identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03B6CE240D54FFA0FD85F8C4BF5FF907.text	03B6CE240D54FFA0FD85F8C4BF5FF907.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cervidellus hueckeswagensis Mach & Pettrich & Villegas & Waldvogel & Schiffer & Holovachov 2025	<div><p>Cervidellus hueckeswagensis sp. nov.</p><p>urn:lsid:zoobank.org:act: 78EECCE1-ADD0-4797-8C41-2A3A320C82AF</p><p>Figs 1–3, 7A–F, Tables 1–2</p><p>Diagnosis</p><p>Cervidellus hueckeswagensis sp. nov. is characterized by a body length of 488–539 µm in females; cuticle with longitudinal incisures; lateral alae with five incisures extending to tail tip; three pairs of asymmetrical lips, pairs of lips separated by V-shaped primary axils with two triangular-acute guarding processes, and each lip asymmetrically triangular with 6–7 tines along its margin; three labial probolae, each bifurcated at one level, at about half of their length; labial probolae lacking additional tines or swellings; pharyngeal corpus 2.1–3.4× as long as isthmus; excretory pore at corpus/isthmus level; vulva continuous with body contour; spermatheca 7–15 µm long; and post-vulval uterine sac 12–32 µm long.</p><p>Etymology</p><p>The specific epithet is derived from the hometown (‘Hückeswagen’) of the first author.</p><p>Type material</p><p>Holotype</p><p>CHILE • ♀; Tarapacá Region, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-68.77528&amp;materialsCitation.latitude=-20.581383" title="Search Plazi for locations around (long -68.77528/lat -20.581383)">Tamarugal Province</a>; 20°34.883′ S, 68°46.517′ W; 4056 m a.s.l.; 4 Oct. 2022; O. Holovachov, L. Villegas, L. Pettrich, A.-M. Waldvogel and P. Schiffer leg.; soil, sample code ALT.22.10 (Fig. 7A–B); SMNH Type-9945; SMNH.</p><p>Paratypes</p><p>CHILE • 6 ♀♀; same data as for holotype; SMNH Type-9945; SMNH .</p><p>Other material examined</p><p>CHILE • 7 ♀♀; Tarapacá Region, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-68.97528&amp;materialsCitation.latitude=-20.339716" title="Search Plazi for locations around (long -68.97528/lat -20.339716)">Tamarugal Province</a>; 20°20.383′ S, 68°58.517′ W; 4196 m a.s.l.; 3 Oct. 2022; O. Holovachov, L. Villegas, L. Pettrich, A.-M. Waldvogel and P. Schiffer leg.; soil under Azorella compacta Phil., sample code ALT.22.08 (Fig. 7E–F); SMNH .</p><p>Associated molecular data</p><p>CHILE • 1 ♀; Tarapacá Region, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-69.042786&amp;materialsCitation.latitude=-20.335833" title="Search Plazi for locations around (long -69.042786/lat -20.335833)">Tamarugal Province</a>; 20°20.150′ S, 69°02.567′ W; 3886 m a.s.l.; 3 Oct. 2022; O. Holovachov, L. Villegas, L. Pettrich, A.-M. Waldvogel and P. Schiffer leg.; soil under unidentified bush, sample code ALT.22.04; GenBank no: PQ587598 (18S) • 1 ♀; Antofagasta Region, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-67.848885&amp;materialsCitation.latitude=-23.516117" title="Search Plazi for locations around (long -67.848885/lat -23.516117)">El Loa Province</a>; 23°30.967′ S, 67°50.933′ W; 3830 m a.s.l.; 9 Oct. 2022; Holovachov, A.-M. Waldvogel, L. Schiffer and P. Schiffer leg.; soil under dead cactus, sample code EPT.22.3830; GenBank no: PQ587809 .</p><p>Description</p><p>Adult</p><p>Body variably curved ventrad upon fixation (fixed without applying heat). Cuticle coarsely annulated, annuli 2–2.5 µm wide at midbody. Longitudinal incisures present, giving cuticle tiled appearance. Lateral alae start at level of procorpus, and expand to five incisures at metacorpus / ithmus, extending to tail tip. Lip region slightly offset, with six labial and four cephalic papillae. Amphid opening relatively small, oval, located at base of each lateral lip. Three pairs of asymmetrical lips, one dorsal and two ventrolateral. Pairs of lips separated by V-shaped primary axils with two triangular-acute guarding processes. Secondary axils V-shaped, without guarding processes. Each lip asymmetrically triangular with 6–7 elongate-triangular tines along its margin: 2–3 tines facing primary axil, one long tine pointing forward, and three long tines facing secondary axil (Fig. 3). Three labial probolae bifurcated at about half of their length; prongs slender, curved towards each other, without secondary tines or bifurcations. Basal knobs or ridges absent. Stoma about as long as lip region diameter. Cheilorhabdia irregular in latero-median view; metastegostom with large dorsal denticle. Pharynx cephaloboid. Pharyngeal corpus fusiform; isthmus well demarcated from corpus; basal bulb oval with well-developed grinder. Nerve ring surrounding metacorpus but poorly visible in most specimens. Excretory pore at level of corpus-isthmus junction. Deirids at level with isthmus or basal bulb.</p><p>Female</p><p>Reproductive system monodelphic, prodelphic, in dextral position relative to intestine. Ovary reflexed posteriorly at oviduct, straight posterior to vulva. Vulva continuouing with body contour. Spermatheca weakly developed, often hard to recognize. Post-vulval uterine sac usually small, equal to one vulval body diameter. Tail conoid with 14–16 ventral annuli, and terminus minutely rounded. Phasmids located at about 30–56% of tail length.</p><p>Male</p><p>Not found. Species is possibly asexual.</p><p>Differential diagnosis</p><p>Cervidellus hueckeswagensis sp. nov. from Chile is most similar to C. caucetensis Holovachov, Boström &amp; De Ley, 2011 and C. vinciguerrae Clausi, 1998 in the shape of the labial probolae (bifurcated at about middle of their length and with slender, curved inwardly prongs slender, without secondary tines or bifurcations), and to C. bifidihastatus Abolafia &amp; Robles, 2024, C. hamatus Thorne, 1937, C. doorsselaeri (De Clerk &amp; De Ley, 1990) Boström &amp; De Ley 1996 and C. capraeolus (De Ley, Geraert &amp; Coomans, 1990) Boström &amp; De Ley 1996 in the shape of the cephalic probolae (asymmetrically triangular with multiple tines), see Table 2 for more details.</p><p>The new species differs from C. caucetensis in the shape of cephalic probolae and larger number of tines (6–7 vs 4) the mode of reproduction (asexual vs sexual) and the length of the spermatheca (7–15 µm vs 24–35 µm); C. vinciguerrae in the shape of the cephalic probolae and the larger number of tines (6–7 vs 5), the mode of reproduction (asexual vs sexual) and the length of the spermatheca (7–15 µm vs 27–40 µm); C. bifidihastatus in the shape of the labial probolae (without vs with backwardly directed spur-like processes), a different number and shape of tines on the cephalic probolae (two-three tines facing the primary axil, one long tine pointing forward, and three long tines facing the secondary axil vs four small rounded tines facing the primary axil, one rounded tine facing forward, and three larger rounded tines facing the secondary axil), the mode of reproduction (asexual vs sexual) and the length of the spermatheca (7–15 µm vs 16–19 µm); C. hamatus in the shape of the labial probolae (without vs with bifurcation), a different number and shape of tines on the cephalic probolae (2–3 tines facing the primary axil, one long tine pointing forward, and three long tines facing the secondary axil vs three small tines facing the primary axil, no tine pointing forward and three large tines facing the secondary axil) and the length of the spermatheca (7–15 µm vs 16–37 µm); C. doorsselaeri in the shape of the labial probolae (without vs with bifurcation) and a different number and shape of tines on the cephalic probolae (two-three tines facing the primary axil, one long tine pointing forward, and three long tines facing the secondary axil vs two round tines facing the primary axil, three tines pointing forward and one rounded and two acute tines facing the secondary axil); C. capraeolus in the shape of labial probolae (without vs with bifurcation and additional processes), different number and shape of tines on the cephalic probolae (2–3 tines facing the primary axil, one long tine pointing forward, and three long tines facing the secondary axil vs two round tines facing the primary axil, two tines pointing forward and four tines facing the secondary axil) and the length of the post-vulval uterine sac (12–32 µm vs 4–6 µm).</p></div>	https://treatment.plazi.org/id/03B6CE240D54FFA0FD85F8C4BF5FF907	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Mach, Niclas;Pettrich, Laura;Villegas, Laura;Waldvogel, Ann-Marie;Schiffer, Philipp H.;Holovachov, Oleksandr	Mach, Niclas, Pettrich, Laura, Villegas, Laura, Waldvogel, Ann-Marie, Schiffer, Philipp H., Holovachov, Oleksandr (2025): Description of Cervidellus hueckeswagensis sp. nov. and Stegelleta waltrautae sp. nov. (Nematoda: Rhabditida: Cephalobidae) from the Atacama Desert in Chile. European Journal of Taxonomy 1012: 222-238, DOI: 10.5852/ejt.2025.1012.3051, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3051/13651
03B6CE240D5DFFACFDD6FEA8BF3DFD5B.text	03B6CE240D5DFFACFDD6FEA8BF3DFD5B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Stegelleta waltrautae Mach & Pettrich & Villegas & Waldvogel & Schiffer & Holovachov 2025	<div><p>Stegelleta waltrautae sp. nov.</p><p>urn:lsid:zoobank.org:act: 05D4E5B5-8719-446C-BC1F-F341D15E595D</p><p>Figs 4–6, 7G–H, Tables 3–4</p><p>Diagnosis</p><p>Stegelleta waltrautae sp. nov. is characterized by a body length of 531–631µm in females and 576– 583 µm in males; cuticle divided by longitudinal incisures into about 30 irregular ridges; lateral alae with five incisures extending to tail terminus; three pairs of asymmetrical lips, pairs of lips separated by U-shaped primary axils without guarding processes, and asymmetrically trapezoidal lips with two elongate-triangular tines along margin: one longer tine facing the primary axil and one shorter tine facing secondary axil; three labial probolae each bifurcated at about half of their length; prongs slender, apically curved towards each other, without secondary tines or bifurcations; pharyngeal corpus 3.1–4.7 × as long as isthmus; nerve ring and excretory pore at isthmus / basal bulb level; vulva convex with less thickened anterior vulval lip and more thickened posterior vulval lip; spermatheca 28.0–36.0 µm long; and post-vulval uterine sac 25.0–39.0 µm long.</p><p>Etymology</p><p>The new species is named in honor of the first author’s grandmother, who recently passed away.</p><p>Type material</p><p>Holotype</p><p>CHILE • ♀; Atacama Region, Copiapo Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-71.04778&amp;materialsCitation.latitude=-27.793884" title="Search Plazi for locations around (long -71.04778/lat -27.793884)">Totoral</a>; 27°47.633′ S, 71°02.867′ W; 189 m a.s.l.; 3 Oct. 2023; O. Holovachov and P. Schiffer leg.; dune area, soil under a cactus, sample code TDT.23.14 (Fig. 7G–H); SMNH Type-9946; SMNH.</p><p>Paratypes</p><p>CHILE • 4 ♀♀, 2 ♂♂; same collection data as for holotype; SMNH Type-9946; SMNH .</p><p>Description</p><p>Adult</p><p>Body straight or slightly curved ventrad upon fixation (fixed without applying heat). Cuticle coarsely annulated, annuli 2.5–3.0 µm wide at midbody. Distinct longitudinal incisures present, giving cuticle tiled appearance, cuticular blocks with rounded corners, arranged in about 30 irregular longitudinal ridges (in addition to lateral alae). Lateral alae start with three incisures at level of metacorpus and expands to five incisures at level with deirid, extending to tail terminus in both sexes. Lip region continuous, with six labial and four cephalic papillae. Amphid opening relatively small, oval, located at the base of each lateral lip. Three pairs of asymmetrical lips, one dorsal and two ventrolateral. Pairs of lips separated by U-shaped primary axils without guarding processes. Secondary axils U- or V-shaped without guarding processes. Each lip asymmetrically trapezoidal with two elongate-triangular tines along margin: one longer tine facing primary axil and one shorter tine facing secondary axil. Three labial probolae, bifurcated at about half of their length; prongs slender, apically curved towards each other, without secondary tines or bifurcations. Longitudinal swellings or ridges absent. Stoma slightly longer than lip region diameter. Cheilorhabdia bacilliform in latero-median view; metastegostom with small dorsal denticle. Pharynx cephaloboid. Pharyngeal corpus cylindrical; isthmus less well demarcated from corpus; basal bulb oval with well-developed grinder. Nerve ring surrounding isthmus. Excretory pore at level of isthmus-bulb junction. Deirids at level with basal bulb.</p><p>Female</p><p>Reproductive system monodelphic, prodelphic, in dextral position relative to intestine. Ovary reflexed posteriorly at oviduct, straight or with two flexures posterior to vulva. Vulval region convex, well pronounced with less thickened anterior vulval lip and more thickened posterior vulval lip. Spermatheca well developed. Post-vulval uterine sac usually small, about equal to one vulval body diameter. Tail conoid with 14–15 ventral annuli, and rounded tip. Phasmids located at about 32–42% of tail length.</p><p>Male</p><p>Reproductive system monorchic, dextral in position; testis usually reflexed ventrad anteriorly. Spicules paired and symmetrical, curved ventrad; with oval manubrium and subcylindrical, gradually narrowing shaft. Gubernaculum curved plate. Genital papillae distributed as follows: two pairs of subventral precloacal papillae (33–35 µm and 79–80 µm from cloacal aperture, respectively) and one subventral pair at level of cloacal aperture; single midventral papilla on anterior cloacal lip; one caudal pair subventrally few annuli posterior to cloacal aperture, second caudal pair laterally at mid-tail very close to the phasmids and three more pairs (one subdorsal, one subventral and one lateral) near tail terminus. Phasmids located at about 33–40% of tail length. Tail conoid with short bluntly rounded mucro.</p><p>Differential diagnosis</p><p>Stegelleta waltrautae sp. nov. from Chile is most similar to S. tuarua Yeates, 1967 in having a large number of longitudinal lines (rows of cuticular blocks) and in having lateral alae (filed) with five incisures, thus differing from the remaining six species of the genus (Table 4; Boström &amp; Holovachov 2014). The new species, however, differs from S. tuarua in the shape of the cephalic probolae (with two acute tines vs single tine in S. tuarua) and distinctly protruding vulva (vs continuous with body contour).</p></div>	https://treatment.plazi.org/id/03B6CE240D5DFFACFDD6FEA8BF3DFD5B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Mach, Niclas;Pettrich, Laura;Villegas, Laura;Waldvogel, Ann-Marie;Schiffer, Philipp H.;Holovachov, Oleksandr	Mach, Niclas, Pettrich, Laura, Villegas, Laura, Waldvogel, Ann-Marie, Schiffer, Philipp H., Holovachov, Oleksandr (2025): Description of Cervidellus hueckeswagensis sp. nov. and Stegelleta waltrautae sp. nov. (Nematoda: Rhabditida: Cephalobidae) from the Atacama Desert in Chile. European Journal of Taxonomy 1012: 222-238, DOI: 10.5852/ejt.2025.1012.3051, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3051/13651
