taxonID	type	description	language	source
03A16432FFE8A8215DAACC0E8954F80C.taxon	description	(Figs. 2 – 4)	en	Amaral, I. F., Cifuentes-Vargas, E. T., Salles, F. F. (2025): Cloeodes tovauna: a new species previously identified as Cloeodes aymore Massariol & Salles, 2011 (Ephemeroptera: Baetidae). Zootaxa 5594 (2): 355-366, DOI: 10.11646/zootaxa.5594.2.7, URL: https://doi.org/10.11646/zootaxa.5594.2.7
03A16432FFE8A8215DAACC0E8954F80C.taxon	diagnosis	Diagnosis. Nymph. 1) Body size 5.5 – 6.0 mm; 2) Body color pattern dark brown on head, thorax and abdominal segments I, V, VI and X; 3) Labrum with dorsal arc of setae composed of 1 + 0 + 3 setae (Fig. 3 A); 4) Inner margin of labial palp segment III with robust pectinate setae (Fig. 3 I); 5) Hind wing pads present; 6) Fore femur with apex projected, with 2 blunt setae (Fig. 4 A); 7) Posterior margins of terga II-X with spines of the same length; 8) Paraprocts with 15 – 19 spines (Figs. 4 E, F); 9) Cercus with spines on every two segments. Adults. 1) Turbinate eyes of males separated by approximately the width of a lateral ocelli; upper surface oblong, yellowish-brown (Fig. 5 A); 2) Thorax dark brown (Fig. 5 A); 3) Terga I, VII – X brown; tergum IV with median pair of brown spots; 4) Hind wing present (Fig. 5 C); 5) Forceps brown, lighter towards apex, forceps segment III oval; 6) Posterior margin of subgenital plate slightly convex at middle.	en	Amaral, I. F., Cifuentes-Vargas, E. T., Salles, F. F. (2025): Cloeodes tovauna: a new species previously identified as Cloeodes aymore Massariol & Salles, 2011 (Ephemeroptera: Baetidae). Zootaxa 5594 (2): 355-366, DOI: 10.11646/zootaxa.5594.2.7, URL: https://doi.org/10.11646/zootaxa.5594.2.7
03A16432FFE8A8215DAACC0E8954F80C.taxon	description	Description. Nymph. Length (in mm). Body: 5.5 – 6.0; 1.9; cerci: 2.7; terminal filament: 2.9; antenna: 1.9. Coloration (Fig. 2). General coloration dark brown with yellowish-white (Fig. 2 A, B). Head dark brown with yellowish-white stripe between inner margin of compound eye and lateral ocellus (in males, reaching the base of turbinate eyes). Labrum and mandible darkish brown; hypopharynx, maxilla and labium light brown (Figs. 2 C, D). Thorax dark brown to light brown. Fore wing pads light brown at base and yellowish-white distally. Legs yellowish-white, mid and hind femora with apical dark brown band reaching ½ of its length. Abdominal segments I, V, VI and X dark brown; segments II, III, IV, VII, VIII and IX yellowish-white; terga II and III with sublateral marks light brown; tergum IV with sublateral marks orangish brown; tergum V with anteromedial subtriangular yellowish-white mark; tergum VI with yellowish-white sublateral area. Sterna yellowish-white, except for V, VI and X dark brown. Filaments light brown, with a yellowish-white band subdistally and a brown band distally. Head. Labrum (Fig. 3 A). Length about 0.7 × maximum width; dorsal arc of setae composed of 1 + 0 + 3 long setae; ventrally with 6 – 10 spine-like setae near lateral margin. Left mandible (Figs. 3 B, C). Margin between prostheca and mola slightly convex. Right mandible (Figs. 3 D, E). Prostheca slender and pectinate, bifurcated at the middle; inner lobe long, outer lobe divided in the middle. Margin between prostheca and mola slightly convex. Hypopharynx as in Fig. 3 F. Maxilla (Fig. 3 G, H). Canines and distal dentiseta stout. Medially with 1 spine-like seta + 6 apically pointed setae. Maxillary palp as long as galea-lacinia; palp segment II 1.5 × length of segment I. Labium (Fig. 3 I, J). Base of glossa expanded (reaching at least half of paraglossa); inner and outer margins, respectively, with around 17 and 12 pointed setae increasing in length apically; ventral surface with one row of 7 short setae near the inner margin. Paraglossa apex with 2 rows of spine-like setae; outer margin with 3 pectinate setae; dorsally with arc of 3 setae close to inner margin; ventrally with arc of 7 – 8 setae close to inner margin. Labial palp with segment I 0.8 × length of segments II and III combined; segment II dorsally with oblique row of 6 – 7 setae; segment III rounded with 2 – 5 robust pectinate setae in inner margin; ventrally covered with several spine-like simple setae along margins and ventral surface. Thorax. Hind wing pad present. Fore leg (Fig. 4 A). Ratio 1.1: 1 (0.6 mm): 1: 0.2. Femur. Length about 4.4 × maximum width; outer margin with row of 8 – 9 clavate setae (length of setae about 0.1 × maximum width of femur); anterior and posterior margins parallel; submarginal setae absent; anterior surface and inner margin with around 36 simple spine-like setae; posterior surface with around 27 scattered setae near inner margin; apex projected; distal projection 0.1 × width of femur; with 2 blunt, distally expanded setae. Tibia. Outer margin with few simple setae; inner margin with two serrate setae on apex; subtending bristle blunt, distally expanded, extending apex of tibia. Tarsus. Ventrally with row of 13 – 15 + 1 spine-like short setae. Tarsal claw. 0.2 × length of tarsus. Mid and hind legs (Figs. 4 B, C). Similar to fore leg, except for subapical projections of femur less developed and type of setae on inner apex of tibia on (simple instead of serrate). Abdomen. Terga (Fig. 4 D). Spines present on posterior margins of segments II – X (occasionally reduced in segment II); spines approximately with the same length and not slanted, larger spines 2.5 × longer than wide. Sterna. Spines present on posterior margin of segments IV – IX. Gills. Outer margin serrated on distal 1 / 4; inner margin slightly serrated; few secondary tracheae extending from main trunk to inner and outer margins. Gill I about 3 / 4 length of segment II, oval. Gill IV 1 1 / 2 length of segment V, asymmetric, broad at the middle. Gill VII about 1 1 / 2 length of segment VIII. Paraproct (Figs. 4 E – F) with 15 – 19 spines. Cercus with large spines on every two segments. Terminal filament without large spines. Male imago (Fig. 5). Length (in mm). Body: 5.8 – 6.0; antenna: 0.9; fore wing: 6.5; hind wing: 1.13 – 1.15; tibia I: 2.1; tibia II: 0.9; tibia III: 0.8; caudal filaments: broken. Coloration (Fig. 5). Head dark brown with yellowish-white stripe between inner margin of compound eyes to base of antenna; yellowish-brown mark between inner margin of lateral ocelli and surrounding middle ocelli. Antenna light brown. Lateral ocelli cream, surrounded with black. Compound eye cream. Turbinate eyes yellowish-brown; basal ½ of stalk brown, remainder cream. Pronotum dark brown. Mesothorax mostly dark brown; antelateroparapsodal and recurrent scuto-scutellar sutures cream. Metanotum dark brown. Fore wing hyaline; veins light brown, dark brown in distal ½ of C – MP. Hind wing hyaline; veins light brown. Foreleg trochanter yellowish-brown with subapical orange mark; trochanter, femur, tibia and tarsi light brown; mid and hindleg similar, light brown. Terga I, VII – X brown; II – V cream; VI light brown; terga II – VI with lateral brown area; tergum IV with median pair of brown spots; posterior margin of terga II – VI with membrane tinged with brown. Sterna I, VII – X brown; II – VI cream. Tracheation black. Forceps predominantly brown, lighter towards apex. Head (Fig. 5 A). Turbinate eyes separated approximately by the width of lateral ocellus; inner margin parallel; upper surface oblong; length 1.35 × width; stalk height 0.66 × width of dorsal portion. Thorax (Fig. 5 B). Anteronotal protuberance rounded. Metascutellar protuberance posteriorly pointed. Fore wing (Fig. 5 C). Stigmatic area with 7 short veins, some incomplete; marginal intercalary veins paired, single between RA-RP 1 (sometimes paired), ICu 1 - ICu 2 and ICu 2 - CuP, absent between veins Sc-RA and CuP-A. Hind wing (Fig. 5 C). With 2 complete longitudinal veins. Costal process hooked, located on basal third. Legs. Foreleg: tibia 0.6 × length of femur; tarsi 0.05 × length of femur; tarsus segments in decreasing order: 2, 3, 4, 5, 1; mid and hindleg similar. Abdomen (Fig. 5 B). Genitalia. (Fig. 5 D). Forceps segment I subrectangular; 0.4 × length of segment II; distance between base of forceps 1.0 × distance between lateral margins of forceps. Segment II of forceps with medial constriction. Segment III oval. Posterior margin of subgenital plate slightly convex at middle. Female imago (Fig. 6 A) Length (in mm). Body: 6.2; antenna: broken; fore wing: 6.4; hind wing: 1; tibia I: 0.9; tibia II: 0.8; tibia III: 0.8; caudal filaments: broken. Head (Fig. 6 A). Coloration light brown; compound eyes, lateral and middle ocelli similar to male imago. Antenna light brown. Thorax (Fig. 6 B). Pronotum light brown with brown marks. Mesothorax predominantly brownish-grey; antelateroparapsodal and medioparapsidal sutures cream; mediolongitudinal suture brown. Metanotum brown. Forewing (Fig. 6 C). Hyaline; longitudinal and cross yellowish-brown. Stigmatic area with 5 to 7 cross veins touching or almost touching subcostal vein and 2 veins not touching subcostal vein; marginal intercalary veins similar to the male imago. Hindwing (Fig. 6 C). Similar to male imago. Legs. Foreleg 2. 9 mm; trochanter yellowish-brown with subapical orange mark; trochanter, femur, tibia and tarsi light brown; middle and hindleg similar. Abdomen (Fig. 6 B). Tergum I dark brown; terga II, V – X brown; terga III and IV light brown; tergum IV with median pair of brown spots. Sterna light brown with sublateral dark brown subtriangular marks on segments II−VIII. Female subimago (Figs. 7 A – B). Body color pattern and lengths similar to female imago but opaque.	en	Amaral, I. F., Cifuentes-Vargas, E. T., Salles, F. F. (2025): Cloeodes tovauna: a new species previously identified as Cloeodes aymore Massariol & Salles, 2011 (Ephemeroptera: Baetidae). Zootaxa 5594 (2): 355-366, DOI: 10.11646/zootaxa.5594.2.7, URL: https://doi.org/10.11646/zootaxa.5594.2.7
03A16432FFE8A8215DAACC0E8954F80C.taxon	materials_examined	Type Material. Holotype: Male imago (in alcohol) with corresponding nymphal exuviae on slide, BRAZIL, Minas Gerais, Araponga, Parque Estadual da Serra do Brigadeiro; Trilha para Pedra do Pato (20 º 44 ’ 38 ” S; 42 º 28 ’ 03 ” W), 23 / VI / 2024, Amaral, I. F., Cifuentes-Vargas, E. T., Salles, F. F. & Lima M. M. cols. (UFVB: EP 02956). Paratypes: One female imago with corresponding nymphal exuviae and one female subimago with corresponding nymphal exuviae on slide, same data as holotype (MNRJ). 3 nymphs, Brazil, Minas Gerais, Araponga, Parque Estadual da Serra do Brigadeiro; início Trilha Pedra do Pato (20 ° 43 ’ 59 ” S 42 ° 28 ’ 12 ” W), 07 / VI / 24, Salles, F. F., Neto, P. B., Sarmento, F. R. P., Cifuentes-Vargas, E. T. & Alvim, J. B. cols. (UFVB: EP 02958). 6 nymphs (three dissected), Brazil, Minas Gerais, Araponga, Parque Estadual da Serra do Brigadeiro; Trilha das águas (20 ° 43 ’ 52 ” S; 42 ° 27 ’ 54 ” W), 28 / V / 2024, Amaral, I. F., Viana, A. D. L. & Correia, M. C. D. cols. (UFVB: EP 02959). One female imago with corresponding nymphal exuviae on slide, Brazil, Minas Gerais, Araponga, Parque Estadual da Serra do Brigadeiro; Trilha das águas (20 ° 43 ’ 52 ” S; 42 ° 27 ’ 54 ” W), 28 / V / 2024, Amaral, I. F., Viana, A. D. L. & Correia, M. C. D. cols. (UFVB: EP 02957). Additional material. 7 nymphs (two dissected) and 1 male imago, Brazil, Minas Gerais, Catas Altas, Cachoeira da Santa (20 º 05 ’ 17 ” S; 43 º 25 ’ 06 ” W), 25 / I / 2015, Salles, F. F. col. UFVB: EP 02962. 2 nymphs, Brazil, Minas Gerais, Ouro Preto, Parque das Andorinhas, Cachoeira Véu das Noivas (20 º 21 ’ 30 ” S; 43 º 29 ’ 18 ” W), 18 / X / 2023, Amaral, I. F., Morais, C. M. A. & Domingues, S. P. cols. UFVB: EP 02963. One nymph (dissected), Brazil, Minas Gerais, Ouro Preto, Parque das Andorinhas, Cachoeira Véu das Noivas (20 º 21 ’ 30 ” S; 43 º 29 ’ 18 ” W), 15 / II / 2024, Amaral, I. F., Silva, E. F. cols. UFVB: EP 02964. Material examined of C. aymore: Holotype. Male imago with corresponding nymphal exuviae, BRAZIL, Espírito Santo, Parque Nacional do Caparaó, Ibitirama, Pedra Roxa River (20 ° 23 ’ 48.1 ’ ‘ S; 41 ° 44 ’ 08.1 ’’ W), 20 / IV / 2008, Salles F. F., Massariol, F. C., Boldrini, R., Lima, M. M. & Britto, P. V. A. cols. (INPA). Paratype. Nymphal exuviae on slide, BRAZIL, Espírito Santo, Parque Nacional do Caparaó, Ibitirama, Tecnotruta River, propriedade “ Sonho meu ”, 21 / iv / 2008, rock, moderate current, Salles F. F., Massariol, F. C., Boldrini, R., Lima, M. M. & Britto, P. V. A. cols. Slide of nymph PT 09, Brazil, Espírito Santo, Espera Feliz, Rio São Domingos, Cachoeira do Aurélio (20 ° 28 ’ 57.5 ’’ S, 41 ° 50 ’ 14.5 ’’ W 1800 m), 22 / IV / 2008. Slide of nymph PT 11, Brazil, Espírito Santo, Espera Feliz Rio São Domingos, Cachoeira Sete Pilões (20 ° 28 ’ 57.1 ’’ S, 41 ° 49 ’ 50.4 ’’ W 1850 m), 25 / III / 2009.	en	Amaral, I. F., Cifuentes-Vargas, E. T., Salles, F. F. (2025): Cloeodes tovauna: a new species previously identified as Cloeodes aymore Massariol & Salles, 2011 (Ephemeroptera: Baetidae). Zootaxa 5594 (2): 355-366, DOI: 10.11646/zootaxa.5594.2.7, URL: https://doi.org/10.11646/zootaxa.5594.2.7
03A16432FFE8A8215DAACC0E8954F80C.taxon	etymology	Etymology. The specific epithet is a combination of words derived from the Tupi-Guarani: ‘ tova’ (face) and ‘ una’ (black), alluding to the color pattern of the head. Name in apposition.	en	Amaral, I. F., Cifuentes-Vargas, E. T., Salles, F. F. (2025): Cloeodes tovauna: a new species previously identified as Cloeodes aymore Massariol & Salles, 2011 (Ephemeroptera: Baetidae). Zootaxa 5594 (2): 355-366, DOI: 10.11646/zootaxa.5594.2.7, URL: https://doi.org/10.11646/zootaxa.5594.2.7
03A16432FFE8A8215DAACC0E8954F80C.taxon	distribution	Distribution. Southeastern Brazil, Minas Gerais State (Araponga, Catas Altas, Ouro Preto); Espírito Santo State (Alto Caparaó, Espera Feliz).	en	Amaral, I. F., Cifuentes-Vargas, E. T., Salles, F. F. (2025): Cloeodes tovauna: a new species previously identified as Cloeodes aymore Massariol & Salles, 2011 (Ephemeroptera: Baetidae). Zootaxa 5594 (2): 355-366, DOI: 10.11646/zootaxa.5594.2.7, URL: https://doi.org/10.11646/zootaxa.5594.2.7
