identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03B5846AFF8BFFC6FD06F9C1FC633AF5.text	03B5846AFF8BFFC6FD06F9C1FC633AF5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Belladessus soqtapata Miller 2020	<div><p>Belladessus soqtapata Miller, new species</p><p>Zoobank.org/ urn:lsid:zoobank.org:act: A08412EA-13C8-4245-ACAB-5A334C740BA9</p><p>(Figs. 1–5, 8–13)</p><p>Type locality. Peru, Cusco Prov, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-70.84965&amp;materialsCitation.latitude=-13.345235" title="Search Plazi for locations around (long -70.84965/lat -13.345235)">Soqtapata Conservation Reserve</a>, 13.345235°S 70.849646°W, 988 m elev, pools in small stream (Figs. 8, 9).</p><p>Diagnosis. This species shares the following character states with other Belladessus within Bidessini: 1) Absence of an occipital line (Fig. 1); 2) presence of a basal pronotal stria (Fig. 1); 3) presence of a basal elytral stria (Fig. 1); 4) absence of a sutural elytral stria (Fig. 1); 5) absence of a transverse carina on the elytral epipleuron at the humeral angle; and 6) presence of a distinct marginal bead on the anterior clypeal margin (Fig. 1). From other Belladessus, this species differs in having males. Males have a distinctive median lobe that is exceptionally broad in lateral aspect (Fig. 3) and apically bifid in ventral aspect (Fig. 4). Also, the complex dorsal color pattern with maculae and fasciae is distinctive (Fig. 1). The pronotum is entirely pale orange in other species of Belladessus (Figs. 6, 7), but in B. soqtapata there is a conspicuous posteromedial marginal dark, well-defined border (Fig. 1). Finally, the anterior clypeal margin is distinctly and strongly beaded as in B. puella, but unlike B. femineus, which has this bead present but less distinctly impressed.</p><p>Description. Measurements: TL = 1.9–2.1 mm, GW = 1.1–1.2 mm, PW = 0.8–1.0 mm, HW = 0.6–0.7 mm, EW = 0.4–0.5 mm, TL/GW = 1.7–1.8, HW/EW = 1.6–1.7, FL/RL = 1.7–1.8. Body shape oval, lateral outline slightly discontinuous between pronotum and elytron, posterior apex pointed (Fig. 1). Coloration: Head yellow-orange, slightly darker anteriorly. Pronotum yellow, narrowly dark along posterior margin laterally. Elytron maculate, dark brown and yellow, with large brown regions anteromedially, lateromedially, medially, and in longitudinal vittae subapically on each elytron as well as in other small areas (Fig. 1). Sculpture and structure: Head shiny and smooth with small micropunctures; anterior clypeal margin produced, distinctly and broadly beaded; eyes moderately large (HW/EW = 1.6–1.7). Pronotum smooth and shiny, with few micropunctures; pronotal plica strongly impressed, curved, extending about onehalf distance across pronotum; lateral pronotal margin distinctly, narrowly beaded. Elytron with lateral margin broadly rounded; surface shiny with dispersed micropunctures; elytral stria elongate, distinctly longer than length of pronotal stria. Prosternum medially short, carinate, prosternal process broad, deeply longitudinally grooved. Metaventrite and metaventrital wings smooth and shiny with few micropunctures. Metacoxa smooth and shiny, with few micropunctures; metacoxal lines distinct, region between metacoxal lines moderately broad; metatrochanter large relative to metafemur (RL/FL = 1.7–1.8; Fig. 2). Male genitalia: Median lobe in lateral aspect extremely broad, broadly and evenly curved along ventral margin, dorsal margin slightly curved, apex pointed (Fig. 3); in ventral aspect slender, apically bifid, sides subparallel, slightly convergent (Fig. 4); lateral lobe in lateral aspect robust, apical segment with apex rounded, with distinct apical tooth-shaped lobe (Fig. 5).</p><p>Sexual Dimorphism. Males have the pro- and mesotarsomeres I–III somewhat but distinctly broader than in females.</p><p>Variation. There is some minor variation in extent of coloration of the dorsal surface but otherwise specimens are similar.</p><p>Etymology. This species is named soqtapata, the Quechua word for “six floors,” and the name of the Soqtapata Conservation Reserve, the type locality of the species.</p><p>Distribution. Belladessus soqtapata is only known from the type locality (Figs. 8, 9). However, this habitat type extends north and south along the base of the Andes Mountains for many miles, suggesting the species may be considerably more widespread.</p><p>Habitat. Soqtapata is a conservation reserve at the base of a steep area of the east slope of the Andes Mountains. The type locality is at 988 m elevation in dense tropical forest (Figs. 10–13). The specimens were found in small rock pools in a trickling spring in the heavily shaded forest adjacent to the larger stream in a former channel of the stream (Figs. 10–13). Specimens were not found in the larger stream or side pools, only in the rock pool habitat of the small trickling spring. The pools had extensive leaf-pack and areas with denser algae. Specimens were uncommon despite extensive search. The only other diving beetles found in this habitat were members of a rarely collected species of Laccophilus Leach (not identified to species).</p><p>Discussion. This species includes specimens with the same diagnostic features of Belladessus, but clearly there are male specimens involved, which are not known from the other two species of Belladessus . The relationship of this species to the other two species in the genus is not perfectly clear, but specimens are similar in general appearance and diagnostic combination within Bidessini. All share a somewhat similar habitus and dorsally complex coloration (Figs. 1, 6, 7) (Miller and Short 2015).</p><p>Type Material. Holotype in MUSM, male, labeled “ PERU Cusco Dist, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-70.84965&amp;materialsCitation.latitude=-13.345235" title="Search Plazi for locations around (long -70.84965/lat -13.345235)">Soqtapata Cons Res</a>, 988m 13.345235°S 70.849646°W pools in small stream, 5 Jun 2019 / HOLOTYPE Belladessus soqtapata Miller, 2020 [red label with double black line border].” 2 paratypes labeled same as holotype, except “…/ PARATYPE Belladessus soqtapata Miller, 2020 [blue label with black line border].” 1 paratype, labeled “ PERU: Cusco Dept. Conc. Soqtapata, 3Aug2018 13.34714°S 70.84400°W Pools in stream KBM03081801 KBMiller/ PARATYPE Belladessus soqtapata Miller, 2020 [blue label with black line border].”</p></div>	https://treatment.plazi.org/id/03B5846AFF8BFFC6FD06F9C1FC633AF5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Miller, Kelly B.	Miller, Kelly B. (2020): Belladessus soqtapata, New Species from Peru (Coleoptera: Adephaga: Dytiscidae: Hydroporinae: Bidessini). The Coleopterists Bulletin 74 (2): 441-446, DOI: 10.1649/0010-065X-74.2.441, URL: http://dx.doi.org/10.1649/0010-065x-74.2.441
03B5846AFF8FFFC6FCE5FE55FB2F3F61.text	03B5846AFF8FFFC6FCE5FE55FB2F3F61.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Belladessus Miller and Short	<div><p>KEY TO SPECIES OF BELLADESSUS</p><p>1. Anterior clypeal margin thickened and distinctly but weakly bordered; eastern Venezuela and Suriname (Fig. 8) ............ B. femineus</p><p>1ʹ. Anterior clypeal margin strongly modified, flattened and prominently beaded ............... 2</p><p>2. Males present; male median lobe very broad and evenly curved in lateral aspect (Fig. 2), slender and apically bifid in ventral aspect (Fig. 3); basal elytral stria distinctly longer than basal pronotal stria (Fig. 1); dorsal color pattern complex, including extensive maculations and fasciae, with dark brown along posterior margin of pronotum (Fig. 1); Peru (Fig. 8) ........................ B. soqtapata, new species</p><p>2ʹ. Males apparently absent; basal elytral stria similar in length to basal pronotal stria (Fig. 7); dorsal color pattern simpler, restricted to a few larger maculae, entire pronotum yellow-orange (Fig. 7); western Venezuela (Fig. 8) ............. ................................................. B. puella</p></div>	https://treatment.plazi.org/id/03B5846AFF8FFFC6FCE5FE55FB2F3F61	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Miller, Kelly B.	Miller, Kelly B. (2020): Belladessus soqtapata, New Species from Peru (Coleoptera: Adephaga: Dytiscidae: Hydroporinae: Bidessini). The Coleopterists Bulletin 74 (2): 441-446, DOI: 10.1649/0010-065X-74.2.441, URL: http://dx.doi.org/10.1649/0010-065x-74.2.441
03B5846AFF8FFFC6FD15FBA1FB0D3F8D.text	03B5846AFF8FFFC6FD15FBA1FB0D3F8D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Belladessus Miller and Short	<div><p>SPECIES IN THE GENUS BELLADESSUS</p><p>Belladessus femineus Miller and Short, 2015 (Suriname, eastern Venezuela, Fig. 8)</p><p>B. puella Miller and Short, 2015 (western Venezuela, Fig. 8)</p><p>B. soqtapata Miller, new species (Peru, Fig. 8)</p></div>	https://treatment.plazi.org/id/03B5846AFF8FFFC6FD15FBA1FB0D3F8D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Miller, Kelly B.	Miller, Kelly B. (2020): Belladessus soqtapata, New Species from Peru (Coleoptera: Adephaga: Dytiscidae: Hydroporinae: Bidessini). The Coleopterists Bulletin 74 (2): 441-446, DOI: 10.1649/0010-065X-74.2.441, URL: http://dx.doi.org/10.1649/0010-065x-74.2.441
