (Drury, 1773) (Lepidoptera, Erebidae), an invasive alien pest in northern Iran. Journal of Insect Biodiversity and Systematics 9 (1): 81-101, DOI: 10.52547/jibs.9.1.81, URL: https://doi.org/10.52547/jibs.9.1.81
03BDF65F0606CC67FF00436A96F2097D.taxon	distribution	Distribution in Iran. Guilan (Ghahari, 2017; current study), Mazandaran (Ghahari, 2017), West Azerbaijan (Karimpour et al., 2005), Zanjan (Modarres Awal, 1994), Kohgiluyeh and Boyer-Ahmad (Saeidi, 2011). General distribution. Armenia, Austria, Azerbaijan, Belarus, Belgium, British Isles, Bulgaria, Canada (introduced), China, Corse, Croatia, Czech Republic, Denmark, Egypt, Finland, France, Georgia, Germany, Greece, Hungary, India, Iran, Israel, Italy, Japan (Hokkaidō, Honshū, Kyūshū), Korean Peninsula (North Korea), Kyrgyzstan, Latvia, Lithuania, Macedonia, Moldova, Mongolia, Netherlands, New England, Palestine, Poland, Portugal, Romania, Russia (Eastern Siberia, Southern Far East, Western Russia, Western Siberia), Saudi Arabia, Scotland, Serbia, Slovakia, Slovenia, Spain, Sweden, Switzerland, Taiwan, Tajikistan, Tunisia, Turkey, Turkmenistan, Ukraine, USA (introduced), Uzbekistan (Kara et al., 2007; El-Hawagry, 2018; O’Hara et al., 2020) (Fig. 4 G).	en	Karami, Amene, Talebi, Ali Asghar, Gilasian, Ebrahim, Fathipour, Yaghoub, Mehrabadi, Mohammad (2023): Native parasitoids of the fall webworm, Hyphantria cunea (Drury, 1773) (Lepidoptera, Erebidae), an invasive alien pest in northern Iran. Journal of Insect Biodiversity and Systematics 9 (1): 81-101, DOI: 10.52547/jibs.9.1.81, URL: https://doi.org/10.52547/jibs.9.1.81
03BDF65F0606CC67FF00436A96F2097D.taxon	biology_ecology	Hosts. Lepidoptera: Hyphantria cunea (Drury, 1773), Lymantria dispar (Linnaeus, 1758), Euproctis chrysorrhoea (Linnaeus, 1758), Leucoma salicis (Linnaeus, 1758), Arctia caja (Linnaeus, 1758) (Erebidae); Pieris brassicae (Linnaeus, 1758), P. rapae (Linnaeus, 1758), Colias croceus (Fourcroy, 1785) (Pieridae); Malacosoma castrensis (Linnaeus, 1758), M. neustria (Linnaeus, 1758), Dendrolimus pini (Linnaeus, 1758) (Lasiocampidae); Tortrix viridana Linnaeus, 1758 (Tortricidae); Cydalima perspectalis (Walker, 1859) (Crambidae); Mamestra oleracea (Linnaeus, 1758), M. brassicae (Linnaeus, 1758), Autographa gamma (Linnaeus, 1758), Spodoptera exigua (Hübner, 1808), S. littoralis (Boisduval, 1833), Agrotis segetum (Denis & Schiffermüller, 1775), Prodenia litura (Fabricius, 1775), Peridroma saucia (Hübner, 1808), Xestia c-nigrum (Linnaeus, 1758), Simyra dentinosa Freyer, 1838, Mythimna loreyi Duponchel, 1827, M. unipuncta (Haworth, 1809), Trichoplusia ni (Hübner, 1803), Lacanobia oleracea (Linnaeus, 1758), Orthosia gracilis (Denis & Schiffermüller, 1775) (Noctuidae) (Herard et al., 1979; Kara & Tschorsnig, 2003; Karimpour et al., 2005; Kara et al., 2007; Cerretti & Tschorsnig, 2010; Razmi et al., 2011; Saeidi, 2011; Depalo et al., 2012; Tschorsnig, 2017; Martini et al., 2019; O’Hara et al., 2020).	en	Karami, Amene, Talebi, Ali Asghar, Gilasian, Ebrahim, Fathipour, Yaghoub, Mehrabadi, Mohammad (2023): Native parasitoids of the fall webworm, Hyphantria cunea (Drury, 1773) (Lepidoptera, Erebidae), an invasive alien pest in northern Iran. Journal of Insect Biodiversity and Systematics 9 (1): 81-101, DOI: 10.52547/jibs.9.1.81, URL: https://doi.org/10.52547/jibs.9.1.81
03BDF65F0607CC6AFFF346F397D00801.taxon	materials_examined	Material examined. Iran, Guilan province, Rezvanshahr, Paresar, Sandian, (37 ° 57 ′ 22.70 ″ N, 49 ° 12 ′ 82.95 ″ E), 02. ix. 2019, 5 ♂, 3 ♀; on pupae of Hyphantria cunea, leg. A. Karami.	en	Karami, Amene, Talebi, Ali Asghar, Gilasian, Ebrahim, Fathipour, Yaghoub, Mehrabadi, Mohammad (2023): Native parasitoids of the fall webworm, Hyphantria cunea (Drury, 1773) (Lepidoptera, Erebidae), an invasive alien pest in northern Iran. Journal of Insect Biodiversity and Systematics 9 (1): 81-101, DOI: 10.52547/jibs.9.1.81, URL: https://doi.org/10.52547/jibs.9.1.81
03BDF65F0607CC6AFFF346F397D00801.taxon	distribution	Distribution in Iran. Alborz (Zamani et al., 2005), Zanjan (Ghavami & Djalilvand, 2015), Fars (Sadeghi et al., 2013), Guilan (Ejlali et al., 2008; current study). General distribution. Albania, Andorra, Angola, Austria, Belgium, Bosnia and Herzegovina, Brazil, Bulgaria, Cameroon, Cape Verde, China, Colombia, Congo, Costa Rica, Croatia, Cuba, Egypt, France, Georgia, Ghana, Greece, India, Iran, Italy, Japan, Kuwait, Macedonia, Malaysia, Malta, Mexico, Montenegro, Netherlands, New Zealand, Norway, Pakistan, Philippines, Portugal, Puerto Rico, San Marino, Saudi Arabia, Serbia, Slovenia, South Korea, Spain, Sri Lanka, Tanzania, Thailand, Togo, Turkey, United Kingdom, United States of America, Venezuela (Brown & Oliver, 2007; Disney, 2008; Wakid, 2008; Disney et al., 2010; Seebens et al., 2017; Cham et al., 2018; Khameneh et al., 2018; Debnath & Roy, 2019) (Fig. 4 H).	en	Karami, Amene, Talebi, Ali Asghar, Gilasian, Ebrahim, Fathipour, Yaghoub, Mehrabadi, Mohammad (2023): Native parasitoids of the fall webworm, Hyphantria cunea (Drury, 1773) (Lepidoptera, Erebidae), an invasive alien pest in northern Iran. Journal of Insect Biodiversity and Systematics 9 (1): 81-101, DOI: 10.52547/jibs.9.1.81, URL: https://doi.org/10.52547/jibs.9.1.81
03BDF65F0607CC6AFFF346F397D00801.taxon	biology_ecology	Hosts. Lepidoptera: Hyphantria cunea (Drury, 1773) (Erebidae), Peridroma saucia (Hübner, 1808) (Noctuidae); Diptera: Aneomochtherus mundus (Loew, 1849) (Asilidae), Palaeosepsis sp. (Sepsidae); Hemiptera: Triatoma brasiliensis Neiva, 1911 (Reduviidae); Hymenoptera: Apis mellifera Linnaeus, 1758 (Apidae); Blattodea: Macrotermes gilvus (Hagen, 1858) (Termitidae); Ixodida: Dermacentor nitens Neumann, 1897 (Ixodidae); Rhipicephalus microplus Canestrini, 1888 (Ixodidae); Otobius megnini Dugès, 1883 (Argasidae); Amblyomma variegatum Fabricius, 1794 (Ixodidae); Orthoptera: Zonocerus variegatus (Linnaeus, 1758) (Pyrgomorphidae); Coleoptera: Curculio caryae Horn, 1873 (Curculionidae); Macrodactylus marinus (Scarabaeidae); Araneae: Brachypelma vagans (Ausserer, 1875) (Theraphosidae); Mantodea: Parastagmatoptera tessellata Saussure & Zehntner, 1894 (Mantidae); Agaricales: Agaricus bisporus (Lange, 1946) (Agaricaceae); Scorpiones: Mesobuthus eupeus (Koch, 1839) (Buthidae) (Gregorio & Leonide, 1980; Harrison & Gardner, 1991; Arredondo-Bernal & Trujillo-Arriaga, 1994; Downie et al., 1995; Zamani et al., 2005; Costa et al., 2007; Ejlali et al., 2008; Disney, 2008; Abdi Goodarzi et al., 2012; Mongiardino Koch et al., 2013; Machkour-M'Rabet et al., 2015; Zhang et al., 2017; Marchiori, 2020; Noknoy et al., 2020). Species frequency and rate of parasitism. Four parasitoid species were found as the major insect parasitoids of the fall webworm. These were Chouioia cunea Yang, 1989 (Hym., Eulophidae), Psychophagus omnivorus (Walker, 1835) (Pteromalidae), Compsilura concinnata (Meigen, 1824) and Exorista larvarum (Linnaeus, 1758) (Diptera, Tachinidae) which appeared to have considerable influence on the population of this pest. Among the hymenopterous parasitoid species, C. cunea and P. omnivorus showed the highest number of H. cunea parasitoids in the collected pupae samples. Of these, C. cunea was the most abundant parasitoid species and may be important in the initial regulation of moth populations. Exoriata larvarum and C. concinnata were also the dominant larval parasitoids of this pest. Compsilura concinnata occupied the position as the second most abundant parasitoid of H. cunea. The gregarious endoparasitoid E. larvarum, despite the lower population than C. concinnata, appeared to have considerable influence on the population of this moth pest. The rest of the species were in relatively low densities, with fewer than ten individuals each and did not play an important role as regulating agents of H. cunea (Fig. 5). There was a significant difference in the contribution of each parasitoid species to overall parasitism (66.3 %) achieved by the parasitoid complex. Chouioia cunea was the most predominant and effective species with the highest parasitism percentage among collected parasitoids (24.33 %) and is followed by P. omnivorus (15.5 %) and C. concinnata (13.73 %). Exorista larvarum was the fourth most prevalent parasitoid recovered in this study (12.57 %) which can be a major factor for the population dynamics of this pest. But the remaining species were rare, with fewer than ten individuals each and overall represent 0.53 % of the parasitoids reared from H. cunea (Fig. 6).	en	Karami, Amene, Talebi, Ali Asghar, Gilasian, Ebrahim, Fathipour, Yaghoub, Mehrabadi, Mohammad (2023): Native parasitoids of the fall webworm, Hyphantria cunea (Drury, 1773) (Lepidoptera, Erebidae), an invasive alien pest in northern Iran. Journal of Insect Biodiversity and Systematics 9 (1): 81-101, DOI: 10.52547/jibs.9.1.81, URL: https://doi.org/10.52547/jibs.9.1.81
