taxonID	type	description	language	source
03A3AD0FFF82FFE3A0A0FD7CFB72FCDA.taxon	description	(Fig. 1)	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF82FFE3A0A0FD7CFB72FCDA.taxon	materials_examined	HOLOTYPE. — Madagascar • Tsimbazaza; on soil; 3. VI. 1976; P. Tixier 8027; Holotype: MNHN [PC 0786650] (mixed with F. rosulatus P. de la Varde).	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF82FFE3A0A0FD7CFB72FCDA.taxon	diagnosis	DIAGNOSIS. — This new species is characterized by the completely limbate vaginant laminae on all leaves, the limbidia frequently extending onto the apical lamina; the laminal cells tiny, smooth, convex; the costae excurrent; and axillary antheridial buds frequent. Fissidens colliculosus Brugg. - Nann., sp. nov., is most likely to be confused with other semilimbate species with convex or mammillose cells. Of these, F. submarginatus Bruch in Krauss differs in its highly and sharply mammillose cells, and F. subundatus Dury in its convex cells each with a small papilla (those of the new species are convex and smooth), and its corticolous habitat. Fissidens reflexus Hampe and F. subscleromitrius Bizot & Dury have shorter limbidia.	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF82FFE3A0A0FD7CFB72FCDA.taxon	etymology	ETYMOLOGY. — The occurrence of small, convex, smooth cells is remarkable in a subgen. Polypodiopsis species that typically have pluripapillose, mammillose or smooth, ± plane cells. The species is named for these convex cells that produce a somewhat colliculose (collis: hill; - osus: rich in) leaf surface.	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF82FFE3A0A0FD7CFB72FCDA.taxon	description	DESCRIPTION Stem with central strand, unbranched to heavily branched, not heterocaulous, pinnately foliated, 5 - 11 × 1 - 2 mm, rhizoids brown, smooth; axillary nodules not differentiated; leaves distant to close, up to 16 pairs, crispate when dry, soft, lanceolate to elliptic-lanceolate with an acute, acuminatemucronate apex, 1.0 - 1.5 × 0.3 - 0.4 mm, 3 - 4.5 times as long as wide, margin subentire where elimbate; limbidia on the vaginant laminae of all leaves, extending the complete length of the vaginant lamina, frequently extending onto the apical lamina, confluent at the apex of the vaginant lamina or not, up to 16 µm wide, marginal, unistratose; vaginant lamina 2 / 3 - 3 / 4 the leaf length, subequal, near the insertion about as wide as the stem, unistratose; dorsal lamina variable below, often rounded, plane, less often undulate, reaching the insertion, rarely decurrent; dorsal and apical lamina unistratose, unito bistratose near the costa; costa excurrent, bryoides-type; lumina mid dorsal laminal cells 3.5 - 6.5 × 2.5 - 5.5 µm, walls thin, up to 1 µm diam., convex, smooth; lumina mid vaginant laminal cells 4.5 - 7.0 × 2.5 - 5.5 µm, walls up to 1 µm diam., convex, smooth. Gemmae not seen. Perichaetia and perigonia on the same or on different stems, perigonia terminal on stems and branches; perigonial branches bud like and short, ± 0.5 mm long (stalk included) (Fig. 1 D) to branch like and to up to 0.9 mm long, antheridia 170 - 200 µm long; perichaetia terminal, perichaetial leaves 1.3 - 1.9 mm long, archegonia 250 µm long; calyptra smooth, 0.6 mm long. Sporophyte, seta 4 mm long, smooth, 1 per perichaetium; theca erect, cylindric, 0.8 - 1.0 × 0.3 mm, ± 32 columns of oblong exothecial cells; peristome-type unknown, tooth base 48.5 µm wide; operculum rostrate, 0.4 mm long; spores not observed. The semilimbate leaves and the low number of columns of exothecial cells indicate that the new species belongs to subgenus Polypodiopsis (Müll. Hal.) Broth. This subgenus is characterized by c. 32 columns of exothecial cells and probably scariosus-type peristomes. However, since the peristome teeth of F. colliculosus Brugg. - Nann., sp. nov., are broken the type cannot be determined with certainty.	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF82FFE3A331FC1DFAEBF85F.taxon	description	(Fig. 2)	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF82FFE3A331FC1DFAEBF85F.taxon	materials_examined	— Lectotype (designated by Slavick 1992 as holotype): Niger Expedition; Barter 1425 (NY, mixed with e. g. F. porrectus Mitt.; isolectotypes: PC, BM).	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF82FFE3A331FC1DFAEBF85F.taxon	materials_examined	— Lectotype (designated by Bruggeman-Nannenga 1993, here refined with barcode): [Sudan], Central-Africa • im Lande der Bongo bei Kulongo der Höhle von Gubbihih; Schweinfurth 2234 MNHN [PC 10106795]; isolectotypes: PC and H-BR as Schweinfurth s. n., syn. nov.	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF82FFE3A331FC1DFAEBF85F.taxon	materials_examined	— Holotype: Madagascar • au nord de Maromandia; au bord d’un marigot permanent, sous forêt, sur un arbre; 50 m alt.; Cremers 2734; MNHN [PC 0129997]; isotype: BR, syn. nov.	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF82FFE3A331FC1DFAEBF85F.taxon	diagnosis	DIAGNOSIS. — Fissidens pellucidus var. ripensis differs from F. p. var. pellucidus by mucronate apices and taller plants with up to 55 leaf pairs. Leaves of the variety can be elimbate, but typically have weak limbidia on the basal parts of some or all or most vaginant laminae. Leaves of var. pellucidus are as a rule elimbate. Most collections of var. ripensis are from semiaquatic places, but it is not restricted to these.	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF82FFE3A331FC1DFAEBF85F.taxon	materials_examined	OTHER EXAMINED MATERIAL (Specimens used in Fig. 2). — Madagascar • Atsinanana (Toamasina); Secondary forest, on rock; 17 ° 42 ’ 1 ” S, 49 ° 27 ’ 18 ” E; Brinda et Andramiarisoa 12757 (MO, private herbarium Bruggeman-Nannenga) • Tamatave, Forest Reserve atTampolo, 95 km N of Tamatave; on root in boggy area; 17 ° 6 ’ S, 49 ° 24 ’ E; Crosby 6700 (PC, private herbarium Bruggeman-Nannenga) • SW Madagascar, Isalo National park, ‘ Piscine Naturelle’ 5 km W of Ranohira; sandstone canyon with Pandanus near the streamlet; Orban 9455 / CR; EGR; type specimen of F. cremersii MNHN [PC 0129997].	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF80FFE1A3A9FC5DFBB5FAD6.taxon	materials_examined	— Lectotype (designated as holotype byMagill 1981, here refined with barcode): South Africa • Port St John’s. 1920; Wager 927; BM [BM 000871817]; isolectotype: PRE.	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF80FFE1A3A9FC5DFBB5FAD6.taxon	materials_examined	— Holotype: Zambia • Lusaka Distr.; on damp vertical earth bank in Syzygium thicket by Bothasrus road 8 km from the Wilfred Watson turn-off from the Lusaka / Mumbwa road; c. 1275 m a. s. l.; 10. II. 1975; Townsend 75 / 21; L; isolectotype: E, syn. nov.	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF80FFE1A14CFF7EFADFFCDA.taxon	description	(Fig. 3)	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF80FFE1A14CFF7EFADFFCDA.taxon	materials_examined	HOLOTYPE. — Kapverden • Santo Antão, ribeira das Pombas; oberer Bereich, auf feuchten Steinen im obersten Bereich der Pflanzungen, nordostexponiert; 260 m alt.; 10. II. 1995; Lindlar 405; L.	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF80FFE1A14CFF7EFADFFCDA.taxon	diagnosis	DIAGNOSIS. — This subspecies is characterized by the pale green color of young plants, the mostly distant, lanceolate leaves; the characteristic (but variable) apex, typically narrow and terminated by a large, pointed cell; rounded dorsal lamina bases, limbidia restricted to the leaves of vaginant laminae of perichaetial stems; laminal cells with 2 - 4 punctate papillae over the lumen; slightly to half-open vaginant laminae and costae that end 2 - 10 cells below the apex. The antheridia are terminal on short stems and branches, and the leaves are inflexed with a slightly incurved apex when dry. The new variety differs from subsp. sciophyllus in the typically slender, tapering leaf tips and shorter costae that do not reach the apex and may end as many as 10 cells below it. Leaf tips of var. sciophyllus are mostly less narrowly acute with percurrent to shortly excurrent costae. At first sight F. s. subsp. capeverdensis may resemble the smooth-celled F. pellucidus. Both are pale green when young and have distant, lanceolate leaves. However, the pluripapillose laminal cells of the new subspecies make distinction easy.	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF80FFE1A14CFF7EFADFFCDA.taxon	etymology	ETYMOLOGY. — This subspecies is endemic to the Cape Verde Islands. This is reflected in the name.	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF80FFE1A14CFF7EFADFFCDA.taxon	distribution	DISTRIBUTION. — Endemic to the Cape Verde Islands where it known from Santo Antão, Brava, Fogo, Santiago and São Nicolau.	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF80FFE1A14CFF7EFADFFCDA.taxon	description	SUBSTRATE. — Stones, less often soil. ELEVATION. — 260 - 1220 m alt.	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF80FFE1A14CFF7EFADFFCDA.taxon	materials_examined	PARATYPES. — Kapverden • Brava, an der Strasse zum Flughafen; c. 600 m alt.; hinter Cova Rodela. Felsrinne, nordexponiert; 580 m alt.; 30. I. 1995; Lindlar 91 (private herbarium Bruggeman-Nannenga) • Santiago, Serra de Malagueta; Ost- bzw. Nordosthänge im Südwesten von Cha de Figueiras, auf Erde und Gestein; ost- nordostexponiert; 770 - 800 m alt.; 1. II. 1995; Lindlar 107 (private herbarium Bruggeman-Nannenga) • Santo Antão, ribeira das Pombas; oberer Bereich, auf feuchten Steinen im obersten Bereich der Pflanzungen; nordostexponiert; 260 m alt.; 10. II. 1995; Lindlar 404 and 410 (all private herbarium Bruggeman-Nannenga).	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF80FFE1A14CFF7EFADFFCDA.taxon	discussion	REMARKS Leaf tips vary from tips that, apart from the shorter costa, resemble those of subsp. sciophyllus (Fig. 3 F) to long, slender and attenuated (Fig. 3 G). Some of this variability is shown in Fig. 3 F-I. Earlier reported from the Cape Verde islands as F. sciophyllus (Frahm et al. 1996) and F. danckelmannii Müll. Hal. (Bizot 1969; Potier de la Varde 1943).	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF80FFE1A14CFF7EFADFFCDA.taxon	description	DESCRIPTION BASED ON LINDLAR 405 (HOLOTYPE), LINDLAR 404 (CROSS-SECTIONS) AND LINDLAR 107 (PERIGONIA) Growing in loose mat. Stem with central strand, not heterocaulous, unbranched, 4 - 5 × 1.5 mm, pinnately foliated; rhizoids brown, smooth; axillary nodules not differentiated; leaves pale green, distant to close, up to 13 pairs, reflexed when dry, lanceolate with a long, narrow, acute-acuminate apex ending in a large pointed cell, 0.7 - 1.1 × 0.2 - 0.3 mm, 3.6 - 4.7 times as long as wide, margin subentire to crenulate, elimbate except for the vaginant laminae of the 4 upper leaf pairs of perichaetial stems that have a unistratose (estimated), more or less marginal limbidium; limbidium extending up to 1 / 2 the vaginant lamina in perichaetial leaves; vaginant lamina 1 / 2 - ª ⁄ 5 the leaf length, rounded and less wide than the stem at the insertion, unistratose, unequal with the minor lamella ending ± midway between costa and margin; dorsal lamina rounded and slightly undulate at the insertion, reaching the insertion, not decurrent; dorsal and apical lamina unistratose; costa ending 2 - 7 (up to 10 in some collections) cells below the apex, bryoides-type; mid dorsal laminal cells (4.5 -) 6 - 8 × (3 -) 4.0 - 7 µm, with 2 - 4 tiny punctate papillae; mid vaginant laminal cells 7 - 9.5 (- 11) × (4.5 -) 5.5 - 8 µm, plane, with 3 - 4 punctate papillae over the lumen. Gemmae not observed. Perigonia terminal on 1 - 1.5 mm long stems and on branches; antheridia c. 150 µm long; perichaetia terminal, perichaetial leaves c. 1.5 mm long, archegonia c. 300 µm long; calyptra not observed. Sporophyte, seta 4 mm long, smooth, theca erect, ± cylindric, 0.6 × 0.3 mm, ± 32 columns of collenchymatous exothecial cells; peristome type unknown, tooth base c. 32 µm wide; operculum not observed; spores not observed.	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF80FFEFA3DBFA1BFBAFFCFA.taxon	description	(Fig. 4)	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF80FFEFA3DBFA1BFBAFFCFA.taxon	materials_examined	— Lectotype (designated byTouw 2013): Ceylon [Sri Lanka] • König 910, 114 - 864 = L 0041421 (herb. Van Royen; labelled by Sande Lacoste; analytic drawings by Iwatsuki).	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF80FFEFA3DBFA1BFBAFFCFA.taxon	materials_examined	— Lectotype (designated as holotype byMagill 1981, here refined as “ specimen in herbarium Hampe ”): South Africa • Natal, prope Umpumulo; Borgen 7; BM-herb. Hampe; isolectotypes: BM, PC, H-SOL, syn. nov.	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF80FFEFA3DBFA1BFBAFFCFA.taxon	materials_examined	— Lectotype: (designated by Bruggeman-Nannenga 1993, here refined with barcode): Musci Africani in Camerunia a P. Dusén collecti coll. a 13, BM [BM 000879000]; isolectotype: BM, syn. nov.	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF80FFEFA3DBFA1BFBAFFCFA.taxon	materials_examined	— Lectotype (designated by Bruggeman-Nannenga 1993 here refined with barcode): Gabon • Dibwangui; Le Testu 5631; MNHN [PC 0106495], syn. nov.	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF80FFEFA3DBFA1BFBAFFCFA.taxon	materials_examined	— Lectotype (Bruggeman-Nannenga 1993 here refined with barcode): Gabon • dans la Basomba; Le Testu 5633; MNHN [PC 0106554]; isolectotype: PC, syn. nov.	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF80FFEFA3DBFA1BFBAFFCFA.taxon	materials_examined	— Lectotype (designated by Bruggeman-Nannenga 1993 here refined with barcode): Central African Republic • Mbaïki; Tisserant M 738; MNHN [PC 0029462]; isolectotype: PC, syn. nov.	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF80FFEFA3DBFA1BFBAFFCFA.taxon	materials_examined	— Lectotype (designated by Bruggeman-Nannenga 1997 here refined with barcode): Burkina Faso • Haute Volta; chutes de la Comoe près de Banfora; Chevalier s. n.; PV 9935; MNHN [PC 0095359]; isolectotype: MNHN [PC 0095358], syn. nov.	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF80FFEFA3DBFA1BFBAFFCFA.taxon	materials_examined	— Lectotype (designated by Bruggeman-Nannenga 1993 here refined with barcode): Nigeria • Jos; Thorold 306; MNHN [PC 0095365]; isolectotype: MNHN [PC 0073009], syn. nov.	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF80FFEFA3DBFA1BFBAFFCFA.taxon	materials_examined	— Lectotype (designated as holotype by Bruggeman-Nannenga 1999, here refined with barcode): Seychelles • Mahé, Evéché de Victoria, la Rosière; sur les murs de granit ombragés; 20 m alt.; 20. I. 1974; Onraedt 75 - S- 003; MNHN [PC 0096369].	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF80FFEFA3DBFA1BFBAFFCFA.taxon	diagnosis	DIAGNOSIS (distinctive features bold). — Fissidens ceylonensis is a variable, palaeotropical species characterized by stiff, opaque leaves, marginal papillae that obscure the cells, long vaginant laminae reaching 2 / 3 - ª ⁄ 5 the leaf length and irregular, typically intramarginal, limbidia. The distribution and extension of the limbidia is variable. They vary between restricted to the upper leaves of perichaetial stems (e. g. in the type specimen of F. sarcophyllus) and present on the vaginant laminae of all or most leaves of both vegetative and perichaetial stems. Limbidia can exist of a single cell or extend over a large part of the vaginant lamina, but always end below the apex of the vaginant lamina. Limbidia are frequently interrupted, irregular and typically at least in part intramarginal, limbidia of perichaetial leaves intramarginal or marginal. Limbidia are unistratose or, rarely, partly bistratose. The shape of the leaves varies between lanceolate and oblong-lanceolate. The costa typically is percurrent to excurrent in short mucro. Rarely it ends a few cells below the apex (e. g. in the type specimen of F. borgenii). The African F. schweinfurthii resembles F. ceylonensis in the obscure, stiff leaves, long vaginant laminae, marginal papillae, lanceolate to oblong leaves and percurrent to excurrent costae. This species is easily separated by its better developed, limbidia that are present on the vaginant laminae of all leaves, reach the apex of the vaginant laminae and are marginal throughout, except sometimes for a short intramarginal piece near the insertion.	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF80FFEFA3DBFA1BFBAFFCFA.taxon	materials_examined	OTHER EXAMINED MATERIAL (Specimens used in Fig. 4). — Democratic Republic of the Congo • Bas-Uele; 5 km à l’ouest de Likali, termitière dans la forêt secondaire; Lisowski 47587; KRAM (private herbarium Bruggeman-Nannenga). Ghana • Central Region, Akroform-Akwekrom, Nimere R.; riverside rocks under roadbridge; 5 ° 27 ’ 760 N, 1 ° 30 ’ 236 W; R. Agyei, G. Ameka, C. Dankwah & N. G. Hodgetts 10217 (private herbarium Hodgetts; small collection mixed with other species). Guinea • Nzérékoré Region, Macenta Prefecture, above the village of Zoubrouma; rocks in the forest; Lisowski 129 and 225; KRAM (private herbarium Bruggeman-Nannenga). Seychelles • West coast of southern Mahé island; Point Maravi at the S end of Anse Gaulettes beach; Carophyllum, coconut and seminatural thicket on granitic rocks, rupicolous; 4 ° 41 ’ S, 55 ° 28 ’ 35 ” E; Kis 9329 / X; EGR, L • Northern tip of Mahé Island, Montagne Glacis above Rivière Glacis valley; plantation forest and thickets, rupicolous; 4 ° 34 ’ 30 ” S, 55 ° 26 ’ 25 ” E; 4 ° 34 ’ 40 ” S, 55 ° 26 ’ 35 ” E; Kis 9340 / BM; EGR, L. DISTRIBUTION. — This is the first record for mainland Africa of this species under the name F. ceylonensis. Formerly, African specimens were known as e. g. F. borgenii. Further distribution: Seychelles, Asia, Polynesia and Australia.	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF80FFEFA3DBFA1BFBAFFCFA.taxon	discussion	REMARK All new synonyms above were earlier (Bruggeman-Nannenga 1993, 1997) reduced to F. borgenii and thus are no more than the logical result of the synonymy of F. borgenii with F. ceylonensis. This list of synonyms is not complete as only African synonyms are included.	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF8EFFEFA387FC3DFB5CF9D9.taxon	materials_examined	— Lectotype (designated here): Uganda • Ankole district, Moutozho; sur rochers ensolleillées; 1300 m alt.; XII. 1967; Balázs 96 G; MNHN [PC 0073300].	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF8EFFEFA387FC3DFB5CF9D9.taxon	discussion	REMARKS The lectotypification of this species is rather complicated. On page 11 of Bizot (1973) is a footnote that says that holotypes of new species described in the paper are in DI (where Bizot used to work; after his death in 1979 his herbarium went to PC) and isotypes in EGR. Thus Pócs’ 1977 ‘ typification’ of F. cuynetii that says holotype in EGR, isotype in PC, is not effective because it is in contradiction with the protologue. A further complication is that the protologue gives the collection number Balázs 96, whereas in Bizot’s herbarium in PC there are Balázs 96 D MNHN [PC 0073299] and Balázs 96 G MNHN [PC 0073300]. Balázs 96 G MNHN [PC 0073300] is here designated lectotype because it has a note in Bizot’s handwriting saying “ sp. nov. ” and all information on its label is by Pócs (who donated the material to Bizot for identification), whereas the information on 96 D is just a handwritten copy of MNHN [PC 0073300] by Bizot.	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF8EFFEDA3A1F918FD05FDE5.taxon	materials_examined	— Lectotype (designated here): Mayotte • Marie 206 (partly unreadable): BM [BM 000871897]; isolectotypes: MNHN [PC 0129799].	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF8EFFEDA3A1F918FD05FDE5.taxon	description	Fissidens alomoides Müll. Hal. ex Dusén, Kongliga Svenska Vetenskaps Academiens Handlingar 28 (2): 10, 11: f. a-b, pl. II, f. 3. (Dusén 1895).	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF8EFFEDA3A1F918FD05FDE5.taxon	materials_examined	— Lectotype (designated here): Cameroon • Dusén; Musci Afr. Camerunia nr. 505; S [B 107445]; isolectotypes: H-BR, MNHN [PC 0029441] and S [B 107447, B 107448, B 107449, B 107450], syn. nov. Isotypes have been distributed as: Musci Africani in Camerunia a P. Dusén collecti 505 (also miswritten as 305).	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF8EFFEDA3A1F918FD05FDE5.taxon	description	DESCRIPTION Fissidens hymenodon is a corticolous species that is characterized by oblong to oblong-lanceolate leaves with broadly acute to obtuse, often rounded apices, apex not terminated by a pointed cell, costae ending 2 - 6 cells below the apex, limbidia restricted to the upper leaves of perichaetial leaves, 1 - 2 papillose laminal cells and scariosus-type peristomes.	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF8CFFEDA16BFA5BFB55FB59.taxon	description	(Fig. 5)	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF8CFFEDA16BFA5BFB55FB59.taxon	materials_examined	— Lectotype (designated here): Madagascar • Pervillé 51 A; BM [BM 000871895] mixed with the lectotype of F. madecassus Schimp. ex Müll. Hal.; isolectotype: BM [BM 000871896]; two stems mixed between six stems of the isolectotype of F. madecassus.	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF8CFFEDA16BFA5BFB55FB59.taxon	materials_examined	— Lectotype (designated here): Uganda • Musandama; Budd s. n.; 1926; BM [BM 000871873]; isolectotype: MNHN [PC 0095552], syn. nov.	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF8CFFEDA16BFA5BFB55FB59.taxon	materials_examined	— Lectotype (designated byPócs 1977 as holotype): Tanzania • Morogoro District, Uluguru Mts, on top of Bondwa Peak; on the ground of subalpine heath; 2100 m alt.; 12. X. 1969; Pócs & Gibbon 6052 / CR (EGR; isolectotypes: MNHN [PC 0132125] and DSM), syn. nov.	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF8CFFEDA16BFA5BFB55FB59.taxon	diagnosis	DIAGNOSIS (distinctive features in bold). — Plants often rusty brown, leaves narrow, 3.5 - 5 (- 6) times as long as wide, lanceolate with narrow acute to acute-acuminate tips; limbidium thick yet unistratose (Fig. 5 J), smooth to denticulate, marginal, present on the vaginant laminae of all leaves, typically extending the whole length of the vaginant lamina, often confluent at the apex and frequently extending onto the apical lamina; vaginant lamina 2 / 3 - 3 / 4 the leaf length; costa percurrent to excurrent, or ending 1 - 2 cells below the apex; laminal cells can have 1 - 2 or 2 - 4 papillae per cell (rarely 1 (- 2), or 2 - 6), punctate, less often irregular (Fig. 5 I) papillae; antheridia terminal on short stems (rhizautoicous), less often terminal on longer stems or on short perigonial branches in the axils of lower leaves; setae 3 - 10 mm long, theca cylindric to strumulose, with 32 - 58 columns of exothecial cells. Some collections have thecae with ± 32 as well as with 40 or more exothecial cells. Since ± 32 columns are typical for subg. Polypodiopsis 32 is not conclusive. When, however, there are 40 or more exothecial cells, this is diagnostic for F. ferrugineus.	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF8CFFEDA16BFA5BFB55FB59.taxon	materials_examined	OTHER EXAMINED MATERIAL (Specimens used for Fig. 5). — Madagascar • Pervillé 51 A; BM [BM 000871895], lectotype F. ferrugineus. Tanzania • Meru Mts; in the crater ground near Njeku; swampy moorland patches, rupicolous; 2600 m alt.; Pócs et al. 8670 / D; EGR, L • Kilimanjaro Mts, Karanga River Gorge N of Kibosho; Montane evergreen forest on the steep; rocky E facing slope; with Albizia gummifera (J. F. Gmel.) C. A. Sm.; on soil covered rock; 1880 m alt.; Pócs, Mialla & Margesi 90068 / G; EGR • Southern Highlands, Kipengere Range, above Salala Falls below Mt Ishinga; in deep ravine of streamlet, on half shady rock; 2380 - 2400 m alt.; 26. V. 1987; Pócs, Iversen, Persson & Petterson 87146 / O; EGR, private herbarium Bruggeman-Nannenga • Ukaguru Mts in Kilos District, on Mamiwa ridge; mossy elfin forest, rupicolous; 2150 m alt.; Pócs, Chamshama & Sylvander 86100 / T; EGR, L • Ukaguru Mts, NE slope of Mamwira W of Masenge village; on shady rocks in bamboo (Arundinaria alpina K. Schum.), thicket; 2050 m alt.; Pócs 88015 / C; EGR. La Réunion • G. de L’ Isle s. n. (PC) • Forêt de Bélouve on the Plateau des Cochons, Trou de Fer trail; 21 ° 40 ’ S, 55 ° 32.3 ’ E; 21 ° 40 ’ S, 55 ° 32.4 ’ E; Bruggeman-Nannenga 10995; L.	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF8CFFEDA0F2FD3FFE2AFA18.taxon	description	Fissidens ferrugineus and F. schweinfurthii were considered synonyms of F. intramarginatus (Bruggeman-Nannenga 1993, 1997). Fissidens intramarginatus s. l. is characterized by pluripapillose laminal cells and vaginant laminae that are limbate in all leaves of both perichaetial and vegetative stems. The limbidia nearly or completely reach the apex of the vaginant laminae and frequently extend onto the apical lamina. It comprises several expressions that are linked by transitional forms (species-complex). Meanwhile, it has become clear that the neotropical concept of F. intramarginatus (Pursell 2007) is narrower than the African concept (Bruggeman-Nannenga 1993, 1997) and that a revision is needed. In anticipation of a revision of the African species of this complex (in preparation). F. ferrugineus and F. schweinfurthii are here reinstated as species. As a rule, these two species are easily distinguished by the papillae. Fissidens schweinfurthii has 5 - 8 marginal (i. e. distributed on or near the cell walls in surface view) papillae that obscure the cells, whereas F. ferrugineus has a variable, lower number (1 - 2 or 2 - 4 per cell) of punctate to irregular papillae over the lumen and more or less clear cells.	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF8CFFE9A3FBFA9AFEB2FA6F.taxon	description	(Fig. 6)	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF8CFFE9A3FBFA9AFEB2FA6F.taxon	materials_examined	— Lectotype (designated by Bruggeman-Nannenga 1997, here refined with barcode): Sudan as Niam-niam-regiones [border area Sudan, Democratic Republic of the Congo and Central African Republic] • in cacumine collis Gumango; 6. II. 1870; Schweinfurth s. n.; MNHN [PC 0095370]; isolectotype: MNHN [PC 0129867].	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF8CFFE9A3FBFA9AFEB2FA6F.taxon	materials_examined	— Lectotype (designated by Bruggeman-Nannenga 1997 here refined with barcode): Madagascar • Nossi Bé, Marie s. n.; BM [BM 001094101]; isolectotypes: H-BR, PC, syn. nov.	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF8CFFE9A3FBFA9AFEB2FA6F.taxon	materials_examined	— Lectotype (designated here): Madagascar • Andriba; 1897; Dorr s. n.; MNHN [PC 0098324]; isolectotype: MNHN [PC 0098325], syn. nov.	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF8CFFE9A3FBFA9AFEB2FA6F.taxon	materials_examined	— Lectotype (designated by Bruggeman-Nannenga 1997, here refined with barcode): Madagascar • Ambohimanga; Décary 9; MNHN [PC 0096226], syn. nov.	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF8CFFE9A3FBFA9AFEB2FA6F.taxon	diagnosis	DIAGNOSIS (distinctive features in bold). — This African species is characterized by stiff plants, leaves close, in transmitted light obscure with pale, per- to short excurrent costa, vaginant laminae long, reaching 2 / 3 - ̚ ⁄ 10 the leaf length, proximally about as wide as the stem, with rounded insertion, limbidia reaching the insertion or ending some distance above, marginal throughout (Fig. 3 G, H) or intramarginal towards the insertion (Fig. 3 F), distally nearly or completely reaching the apex of the vaginant lamina; laminal cells with (4 -) 5 - 8 (- 10) well developed, low, blunt, marginal papillae (Fig. 6 I, J); archegonia typically terminal on short or long stems and / or on short branches; antheridia terminal on dwarf or larger stems, occasionally on short bud-shaped branches. Most specimens of this species are easily recognized by the bold printed traits. Fissidens ceylonensis similarly has obscure leaves, short excurrent to percurrent costae and marginal papillae, but differs in the shorter limbidia that do not reach the apex of the vaginant laminae, and are frequently lacking on some or most leaves.	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF8CFFE9A3FBFA9AFEB2FA6F.taxon	materials_examined	OTHER EXAMINED MATERIAL (specimens used forFig. 6). — Democratic Republic of the Congo • Haute-Shaba; Plateau des Kundelungu, partie supérieure de la Lofoi; Lisowski 4225; KRAM • Haute-Shaba, près de Likasi, rive gauche de la rivière Panda; gallerie forestière; Lisowski 65116; KRAM. Ivory Coast • Boundiali; nr 1 B 3; 9 ° 32 ’ N, 6 ° 29 ’ W; Granitinselberg, auf beschattenen Fels; Porembski & Biedinger 1508; B, private herbarium Bruggeman-Nannenga • Westl. von Bouaké. Granitinselberg (nr IB 24), terrestrisch; in Afrotrilepis Matte; 370 m alt.; Porembski 515; private herbarium Bruggeman-Nannenga. Madagascar • Prov. Majunga; Station forestière d’Ampijoroa, c. 3 km N d’Andranofasika; on soil, base of tree in forest; 16 ° 20 ’ S, 46 ° 51 ’ E; Dorr 3055; MO, L • 60 km NW of Antanannarivo along Mahajanga road; N of Soaniadanana village; granite outcrops in secondary plateau grassland, rupicolous-terricolous; 18 ° 29 ’ 54 ” S, 47 ° 17 ’ 02 ” E; 5. IX. 1994; Kis 9443 EE; L, EGR. Malawi • Southern District, Zomba Plateau; Long 12529; E; private herbarium Bruggeman-Nannenga. Nigeria • Ogun State, Ijebe-Ode; savanne area S of Idowa, bank in partial shade; McFarlane 275; private herbarium Bruggeman-Nannenga, a few mixed between F. pellucidus Hornsch. Sudan • Schweinfurth s. n.; type specimen F. schweinfurthii MNHN [PC 0095370]. Togo • circ. De Klouto; an der Strasse Palimé-Klouto, etwa auf der halben Höhe des Gebirgsrandes, oberhalb des Wasserfalls; erdig-felsige Böschung; 300 - 400 m alt.; 8. X. 1973; Hiepko & Schultze-Motel 393; NY, MO, L.	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF88FFF7A134F9BBFEBDF859.taxon	description	(Fig. 7)	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF88FFF7A134F9BBFEBDF859.taxon	materials_examined	— Lectotype (designated by Slavick 1992 as holotype): Niger - exp. • Barter 1425; NY, isolectotype: PC, BM, H-BR, S.	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF88FFF7A134F9BBFEBDF859.taxon	materials_examined	— Lectotype (designated byMagill 1981, here refined with barcode): South African Republic • Rehmann, Musci austro-africani (1875 - 77): 289; BM [BM 000871824]; isolectotypes: have been distributed as Dr. A. Rehmann, Musci austro-africani (1875 - 77): 289; BM [BM 000871823], PC, S.	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF88FFF7A134F9BBFEBDF859.taxon	materials_examined	— Lectotype (designated by Bruggeman-Nannenga 1993, here refined with barcode): Tanzania • Lutindi; am Boden im Hochwald; Holst 3472; p. p. BM [BM 000871825].	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF88FFF7A134F9BBFEBDF859.taxon	materials_examined	— Lectotype (designated here): Madagascar • Mosses of Nossi Bé; on earth; Marie s. n.; BM [BM 001094100]; isolectotypes: PC, H-BR.	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF88FFF7A134F9BBFEBDF859.taxon	materials_examined	— Lectotype (designated here): Democratic Republic of the Congo [Congo belge] • Sankuru; leg. Em et M. Laurent; 1903; MNHN [PC 0106533]; isolectotypes: MNHN [PC 0106532], HBR.	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF88FFF7A134F9BBFEBDF859.taxon	materials_examined	— Lectotype (designated here): Gabon • Evoungoun; 9. VII. 1925; Le Testu 5374; PV 2221; MNHN [PC 0106481]; isolectotypes: MNHN [PC 0106478, PC 0106480] and H-BR.	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF88FFF7A134F9BBFEBDF859.taxon	materials_examined	— Lectotype (designated here): Gabon • Idembé; Le Testu 6840; MNHN [PC 0106558]; isolectotypes: in PC, BR, BM.	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF88FFF7A134F9BBFEBDF859.taxon	materials_examined	— Lectotype: (designated here): Gabon • Chute de la Moungoungoulou; Le Testu 6866; the number in the protologue (6668) must be an orthographic error and is here corrected (ICNB art. 9.2): MNHN [PC 0106519]; PV 3974; isolectotype: MNHN [PC 0129748].	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF88FFF7A134F9BBFEBDF859.taxon	materials_examined	— Lectotype (designated here): Gabon • l’Ofoué; Eckendorff s. n.; PV 4450; MNHN [PC 0106556]; isolectotypes: PC, NY.	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF88FFF7A134F9BBFEBDF859.taxon	materials_examined	— Lectotype (designated by Bruggeman-Nannenga 1993 here refined with barcode): Gabon • Mont Loumanzoc; Le Testu s. n.; PV 5205; MNHN [PC 0106516]; isotype: MNHN [PC 0106515].	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF88FFF7A134F9BBFEBDF859.taxon	materials_examined	— Lectotype (designated by Bruggeman-Nannenga 1993, here refined with barcode): Gabon • Rocher de Nzang, Nzoumou; Le Testu s. n.; PV 5443; MNHN [PC 0106518]; isolectotype: MNHN [PC 0106517].	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF88FFF7A134F9BBFEBDF859.taxon	materials_examined	— Lectotype (designated here): Guinea • Ziama; Schnell 2620; PV 9954; MNHN [PC 0106522]; isolectotypes: BM, MNHN [PC 0106520; PC 0106521].	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF88FFF7A134F9BBFEBDF859.taxon	materials_examined	— Lectotype (designated by Bruggeman-Nannenga 1993 here refined with barcode): Uganda • Distr. Sese; Wood 1048; PV 14077; MNHN [PC 0106246]; isolectotypes: MNHN [PC 0106244; PC 0106245].	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF88FFF7A134F9BBFEBDF859.taxon	diagnosis	DIAGNOSIS (most important traits in bold). — Fissidens porrectus Mitt. is variable species. It is characterized by guttulate laminal cells (Fig. 7 M) with smooth, firm walls. These two traits are fairly constant. Furthermore, the dorsal and apical lamina typically have a characteristic incrassate border zone (Fig. 7 B, F) while the vaginant lamina is typically limbate (a limbidium is a border of narrow elongated cells). The last two traits, the incrassate border zone and the limbate vaginant lamina, are rather variable. The incrassate border can be (uni-) 2 - 4 - stratose, the cells forming it are usually short and wide (Fig. 7 B, F), less often, elongated (Fig. 7 K). The limbidia on the vaginant laminae are usually well developed and composed of elongated cells, less often they are indistinct (without or with only a few slightly elongated cells). Plant size and development of the limbidia and of the border zone are correlated, the largest plants having the best developed limbidia and the most elongated cells in the border zone (Fig. 7). However, in several specimens this correlation is not apparent. Moreover, irregularities of the border zone are not uncommon. In spite of its confusing variability, F. porrectus is usually easily recognized by the bold printed traits. It is most likely to be confused with F. splendens Brugg. - Nann. which has similar cells and similar incrassate borders, but (contrary to F. porrectus) the border of the vaginant lamina in this species consists entirely of isodiametric cells. Fissidens pellucidus has similar cells and cell walls, but lacks a differentiated, bistratose border zone and limbidia or has its limbidia restricted to the vaginant laminae of upper leaves of perichaetial stems. Specimens with borders of elongated cells can be mistaken for subg. Fissidens species. Species of this subgenus, however, have different sporophytes and eguttulate cells. Fissidens porrectus strongly resembles the elimbate F. pellucidus that also has guttulate cells with firm and smooth cell walls. However, Fissidens pellucidus clearly belongs to subg. Polypodiopsis. While F. pellucidus is elimbate, F. porrectus varies from more or less elimbate to limbate on all laminae. Smooth laminal cells and limbidia on all laminae are characteristic of subgenus Fissidens Hedw. In other words, gametophytes of limbate expressions of F. porrectus (Fig. 5 N) correspond with subg. Fissidens, but expressions with limbidia restricted to the vaginant laminae or lacking fit better in subg. Polypodiopsis. Subgenus Fissidens is further characterized by a subg. Fissidens - sporophyte, i. e. a bryoides - type peristome and 40 or more columns of exothecial cells on the theca. The sporophyte of F. porrectus is clearly a subg. Polypodiopsis- type with ± 32 columns of exothecial cells and a scariosus type peristome. This settles the species in subg. Polypodiopis. This is supported by molecular data (Budke et al. 2023). The incrassate border zone of the dorsal and apical lamina is one of the most conspicuous features of this species. Similar borders are widespread, but rare in the Fissidentaceae and occur in all subgenera, except subg. Fissidens. Thus the taxonomic value of this feature is limited to species level.	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF88FFF7A134F9BBFEBDF859.taxon	materials_examined	OTHER EXAMINED MATERIAL (Specimens used forFig. 7). — Gabon • Evoungoun; 9. VII. 1925; Le Testu 5374; MNHN [PC 010648]; type F. lestestui. Madagascar • Heim s. n.; MNHN [PC 0773734 A]; Niger-Exp., Barter 1425; NY, type F. porrectus. Republic of São Tome & Principe • Principe Island, cross-country route across summit plateau of Mesa; mixed tropical hardwood cloud forest with sedge and fern understory; on volcanic bedrock in rivulet in filtered light; 01 ° 34 ’ 51.7 ” N, 07 ° 21 ’ 04.2 ” E; Shevock et al. 49960 (private herbarium Bruggeman-Nannenga). South African Republic • Rehmann; Musci austro-africani (1875 - 77): 289; BM [BM 000871824]; type F. nitens.	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF88FFF7A134F9BBFEBDF859.taxon	discussion	REMARKS All these names have at different times and in different combinations been considered synonyms (Bizot & Pócs 1979; Bizot et al. 1990; Bruggeman-Nannenga 1993). Thus, none of the above synonyms is truly new. The present concept embraces all previous ones.	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF96FFF7A3CBFF7EFBC9FCFA.taxon	materials_examined	— Holotype: Indonesia • (Moluccas) Saparoea; 1858 - 1860; De Vriese 910, 1141054 = L 0058502; herb. C. M. van der Sande Lacoste; type discussed by Touw 2013.	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF96FFF7A3CBFF7EFBC9FCFA.taxon	materials_examined	— Lectotype (designated here): Madagascar • Kiangara; été 1931; Grainville s. n.; PV 4037; MNHN [PC 0096206; PC 0095457], syn. nov.	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF96FFF7A3CBFF7EFBC9FCFA.taxon	discussion	REMARK ON THE TYPIFICATION Potier de la Varde specified his collections with PV-number. In his herbarium in PC are two collections of F. grainvillei both numbered PV 4037, and both annotated “ sp. nov. ” by Potier de la Varde. MNHN [PC 0095457] consists of a few stems plus description and MNHN [PC 0096206], a slide and drawing. They are clearly parts of the same specimen, viz. PV 4037. Therefore, they are together designated the lectotype (ICBN art. 8.3).	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF96FFF4A3D8FCDDFC80FD26.taxon	description	(Fig. 8)	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF96FFF4A3D8FCDDFC80FD26.taxon	materials_examined	— Lectotype (designated here): Madagascar • Antalaka; sur les arbres d’une forêt primaire littorale dégradée (l’écorce est couverte de terre); Onraedt 71 M 5644; MNHN [PC 0096355]; isolectotype: slide by Theo Arts in BR.	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF96FFF4A3D8FCDDFC80FD26.taxon	discussion	REMARK The protologue does not indicate where the type specimen is deposited. However, in her unpublished thesis (Dury 1974) the author states that the type is in Bizot’s herbarium (now in PC). It appears that in Bizot’s herbarium there is only one F. subundatus specimen with a label matching the protologue. That specimen is designated the lectotype.	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF96FFF4A3D8FCDDFC80FD26.taxon	description	DESCRIPTION (EMENDED) Corticolous, scattered between other mosses; in KOH brown; rhizoids brown, smooth; axillary nodules not differentiated; stem delicate, pale green, unbranched, 2.5 - 5.0 × 2.0 - 2.5 mm, only fertile stems seen; leaves close, 7 - 14 pairs, more or less crispate when dry, oblong-lanceolate, broadly acute with short acumen, 0.8 - 1.5 × 0.25 - 0.4 mm, 3 - 4 (- 4.5) as long as wide, margin finely serrulate where elimbate; limbidium on the vaginant laminae of all leaves, nearly or completely extending the length of the vaginant laminae, occasionally excurrent on the apical lamina, marginal, sometimes intramarginal over a short distance, 14.5 µm wide, unistratose (estimated); vaginant lamina 1 / 2 - ̚ ⁄ 10 the leaf length, slightly open, near the insertion about as wide as the stem, slightly rounded ending concave at the insertion, unistratose (estimated); dorsal lamina slightly rounded at the insertion, reaching the insertion, not decurrent, dorsal and apical lamina unistratose (estimated); costa percurrent; lumina of mid dorsal laminal cells, 3.5 - 7.5 × 3.0 - 5.5 µm, lowly convex with a small central papilla; lumina of mid vaginant laminal cells 5.0 - 8.0 (- 9.0) × 3.0 - 6.5 µm, lowly mammillose with a distinct, tiny papilla. Gemmae not observed. Cladautoicous, perigonia on short, short bud-shaped or longer c. 4.5 mm long branches, antheridia ± 150 µm long; perichaetia terminal, a few archegonia solitary in the axils of upper leaves of perichaetial stems, perichaetial leaves c. 1.3 mm long; archegonia 250 µm long. Sporophyte, seta 1.2 - 2.0 mm long, smooth, 1 per perichaetium; theca 0.45 × 0.2 mm, with circa 32 columns of quadratic-oblong, collenchymatous exothecial cells; peristome probably a scariosus type (only basal OP seen), tooth base 33.5 µm wide; operculum, calyptra and spores not observed. The semilimbate leaves, mammillose cells and the number of exothecial cells classify this species in subg. Polypodiopsis.	en	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
