identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03A3AD0FFF82FFE3A0A0FD7CFB72FCDA.text	03A3AD0FFF82FFE3A0A0FD7CFB72FCDA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Fissidens colliculosus Brugg. - Nann. 2025	<div><p>Fissidens colliculosus Brugg.-Nann., sp. nov.</p><p>(Fig. 1)</p><p>HOLOTYPE.— Madagascar • Tsimbazaza; on soil; 3.VI.1976; P.Tixier 8027; Holotype: MNHN [PC0786650] (mixed with F. rosulatus P. de la Varde) .</p><p>DIAGNOSIS. — This new species is characterized by the completely limbate vaginant laminae on all leaves, the limbidia frequently extending onto the apical lamina; the laminal cells tiny, smooth, convex; the costae excurrent; and axillary antheridial buds frequent. Fissidens colliculosus Brugg.-Nann., sp. nov., is most likely to be confused with other semilimbate species with convex or mammillose cells. Of these, F. submarginatus Bruch in Krauss differs in its highly and sharply mammillose cells, and F. subundatus Dury in its convex cells each with a small papilla (those of the new species are convex and smooth), and its corticolous habitat. Fissidens reflexus Hampe and F. subscleromitrius Bizot &amp; Dury have shorter limbidia.</p><p>ETYMOLOGY. — The occurrence of small, convex, smooth cells is remarkable in a subgen. Polypodiopsis species that typically have pluripapillose, mammillose or smooth, ± plane cells. The species is named for these convex cells that produce a somewhat colliculose (collis: hill; -osus: rich in) leaf surface.</p><p>DESCRIPTION</p><p>Stem with central strand, unbranched to heavily branched, not heterocaulous, pinnately foliated, 5-11 × 1-2 mm, rhizoids brown, smooth; axillary nodules not differentiated; leaves distant to close, up to 16 pairs, crispate when dry, soft, lanceolate to elliptic-lanceolate with an acute, acuminatemucronate apex, 1.0-1.5 × 0.3-0.4 mm, 3-4.5 times as long as wide, margin subentire where elimbate; limbidia on the vaginant laminae of all leaves, extending the complete length of the vaginant lamina, frequently extending onto the apical lamina, confluent at the apex of the vaginant lamina or not, up to 16 µm wide, marginal, unistratose; vaginant lamina 2/3- 3/4 the leaf length, subequal, near the insertion about as wide as the stem, unistratose; dorsal lamina variable below, often rounded, plane, less often undulate, reaching the insertion, rarely decurrent; dorsal and apical lamina unistratose, unito bistratose near the costa; costa excurrent, bryoides-type; lumina mid dorsal laminal cells 3.5-6.5 × 2.5-5.5 µm, walls thin, up to 1 µm diam., convex, smooth; lumina mid vaginant laminal cells 4.5-7.0 × 2.5-5.5 µm, walls up to 1 µm diam., convex, smooth. Gemmae not seen.</p><p>Perichaetia and perigonia on the same or on different stems, perigonia terminal on stems and branches; perigonial branches bud like and short, ± 0.5 mm long (stalk included) (Fig. 1D) to branch like and to up to 0.9 mm long, antheridia 170-200 µm long; perichaetia terminal, perichaetial leaves 1.3-1.9 mm long, archegonia 250 µm long; calyptra smooth, 0.6 mm long. Sporophyte, seta 4 mm long, smooth, 1 per perichaetium; theca erect, cylindric, 0.8-1.0 × 0.3 mm, ± 32 columns of oblong exothecial cells; peristome-type unknown, tooth base 48.5 µm wide; operculum rostrate, 0.4 mm long; spores not observed.</p><p>The semilimbate leaves and the low number of columns of exothecial cells indicate that the new species belongs to subgenus Polypodiopsis (Müll.Hal.) Broth. This subgenus is characterized by c. 32 columns of exothecial cells and probably scariosus-type peristomes. However, since the peristome teeth of F. colliculosus Brugg.-Nann., sp. nov., are broken the type cannot be determined with certainty.</p></div>	https://treatment.plazi.org/id/03A3AD0FFF82FFE3A0A0FD7CFB72FCDA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bruggeman-Nannenga, Maria Alida	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF82FFE3A331FC1DFAEBF85F.text	03A3AD0FFF82FFE3A331FC1DFAEBF85F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Fissidens pellucidus var. ripensis (Bruggeman-Nannenga 2025) Brugg. - Nann. 2025	<div><p>Fissidens pellucidus var. ripensis (Mitt.) Brugg.-Nann., comb. nov., stat. nov.</p><p>(Fig. 2)</p><p>Fissidens ripensis Mitt., Transactions of the Linnean Society of London 23: 55, 6, fig. 15 (Mitten 1860).</p><p>— Lectotype (designated by Slavick 1992 as holotype): Niger Expedition; Barter 1425 (NY, mixed with e.g. F. porrectus Mitt.; isolectotypes: PC, BM).</p><p>Fissidens cryptarum Müll.Hal., Linnaea 39: 352 (Müller 1875).</p><p>— Lectotype (designated by Bruggeman-Nannenga 1993, here refined with barcode): [Sudan], Central-Africa • im Lande der Bongo bei Kulongo der Höhle von Gubbihih; Schweinfurth 2234 MNHN [PC10106795]; isolectotypes: PC and H-BR as Schweinfurth s.n., syn. nov.</p><p>Fissidens cremersii Bizot &amp; Onr., Revue bryologique et lichénologique 42: 846 (Bizot 1974).</p><p>— Holotype: Madagascar • au nord de Maromandia; au bord d’un marigot permanent, sous forêt, sur un arbre; 50 m alt.; Cremers 2734; MNHN [PC0129997]; isotype: BR, syn. nov.</p><p>DIAGNOSIS. — Fissidens pellucidus var. ripensis differs from F. p. var. pellucidus by mucronate apices and taller plants with up to 55 leaf pairs. Leaves of the variety can be elimbate, but typically have weak limbidia on the basal parts of some or all or most vaginant laminae. Leaves of var. pellucidus are as a rule elimbate. Most collections of var. ripensis are from semiaquatic places, but it is not restricted to these.</p><p>OTHER EXAMINED MATERIAL (Specimens used in Fig. 2). — Madagascar • Atsinanana (Toamasina); Secondary forest, on rock; 17°42’1”S, 49°27’18”E; Brinda et Andramiarisoa 12757 (MO, private herbarium Bruggeman-Nannenga) • Tamatave, Forest Reserve atTampolo, 95 km N of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=49.4&amp;materialsCitation.latitude=-17.1" title="Search Plazi for locations around (long 49.4/lat -17.1)">Tamatave</a>; on root in boggy area; 17°6’S, 49°24’E; Crosby 6700 (PC, private herbarium Bruggeman-Nannenga) • SW Madagascar, Isalo National park, ‘ Piscine Naturelle’ 5 km W of Ranohira; sandstone canyon with Pandanus near the streamlet; Orban 9455 / CR; EGR; type specimen of F. cremersii MNHN [PC0129997] .</p></div>	https://treatment.plazi.org/id/03A3AD0FFF82FFE3A331FC1DFAEBF85F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bruggeman-Nannenga, Maria Alida	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF80FFE1A3A9FC5DFBB5FAD6.text	03A3AD0FFF80FFE1A3A9FC5DFBB5FAD6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Fissidens aciphyllus Dixon	<div><p>Fissidens aciphyllus Dixon</p><p>Fissidens aciphyllus Dixon, Transactions of the Royal Society of South Africa 18: 250, pl. 3, f. 2 (Dixon &amp; Wager 1929).</p><p>— Lectotype (designated as holotype byMagill 1981, here refined with barcode): South Africa • Port St John’s. 1920; Wager 927; BM [BM000871817]; isolectotype: PRE.</p><p>Fissidens magnicellulatus Brugg.-Nann., Lindbergia 36: 13, fig. 2. (Bruggeman-Nannenga 2013).</p><p>— Holotype: Zambia • Lusaka Distr.; on damp vertical earth bank in Syzygium thicket by Bothasrus road 8 km from the Wilfred Watson turn-off from the Lusaka/ Mumbwa road; c. 1275 m a.s.l.; 10.II.1975; Townsend 75 / 21; L; isolectotype: E, syn. nov.</p></div>	https://treatment.plazi.org/id/03A3AD0FFF80FFE1A3A9FC5DFBB5FAD6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bruggeman-Nannenga, Maria Alida	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF80FFE1A14CFF7EFADFFCDA.text	03A3AD0FFF80FFE1A14CFF7EFADFFCDA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Fissidens sciophyllus subsp. capeverdensis Brugg. - Nann. 2025	<div><p>Fissidens sciophyllus Mitt. subsp. capeverdensis Brugg.-Nann., subsp. nov.</p><p>(Fig. 3)</p><p>HOLOTYPE. — Kapverden • Santo Antão, ribeira das Pombas; oberer Bereich, auf feuchten Steinen im obersten Bereich der Pflanzungen, nordostexponiert; 260 m alt.; 10.II.1995; Lindlar 405; L.</p><p>DIAGNOSIS. — This subspecies is characterized by the pale green color of young plants, the mostly distant, lanceolate leaves; the characteristic (but variable) apex, typically narrow and terminated by a large, pointed cell; rounded dorsal lamina bases, limbidia restricted to the leaves of vaginant laminae of perichaetial stems; laminal cells with 2-4 punctate papillae over the lumen; slightly to half-open vaginant laminae and costae that end 2-10 cells below the apex. The antheridia are terminal on short stems and branches, and the leaves are inflexed with a slightly incurved apex when dry. The new variety differs from subsp. sciophyllus in the typically slender, tapering leaf tips and shorter costae that do not reach the apex and may end as many as 10 cells below it. Leaf tips of var. sciophyllus are mostly less narrowly acute with percurrent to shortly excurrent costae. At first sight F. s. subsp. capeverdensis may resemble the smooth-celled F. pellucidus . Both are pale green when young and have distant, lanceolate leaves. However, the pluripapillose laminal cells of the new subspecies make distinction easy.</p><p>ETYMOLOGY. — This subspecies is endemic to the Cape Verde Islands. This is reflected in the name.</p><p>DISTRIBUTION. — Endemic to the Cape Verde Islands where it known from Santo Antão, Brava, Fogo, Santiago and São Nicolau.</p><p>SUBSTRATE. — Stones, less often soil.</p><p>ELEVATION. — 260-1220 m alt.</p><p>PARATYPES. — Kapverden • Brava, an der Strasse zum Flughafen; c. 600 m alt.; hinter Cova Rodela. Felsrinne, nordexponiert; 580 m alt.; 30.I.1995; Lindlar 91 (private herbarium Bruggeman-Nannenga) • Santiago, Serra de Malagueta; Ost- bzw. Nordosthänge im Südwesten von Cha de Figueiras, auf Erde und Gestein; ost- nordostexponiert; 770-800 m alt.; 1.II.1995; Lindlar 107 (private herbarium Bruggeman-Nannenga) • Santo Antão, ribeira das Pombas; oberer Bereich, auf feuchten Steinen im obersten Bereich der Pflanzungen; nordostexponiert; 260 m alt.; 10.II.1995; Lindlar 404 and 410 (all private herbarium Bruggeman-Nannenga).</p><p>REMARKS</p><p>Leaf tips vary from tips that, apart from the shorter costa, resemble those of subsp. sciophyllus (Fig. 3F) to long, slender and attenuated (Fig. 3G). Some of this variability is shown in Fig. 3 F-I. Earlier reported from the Cape Verde islands as F. sciophyllus (Frahm et al. 1996) and F. danckelmannii Müll. Hal. (Bizot 1969; Potier de la Varde 1943).</p><p>DESCRIPTION BASED ON LINDLAR 405 (HOLOTYPE), LINDLAR 404 (CROSS-SECTIONS) AND LINDLAR 107 (PERIGONIA) Growing in loose mat. Stem with central strand, not heterocaulous, unbranched, 4-5 × 1.5 mm, pinnately foliated; rhizoids brown, smooth; axillary nodules not differentiated; leaves pale green, distant to close, up to 13 pairs, reflexed when dry, lanceolate with a long, narrow, acute-acuminate apex ending in a large pointed cell, 0.7-1.1× 0.2-0.3 mm, 3.6-4.7 times as long as wide, margin subentire to crenulate, elimbate except for the vaginant laminae of the 4 upper leaf pairs of perichaetial stems that have a unistratose (estimated), more or less marginal limbidium; limbidium extending up to 1/2 the vaginant lamina in perichaetial leaves; vaginant lamina 1/2-ª⁄5 the leaf length, rounded and less wide than the stem at the insertion, unistratose, unequal with the minor lamella ending ± midway between costa and margin; dorsal lamina rounded and slightly undulate at the insertion, reaching the insertion, not decurrent; dorsal and apical lamina unistratose; costa ending 2-7 (up to 10 in some collections) cells below the apex, bryoides-type; mid dorsal laminal cells (4.5-)6-8 ×(3-) 4.0-7 µm, with 2-4 tiny punctate papillae; mid vaginant laminal cells 7-9.5(-11) × (4.5-) 5.5-8 µm, plane, with 3-4 punctate papillae over the lumen. Gemmae not observed.</p><p>Perigonia terminal on 1-1.5 mm long stems and on branches; antheridia c. 150 µm long; perichaetia terminal, perichaetial leaves c. 1.5 mm long, archegonia c. 300 µm long; calyptra not observed. Sporophyte, seta 4 mm long, smooth, theca erect, ± cylindric, 0.6× 0.3 mm, ± 32 columns of collenchymatous exothecial cells; peristome type unknown, tooth base c. 32 µm wide; operculum not observed; spores not observed.</p></div>	https://treatment.plazi.org/id/03A3AD0FFF80FFE1A14CFF7EFADFFCDA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bruggeman-Nannenga, Maria Alida	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF80FFEFA3DBFA1BFBAFFCFA.text	03A3AD0FFF80FFEFA3DBFA1BFBAFFCFA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Fissidens ceylonensis Dozy & Molk.	<div><p>Fissidens ceylonensis Dozy &amp; Molk.</p><p>(Fig. 4)</p><p>Muscorum Frondosorum Novae Species ex Archipelago Indico et Japonica: 7 (Dozy &amp; Molkenboer 1844) (Aug.-Sept. 1844).</p><p>— Lectotype (designated byTouw 2013): Ceylon [Sri Lanka] • König 910, 114-864 = L0041421 (herb. Van Royen; labelled by Sande Lacoste; analytic drawings by Iwatsuki).</p><p>Fissidens borgenii Hampe, Botanische Zeitung (Berlin) 28: 36 (Hampe 1870).</p><p>— Lectotype (designated as holotype byMagill 1981, here refined as “specimen in herbarium Hampe”): South Africa • Natal, prope Umpumulo; Borgen 7; BM-herb. Hampe; isolectotypes: BM, PC, H-SOL, syn. nov.</p><p>Fissidens sarcophyllus Müll.Hal. ex Dusén, Kongliga Svenska Vetenskaps Academiens Handlingar, n.s. 28 (2): 11, fig. a-d; pl. 2, f. 5. (Dusén 1895).</p><p>— Lectotype: (designated by Bruggeman-Nannenga 1993, here refined with barcode): Musci Africani in Camerunia a P. Dusén collecti coll. a13, BM [BM000879000]; isolectotype: BM, syn. nov.</p><p>Fissidens intralimbatulus Broth. &amp; P. de la Varde, Revue bryologique n.s. 1: 87, pl. II, f.1. (Potier de la Varde 1928).</p><p>— Lectotype (designated by Bruggeman-Nannenga 1993 here refined with barcode): Gabon • Dibwangui; Le Testu 5631; MNHN [PC0106495], syn. nov.</p><p>Fissidens paucilimbatus P. de la Varde, Revue bryologique n.s. 1: 88. pl. II, 2. (Potier de la Varde 1928).</p><p>— Lectotype (Bruggeman-Nannenga 1993 here refined with barcode): Gabon • dans la Basomba; Le Testu 5633; MNHN [PC0106554]; isolectotype: PC, syn. nov.</p><p>Fissidens aoristoloma P. de la Varde, Revue bryologique et lichénologique 11: 179, f. III,1. (Potier de la Varde 1939).</p><p>— Lectotype (designated by Bruggeman-Nannenga 1993 here refined with barcode): Central African Republic • Mbaïki; Tisserant M738; MNHN [PC0029462]; isolectotype: PC, syn. nov.</p><p>Fissidens intralimbatulus var. brevifolius P. de la Varde, Revue bryologique et lichénologique 18: 109 (Potier de la Varde 1949).</p><p>— Lectotype (designated by Bruggeman-Nannenga 1997 here refined with barcode): Burkina Faso • Haute Volta; chutes de la Comoe près de Banfora; Chevalier s.n.; PV9935; MNHN [PC0095359]; isolectotype: MNHN [PC0095358], syn. nov.</p><p>Fissidens norkettii P. de la Varde, Revue bryologique et lichénologique 22: 7, fig. 2. (Potier de la Varde 1953).</p><p>— Lectotype (designated by Bruggeman-Nannenga 1993 here refined with barcode): Nigeria • Jos; Thorold 306; MNHN [PC0095365]; isolectotype: MNHN [PC0073009], syn. nov.</p><p>Fissidens subceylonensis Dury &amp; Onr., Revue bryologique et lichénologique 42: 854. (Bizot 1976).</p><p>— Lectotype (designated as holotype by Bruggeman-Nannenga 1999, here refined with barcode): Seychelles • Mahé, Evéché de Victoria, la Rosière; sur les murs de granit ombragés; 20 m alt.; 20.I.1974; Onraedt 75-S-003; MNHN [PC0096369].</p><p>DIAGNOSIS (distinctive features bold). — Fissidens ceylonensis is a variable, palaeotropical species characterized by stiff, opaque leaves, marginal papillae that obscure the cells, long vaginant laminae reaching 2/3-ª⁄5 the leaf length and irregular, typically intramarginal, limbidia. The distribution and extension of the limbidia is variable. They vary between restricted to the upper leaves of perichaetial stems (e.g. in the type specimen of F. sarcophyllus) and present on the vaginant laminae of all or most leaves of both vegetative and perichaetial stems. Limbidia can exist of a single cell or extend over a large part of the vaginant lamina, but always end below the apex of the vaginant lamina. Limbidia are frequently interrupted, irregular and typically at least in part intramarginal, limbidia of perichaetial leaves intramarginal or marginal. Limbidia are unistratose or, rarely, partly bistratose. The shape of the leaves varies between lanceolate and oblong-lanceolate. The costa typically is percurrent to excurrent in short mucro. Rarely it ends a few cells below the apex (e.g. in the type specimen of F. borgenii). The African F. schweinfurthii resembles F. ceylonensis in the obscure, stiff leaves, long vaginant laminae, marginal papillae, lanceolate to oblong leaves and percurrent to excurrent costae. This species is easily separated by its better developed, limbidia that are present on the vaginant laminae of all leaves, reach the apex of the vaginant laminae and are marginal throughout, except sometimes for a short intramarginal piece near the insertion.</p><p>OTHER EXAMINED MATERIAL (Specimens used in Fig. 4). — Democratic Republic of the Congo • Bas-Uele; 5 km à l’ouest de Likali, termitière dans la forêt secondaire; Lisowski 47587; KRAM (private herbarium Bruggeman-Nannenga).</p><p>Ghana • Central Region, Akroform-Akwekrom, Nimere R.; riverside rocks under roadbridge; 5°27’760N, 1°30’236W; R. Agyei, G. Ameka, C. Dankwah &amp; N.G. Hodgetts 10217 (private herbarium Hodgetts; small collection mixed with other species) .</p><p>Guinea • Nzérékoré Region, Macenta Prefecture, above the village of Zoubrouma; rocks in the forest; Lisowski 129 and 225; KRAM (private herbarium Bruggeman-Nannenga).</p><p>Seychelles • West coast of southern Mahé island; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=55.47639&amp;materialsCitation.latitude=-4.6833334" title="Search Plazi for locations around (long 55.47639/lat -4.6833334)">Point Maravi</a> at the S end of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=55.47639&amp;materialsCitation.latitude=-4.6833334" title="Search Plazi for locations around (long 55.47639/lat -4.6833334)">Anse Gaulettes</a> beach; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=55.47639&amp;materialsCitation.latitude=-4.6833334" title="Search Plazi for locations around (long 55.47639/lat -4.6833334)">Carophyllum</a>, coconut and seminatural thicket on granitic rocks, rupicolous; 4°41’S, 55°28’35”E; Kis 9329 /X; EGR, L • Northern tip of Mahé Island, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=55.443058&amp;materialsCitation.latitude=-4.577778" title="Search Plazi for locations around (long 55.443058/lat -4.577778)">Montagne Glacis</a> above <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=55.443058&amp;materialsCitation.latitude=-4.577778" title="Search Plazi for locations around (long 55.443058/lat -4.577778)">Rivière Glacis</a> valley; plantation forest and thickets, rupicolous; 4°34’30”S, 55°26’25”E; 4°34’40”S, 55°26’35”E; Kis 9340 / BM; EGR, L .</p><p>DISTRIBUTION. — This is the first record for mainland Africa of this species under the name F. ceylonensis . Formerly, African specimens were known as e.g. F. borgenii . Further distribution: Seychelles, Asia, Polynesia and Australia.</p><p>REMARK</p><p>All new synonyms above were earlier (Bruggeman-Nannenga 1993, 1997) reduced to F. borgenii and thus are no more than the logical result of the synonymy of F. borgenii with F. ceylonensis . This list of synonyms is not complete as only African synonyms are included.</p></div>	https://treatment.plazi.org/id/03A3AD0FFF80FFEFA3DBFA1BFBAFFCFA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bruggeman-Nannenga, Maria Alida	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF8EFFEFA387FC3DFB5CF9D9.text	03A3AD0FFF8EFFEFA387FC3DFB5CF9D9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Fissidens cuynetii Bizot 1973	<div><p>Fissidens cuynetii Bizot</p><p>Fissidens cuynetii Bizot, Acta Botanica Academiae Scientiarum Hungaricae 18: 21, f. 6: 1-3 (Bizot 1973).</p><p>— Lectotype (designated here): Uganda • Ankole district, Moutozho; sur rochers ensolleillées; 1300 m alt.; XII.1967; Balázs 96 G; MNHN [PC0073300].</p><p>REMARKS</p><p>The lectotypification of this species is rather complicated. On page 11 of Bizot (1973) is a footnote that says that holotypes of new species described in the paper are in DI (where Bizot used to work; after his death in 1979 his herbarium went to PC) and isotypes in EGR. Thus Pócs’ 1977 ‘typification’ of F. cuynetii that says holotype in EGR, isotype in PC, is not effective because it is in contradiction with the protologue. A further complication is that the protologue gives the collection number Balázs 96, whereas in Bizot’s herbarium in PC there are Balázs 96 D MNHN[PC0073299] and Balázs 96 G MNHN[PC0073300]. Balázs 96 G MNHN[PC0073300] is here designated lectotype because it has a note in Bizot’s handwriting saying “sp. nov.” and all information on its label is by Pócs (who donated the material to Bizot for identification), whereas the information on 96 D is just a handwritten copy of MNHN[PC0073300] by Bizot.</p></div>	https://treatment.plazi.org/id/03A3AD0FFF8EFFEFA387FC3DFB5CF9D9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bruggeman-Nannenga, Maria Alida	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF8EFFEDA3A1F918FD05FDE5.text	03A3AD0FFF8EFFEDA3A1F918FD05FDE5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Fissidens hymenodon Besch.	<div><p>Fissidens hymenodon Besch.</p><p>Annales des Sciences Naturelles, Botanique, série 7, 2: 85. (Bescherelle 1885).</p><p>— Lectotype (designated here): Mayotte • Marie 206 (partly unreadable): BM [BM000871897]; isolectotypes: MNHN [PC0129799].</p><p>Fissidens alomoides Müll.Hal. ex Dusén, Kongliga Svenska Vetenskaps Academiens Handlingar 28(2): 10, 11: f. a-b, pl. II, f. 3. (Dusén 1895).</p><p>— Lectotype (designated here): Cameroon • Dusén; Musci Afr. Camerunia nr. 505; S [B107445]; isolectotypes: H-BR, MNHN [PC0029441] and S [B107447, B107448, B107449, B107450], syn. nov. Isotypes have been distributed as: Musci Africani in Camerunia a P. Dusén collecti 505 (also miswritten as 305).</p><p>DESCRIPTION</p><p>Fissidens hymenodon is a corticolous species that is characterized by oblong to oblong-lanceolate leaves with broadly acute to obtuse, often rounded apices, apex not terminated by a pointed cell, costae ending 2-6 cells below the apex, limbidia restricted to the upper leaves of perichaetial leaves, 1-2 papillose laminal cells and scariosus-type peristomes.</p></div>	https://treatment.plazi.org/id/03A3AD0FFF8EFFEDA3A1F918FD05FDE5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bruggeman-Nannenga, Maria Alida	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF8CFFEDA16BFA5BFB55FB59.text	03A3AD0FFF8CFFEDA16BFA5BFB55FB59.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Fissidens ferrugineus Mull. Hal.	<div><p>Fissidens ferrugineus Müll.Hal.</p><p>(Fig. 5)</p><p>Botanische Zeitung (Berlin) 22: 341 (Müller 1864).</p><p>— Lectotype (designated here): Madagascar • Pervillé 51A; BM [BM000871895] mixed with the lectotype of F. madecassus Schimp. ex Müll.Hal.; isolectotype: BM [BM000871896]; two stems mixed between six stems of the isolectotype of F. madecassus .</p><p>Fissidens ugandae Dixon &amp; P. de la Varde in P. de la Varde, Annales de Cryptogamie Exotique 3: 45, 1, f. 3. (P. de la Varde 1930).</p><p>— Lectotype (designated here): Uganda • Musandama; Budd s.n.; 1926; BM [BM000871873]; isolectotype: MNHN [PC0095552], syn. nov.</p><p>Fissidens gibbonii Bizot, Revue bryologique et lichénologique 40 (2): 131 (Bizot 1973).</p><p>— Lectotype (designated byPócs 1977 as holotype): Tanzania • Morogoro District, Uluguru Mts, on top of Bondwa Peak; on the ground of subalpine heath; 2100 m alt.; 12.X.1969; Pócs &amp; Gibbon 6052 /CR (EGR; isolectotypes: MNHN [PC0132125] and DSM), syn. nov.</p><p>DIAGNOSIS (distinctive features in bold). — Plants often rusty brown, leaves narrow, 3.5-5 (-6) times as long as wide, lanceolate with narrow acute to acute-acuminate tips; limbidium thick yet unistratose (Fig. 5J), smooth to denticulate, marginal, present on the vaginant laminae of all leaves, typically extending the whole length of the vaginant lamina, often confluent at the apex and frequently extending onto the apical lamina; vaginant lamina 2/3-3/4 the leaf length; costa percurrent to excurrent, or ending 1-2 cells below the apex; laminal cells can have 1-2 or 2-4 papillae per cell (rarely 1(-2), or 2-6), punctate, less often irregular (Fig. 5I) papillae; antheridia terminal on short stems (rhizautoicous), less often terminal on longer stems or on short perigonial branches in the axils of lower leaves; setae 3-10 mm long, theca cylindric to strumulose, with 32-58 columns of exothecial cells. Some collections have thecae with ± 32 as well as with 40 or more exothecial cells. Since ±32 columns are typical for subg. Polypodiopsis 32 is not conclusive. When, however, there are 40 or more exothecial cells, this is diagnostic for F. ferrugineus .</p><p>OTHER EXAMINED MATERIAL (Specimens used for Fig. 5). — Madagascar • Pervillé 51A; BM [BM000871895], lectotype F. ferrugineus . Tanzania • Meru Mts; in the crater ground near Njeku; swampy moorland patches, rupicolous; 2600 m alt.; Pócs et al. 8670 /D; EGR, L • Kilimanjaro Mts, Karanga River Gorge N of Kibosho; Montane evergreen forest on the steep; rocky E facing slope; with Albizia gummifera (J.F.Gmel.) C.A.Sm.; on soil covered rock; 1880 m alt.; Pócs, Mialla &amp; Margesi 90068 / G; EGR • Southern Highlands, Kipengere Range, above Salala Falls below Mt Ishinga; in deep ravine of streamlet, on half shady rock; 2380-2400 m alt.; 26.V.1987; Pócs, Iversen, Persson &amp; Petterson 87146 / O; EGR, private herbarium Bruggeman-Nannenga • Ukaguru Mts in Kilos District, on Mamiwa ridge; mossy elfin forest, rupicolous; 2150 m alt.; Pócs, Chamshama &amp; Sylvander 86100 / T; EGR, L • Ukaguru Mts, NE slope of Mamwira W of Masenge village; on shady rocks in bamboo ( Arundinaria alpina K.Schum.), thicket; 2050 m alt.; Pócs 88015 / C; EGR .</p><p>La Réunion • G. de L’ Isle s.n. (PC) • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=55.54&amp;materialsCitation.latitude=-21.666666" title="Search Plazi for locations around (long 55.54/lat -21.666666)">Forêt de Bélouve</a> on the Plateau des Cochons, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=55.54&amp;materialsCitation.latitude=-21.666666" title="Search Plazi for locations around (long 55.54/lat -21.666666)">Trou de Fer</a> trail; 21°40’S, 55°32.3’E; 21°40’S, 55°32.4’E; Bruggeman-Nannenga 10995; L .</p></div>	https://treatment.plazi.org/id/03A3AD0FFF8CFFEDA16BFA5BFB55FB59	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bruggeman-Nannenga, Maria Alida	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF8CFFEDA0F2FD3FFE2AFA18.text	03A3AD0FFF8CFFEDA0F2FD3FFE2AFA18.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Fissidens intramarginatus	<div><p>FISSIDENS INTRAMARGINATUS COMPLEX: REINSTATEMENT OF F. FERRUGINEUS AND F. SCHWEINFURTHII</p><p>Fissidens ferrugineus and F. schweinfurthii were considered synonyms of F.intramarginatus (Bruggeman-Nannenga 1993, 1997). Fissidens intramarginatus s.l. is characterized by pluripapillose laminal cells and vaginant laminae that are limbate in all leaves of both perichaetial and vegetative stems. The limbidia nearly or completely reach the apex of the vaginant laminae and frequently extend onto the apical lamina. It comprises several expressions that are linked by transitional forms (species-complex).</p><p>Meanwhile, it has become clear that the neotropical concept of F. intramarginatus (Pursell 2007) is narrower than the African concept (Bruggeman-Nannenga 1993, 1997) and that a revision is needed. In anticipation of a revision of the African species of this complex (in preparation). F. ferrugineus and F. schweinfurthii are here reinstated as species. As a rule, these two species are easily distinguished by the papillae. Fissidens schweinfurthii has 5-8 marginal (i.e. distributed on or near the cell walls in surface view) papillae that obscure the cells, whereas F. ferrugineus has a variable, lower number (1-2 or 2-4 per cell) of punctate to irregular papillae over the lumen and more or less clear cells.</p></div>	https://treatment.plazi.org/id/03A3AD0FFF8CFFEDA0F2FD3FFE2AFA18	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bruggeman-Nannenga, Maria Alida	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF8CFFE9A3FBFA9AFEB2FA6F.text	03A3AD0FFF8CFFE9A3FBFA9AFEB2FA6F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Fissidens schweinfurthii (Mull. Hal.) Paris	<div><p>Fissidens schweinfurthii (Müll.Hal.) Paris</p><p>(Fig. 6)</p><p>Index bryologicus: 484 (Paris 1896). — Conomitrium schweinfurthii Müll.Hal., Linnaea 39: 369. (Müller 1875).</p><p>— Lectotype (designated by Bruggeman-Nannenga 1997, here refined with barcode): Sudan as Niam-niam-regiones [border area Sudan, Democratic Republic of the Congo and Central African Republic] • in cacumine collis Gumango; 6.II.1870; Schweinfurth s.n.; MNHN [PC0095370]; isolectotype: MNHN [PC0129867].</p><p>Fissidens nossianus Besch., Annales des Sciences naturelles, Botanique, série 6, 9: 333 (Bescherelle 1880).</p><p>— Lectotype (designated by Bruggeman-Nannenga 1997 here refined with barcode): Madagascar • Nossi Bé, Marie s.n.; BM [BM001094101]; isolectotypes: H-BR, PC, syn. nov.</p><p>Fissidens motelayi Renauld &amp; Cardot, Prodrome de la Flore Bryologique de Madagascar, des Mascareignes et des Comores: 117 (Renauld 1898).</p><p>— Lectotype (designated here): Madagascar • Andriba; 1897; Dorr s.n.; MNHN [PC0098324]; isolectotype: MNHN [PC0098325], syn. nov.</p><p>Fissidens ferrugineus Müll.Hal. var. pallidus Thér., Recueil des publications de la société havraise d’études diverses 90: 81 (Thériot 1923); F. decaryi Bizot, Revue bryologique et lichénologique 40: 131 (Bizot 1974) (new name and rank).</p><p>— Lectotype (designated by Bruggeman-Nannenga 1997, here refined with barcode): Madagascar • Ambohimanga; Décary 9; MNHN [PC0096226], syn. nov.</p><p>DIAGNOSIS (distinctive features in bold). — This African species is characterized by stiff plants, leaves close, in transmitted light obscure with pale, per- to short excurrent costa, vaginant laminae long, reaching 2/3-̚⁄10 the leaf length, proximally about as wide as the stem, with rounded insertion, limbidia reaching the insertion or ending some distance above, marginal throughout (Fig. 3 G, H) or intramarginal towards the insertion (Fig. 3F), distally nearly or completely reaching the apex of the vaginant lamina; laminal cells with (4-)5-8(-10) well developed, low, blunt, marginal papillae (Fig. 6I, J); archegonia typically terminal on short or long stems and/ or on short branches; antheridia terminal on dwarf or larger stems, occasionally on short bud-shaped branches. Most specimens of this species are easily recognized by the bold printed traits.</p><p>Fissidens ceylonensis similarly has obscure leaves, short excurrent to percurrent costae and marginal papillae, but differs in the shorter limbidia that do not reach the apex of the vaginant laminae, and are frequently lacking on some or most leaves.</p><p>OTHER EXAMINED MATERIAL (specimens used forFig. 6). — Democratic Republic of the Congo • Haute-Shaba; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-6.483333&amp;materialsCitation.latitude=9.533334" title="Search Plazi for locations around (long -6.483333/lat 9.533334)">Plateau des Kundelungu</a>, partie supérieure de la Lofoi; Lisowski 4225; KRAM • Haute-Shaba, près de Likasi, rive gauche de la rivière Panda; gallerie forestière; Lisowski 65116; KRAM.</p><p>Ivory Coast • Boundiali; nr 1 B3; 9°32’N, 6°29’W; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-6.483333&amp;materialsCitation.latitude=9.533334" title="Search Plazi for locations around (long -6.483333/lat 9.533334)">Granitinselberg</a>, auf beschattenen <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-6.483333&amp;materialsCitation.latitude=9.533334" title="Search Plazi for locations around (long -6.483333/lat 9.533334)">Fels</a>; Porembski &amp; Biedinger 1508; B, private herbarium Bruggeman-Nannenga • Westl. von Bouaké. Granitinselberg (nr IB24), terrestrisch; in Afrotrilepis Matte; 370 m alt.; Porembski 515; private herbarium Bruggeman-Nannenga .</p><p>Madagascar • Prov. Majunga; Station forestière d’Ampijoroa, c. 3 km N <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=46.85&amp;materialsCitation.latitude=-16.333334" title="Search Plazi for locations around (long 46.85/lat -16.333334)">d’Andranofasika</a>; on soil, base of tree in forest; 16°20’S, 46°51’E; Dorr 3055; MO, L • 60 km NW of Antanannarivo along Mahajanga road; N of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=47.28389&amp;materialsCitation.latitude=-18.498333" title="Search Plazi for locations around (long 47.28389/lat -18.498333)">Soaniadanana village</a>; granite outcrops in secondary plateau grassland, rupicolous-terricolous; 18°29’54”S, 47°17’02”E; 5.IX.1994; Kis 9443EE; L, EGR .</p><p>Malawi • Southern District, Zomba Plateau; Long 12529; E; private herbarium Bruggeman-Nannenga .</p><p>Nigeria • Ogun State, Ijebe-Ode; savanne area S of Idowa, bank in partial shade; McFarlane 275; private herbarium Bruggeman-Nannenga, a few mixed between F. pellucidus Hornsch .</p><p>Sudan • Schweinfurth s.n.; type specimen F. schweinfurthii MNHN [PC0095370] .</p><p>Togo • circ. De Klouto; an der Strasse Palimé-Klouto, etwa auf der halben Höhe des Gebirgsrandes, oberhalb des Wasserfalls; erdig-felsige Böschung; 300-400 m alt.; 8.X.1973; Hiepko &amp; Schultze-Motel 393; NY, MO, L .</p></div>	https://treatment.plazi.org/id/03A3AD0FFF8CFFE9A3FBFA9AFEB2FA6F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bruggeman-Nannenga, Maria Alida	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF88FFF7A134F9BBFEBDF859.text	03A3AD0FFF88FFF7A134F9BBFEBDF859.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Fissidens porrectus Mitt.	<div><p>Fissidens porrectus Mitt.</p><p>(Fig. 7)</p><p>Transactions of the Linnean Society of London 23: 56, f. 17 (Mitten 1860).</p><p>— Lectotype (designated by Slavick 1992 as holotype): Niger -exp. • Barter 1425; NY, isolectotype: PC, BM, H-BR, S.</p><p>Fissidens nitens Rehmann ex E.S.Salmon, Annals of Botany 13: 121, figs 69-74 (Salmon 1899).</p><p>— Lectotype (designated byMagill 1981, here refined with barcode): South African Republic • Rehmann, Musci austro-africani (1875-77): 289; BM [BM000871824]; isolectotypes: have been distributed as Dr. A. Rehmann, Musci austro-africani (1875-77): 289; BM [BM000871823], PC, S.</p><p>Fissidens nitens var. neglectus E.S.Salmon, Annals of Botany 13: 122, figs 75-76 (Salmon 1899).</p><p>— Lectotype (designated by Bruggeman-Nannenga 1993, here refined with barcode): Tanzania • Lutindi; am Boden im Hochwald; Holst 3472; p.p. BM [BM000871825].</p><p>Fissidens mariei (Besch.) Broth., Die Natürlichen Pflanzenfamilien I (3): 358 (Brotherus 1901); Conomitrium mariei Besch., Annales des Sciences Naturelles, Botanique, série 6, 9: 335 (Bescherelle 1880).</p><p>— Lectotype (designated here): Madagascar • Mosses of Nossi Bé; on earth; Marie s.n.; BM [BM001094100]; isolectotypes: PC, H-BR.</p><p>Fissidens rufolimbatus Cardot, Revue bryologique 35: 65 (Cardot 1908).</p><p>— Lectotype (designated here): Democratic Republic of the Congo [Congo belge] • Sankuru; leg. Em et M. Laurent; 1903; MNHN [PC0106533]; isolectotypes: MNHN [PC0106532], HBR.</p><p>Fissidens letestui P. de la Varde, Bulletin de la Société Botanique de France 74: 144, f. 41 (Potier de la Varde 1927).</p><p>— Lectotype (designated here): Gabon • Evoungoun; 9.VII.1925; Le Testu 5374; PV2221; MNHN [PC0106481]; isolectotypes: MNHN [PC0106478, PC0106480] and H-BR .</p><p>Fissidens pachylomoides P. de la Varde, Revue bryologique 4 n. s.: 57, f. 1: 2a (Potier de la Varde 1931).</p><p>— Lectotype (designated here): Gabon • Idembé; Le Testu 6840; MNHN [PC0106558]; isolectotypes: in PC, BR, BM.</p><p>Fissidens rufolimbatus var. torrentium P. de la Varde, Revue bryologique, n.s. 4: 59, f. 1: 2d, f (Potier de la Varde 1931).</p><p>— Lectotype: (designated here): Gabon • Chute de la Moungoungoulou; Le Testu 6866; the number in the protologue (6668) must be an orthographic error and is here corrected (ICNB art. 9.2): MNHN [PC0106519]; PV3974; isolectotype: MNHN [PC0129748].</p><p>Fissidens pachylomoides var. subdenticulatus P. de la Varde, Revue bryologique et lichénologique 5: 202, f. 1 (Potier de la Varde 1932).</p><p>— Lectotype (designated here): Gabon • l’Ofoué; Eckendorff s.n.; PV4450; MNHN [PC0106556]; isolectotypes: PC, NY.</p><p>Fissidens rufolimbatus var. latifolius P. de la Varde, Revue bryologique et lichénologique 8: 52 (Potier de la Varde 1935).</p><p>— Lectotype (designated by Bruggeman-Nannenga 1993 here refined with barcode): Gabon • Mont Loumanzoc; Le Testu s.n.; PV5205; MNHN [PC0106516]; isotype: MNHN [PC0106515].</p><p>Fissidens rufolimbatus var. ramosus P. de la Varde, Revue bryologique et lichénologique 8: 52 (Potier de la Varde 1935).</p><p>— Lectotype (designated by Bruggeman-Nannenga 1993, here refined with barcode): Gabon • Rocher de Nzang, Nzoumou; Le Testu s.n.; PV5443; MNHN [PC0106518]; isolectotype: MNHN [PC0106517].</p><p>Fissidens schnellii P. de la Varde, Revue bryologique et lichénologique 18: 105, f. 1 (Potier de la Varde 1949).</p><p>— Lectotype (designated here): Guinea • Ziama; Schnell 2620; PV9954; MNHN[PC0106522]; isolectotypes: BM, MNHN[PC0106520; PC0106521].</p><p>Fissidens woodii J.Taylor &amp; P. de la Varde, Revue bryologique et lichénologique 25: 215, f. 1 (Potier de la Varde 1956).</p><p>— Lectotype (designated by Bruggeman-Nannenga 1993 here refined with barcode): Uganda • Distr. Sese; Wood 1048; PV14077; MNHN [PC0106246]; isolectotypes: MNHN [PC0106244; PC0106245].</p><p>DIAGNOSIS (most important traits in bold). — Fissidens porrectus Mitt. is variable species. It is characterized by guttulate laminal cells (Fig. 7M) with smooth, firm walls. These two traits are fairly constant. Furthermore, the dorsal and apical lamina typically have a characteristic incrassate border zone (Fig. 7B, F) while the vaginant lamina is typically limbate (a limbidium is a border of narrow elongated cells). The last two traits, the incrassate border zone and the limbate vaginant lamina, are rather variable. The incrassate border can be (uni-) 2-4-stratose, the cells forming it are usually short and wide (Fig. 7B, F), less often, elongated (Fig. 7K). The limbidia on the vaginant laminae are usually well developed and composed of elongated cells, less often they are indistinct (without or with only a few slightly elongated cells).</p><p>Plant size and development of the limbidia and of the border zone are correlated, the largest plants having the best developed limbidia and the most elongated cells in the border zone (Fig. 7). However, in several specimens this correlation is not apparent. Moreover, irregularities of the border zone are not uncommon. In spite of its confusing variability, F. porrectus is usually easily recognized by the bold printed traits. It is most likely to be confused with F. splendens Brugg.-Nann. which has similar cells and similar incrassate borders, but (contrary to F. porrectus) the border of the vaginant lamina in this species consists entirely of isodiametric cells. Fissidens pellucidus has similar cells and cell walls, but lacks a differentiated, bistratose border zone and limbidia or has its limbidia restricted to the vaginant laminae of upper leaves of perichaetial stems. Specimens with borders of elongated cells can be mistaken for subg. Fissidens species. Species of this subgenus, however, have different sporophytes and eguttulate cells.</p><p>Fissidens porrectus strongly resembles the elimbate F. pellucidus that also has guttulate cells with firm and smooth cell walls. However, Fissidens pellucidus clearly belongs to subg. Polypodiopsis . While F. pellucidus is elimbate, F. porrectus varies from more or less elimbate to limbate on all laminae. Smooth laminal cells and limbidia on all laminae are characteristic of subgenus Fissidens Hedw. In other words, gametophytes of limbate expressions of F. porrectus (Fig. 5N) correspond with subg. Fissidens, but expressions with limbidia restricted to the vaginant laminae or lacking fit better in subg. Polypodiopsis . Subgenus Fissidens is further characterized by a subg. Fissidens -sporophyte, i.e. a bryoides - type peristome and 40 or more columns of exothecial cells on the theca. The sporophyte of F. porrectus is clearly a subg. Polypodiopsis- type with ± 32 columns of exothecial cells and a scariosus type peristome. This settles the species in subg. Polypodiopis. This is supported by molecular data (Budke et al. 2023).</p><p>The incrassate border zone of the dorsal and apical lamina is one of the most conspicuous features of this species. Similar borders are widespread, but rare in the Fissidentaceae and occur in all subgenera, except subg. Fissidens . Thus the taxonomic value of this feature is limited to species level.</p><p>OTHER EXAMINED MATERIAL (Specimens used forFig. 7). — Gabon • Evoungoun; 9.VII.1925; Le Testu 5374; MNHN [PC010648]; type F. lestestui .</p><p>Madagascar • Heim s.n.; MNHN [PC0773734A]; Niger-Exp ., Barter 1425; NY, type F. porrectus .</p><p>Republic of São Tome &amp; Principe • Principe Island, cross-country route across summit plateau of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=7.3511667&amp;materialsCitation.latitude=1.5810277" title="Search Plazi for locations around (long 7.3511667/lat 1.5810277)">Mesa</a>; mixed tropical hardwood cloud forest with sedge and fern understory; on volcanic bedrock in rivulet in filtered light; 01°34’51.7”N, 07°21’04.2”E; Shevock et al. 49960 (private herbarium Bruggeman-Nannenga) .</p><p>South African Republic • Rehmann; Musci austro-africani (1875-77): 289; BM [BM000871824]; type F. nitens .</p><p>REMARKS</p><p>All these names have at different times and in different combinations been considered synonyms (Bizot &amp; Pócs 1979; Bizot et al. 1990; Bruggeman-Nannenga 1993). Thus, none of the above synonyms is truly new. The present concept embraces all previous ones.</p></div>	https://treatment.plazi.org/id/03A3AD0FFF88FFF7A134F9BBFEBDF859	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bruggeman-Nannenga, Maria Alida	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF96FFF7A3CBFF7EFBC9FCFA.text	03A3AD0FFF96FFF7A3CBFF7EFBC9FCFA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Fissidens punctulatus Sande Lac.	<div><p>Fissidens punctulatus Sande Lac.</p><p>Fissidens punctulatus Sande Lac., Species Novae vel Ineditae Muscorum Archipelagi Indici 2, fig. 1C (Sande Lacoste 1872).</p><p>— Holotype: Indonesia • (Moluccas) Saparoea; 1858-1860; De Vriese 910, 1141054 = L0058502; herb. C.M. van der Sande Lacoste; type discussed by Touw 2013.</p><p>Fissidens grainvillei P. de la Varde, Recueil des publications de la société havraise d’études diverses 99 (1): 133 (Thériot 1932).</p><p>— Lectotype (designated here): Madagascar • Kiangara; été 1931; Grainville s.n.; PV4037; MNHN [PC0096206; PC0095457], syn. nov.</p><p>REMARK ON THE TYPIFICATION</p><p>Potier de la Varde specified his collections with PV-number. In his herbarium in PC are two collections of F. grainvillei both numbered PV4037, and both annotated “sp. nov.” by Potier de la Varde. MNHN [PC 0095457] consists of a few stems plus description and MNHN [PC 0096206], a slide and drawing. They are clearly parts of the same specimen, viz. PV4037. Therefore, they are together designated the lectotype (ICBN art. 8.3).</p></div>	https://treatment.plazi.org/id/03A3AD0FFF96FFF7A3CBFF7EFBC9FCFA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bruggeman-Nannenga, Maria Alida	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
03A3AD0FFF96FFF4A3D8FCDDFC80FD26.text	03A3AD0FFF96FFF4A3D8FCDDFC80FD26.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Fissidens subundatus Dury	<div><p>Fissidens subundatus Dury in Bizot</p><p>(Fig. 8)</p><p>Revue bryologique et lichénologique 42: 854 (Bizot 1976).</p><p>— Lectotype (designated here): Madagascar • Antalaka; sur les arbres d’une forêt primaire littorale dégradée (l’écorce est couverte de terre); Onraedt 71 M 5644; MNHN [PC0096355]; isolectotype: slide by Theo Arts in BR.</p><p>REMARK</p><p>The protologue does not indicate where the type specimen is deposited. However, in her unpublished thesis (Dury 1974) the author states that the type is in Bizot’s herbarium (now in PC). It appears that in Bizot’s herbarium there is only one F. subundatus specimen with a label matching the protologue. That specimen is designated the lectotype.</p><p>DESCRIPTION (EMENDED)</p><p>Corticolous, scattered between other mosses; in KOH brown; rhizoids brown, smooth; axillary nodules not differentiated; stem delicate, pale green, unbranched, 2.5-5.0 × 2.0- 2.5 mm, only fertile stems seen; leaves close, 7-14 pairs, more or less crispate when dry, oblong-lanceolate, broadly acute with short acumen, 0.8-1.5× 0.25-0.4 mm, 3-4 (-4.5) as long as wide, margin finely serrulate where elimbate; limbidium on the vaginant laminae of all leaves, nearly or completely extending the length of the vaginant laminae, occasionally excurrent on the apical lamina, marginal, sometimes intramarginal over a short distance, 14.5 µm wide, unistratose (estimated); vaginant lamina 1/2-̚⁄10 the leaf length, slightly open, near the insertion about as wide as the stem, slightly rounded ending concave at the insertion, unistratose (estimated); dorsal lamina slightly rounded at the insertion, reaching the insertion, not decurrent, dorsal and apical lamina unistratose (estimated); costa percurrent; lumina of mid dorsal laminal cells, 3.5-7.5×3.0-5.5 µm, lowly convex with a small central papilla; lumina of mid vaginant laminal cells 5.0-8.0 (-9.0) ×3.0-6.5 µm, lowly mammillose with a distinct, tiny papilla. Gemmae not observed.</p><p>Cladautoicous, perigonia on short, short bud-shaped or longer c. 4.5 mm long branches, antheridia ± 150 µm long; perichaetia terminal, a few archegonia solitary in the axils of upper leaves of perichaetial stems, perichaetial leaves c. 1.3 mm long; archegonia 250 µm long. Sporophyte, seta 1.2-2.0 mm long, smooth, 1 per perichaetium; theca 0.45 × 0.2 mm, with circa 32 columns of quadratic-oblong, collenchymatous exothecial cells; peristome probably a scariosus type (only basal OP seen), tooth base 33.5 µm wide; operculum, calyptra and spores not observed.</p><p>The semilimbate leaves, mammillose cells and the number of exothecial cells classify this species in subg. Polypodiopsis .</p></div>	https://treatment.plazi.org/id/03A3AD0FFF96FFF4A3D8FCDDFC80FD26	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bruggeman-Nannenga, Maria Alida	Bruggeman-Nannenga, Maria Alida (2025): Notes on African Fissidens, including F. colliculosus Brugg. - Nann., sp. nov., F. sciophyllus subsp. capeverdensis subsp. nov. and F. pellucidus var. ripensis comb. nov., stat. nov. Cryptogamie, Bryologie 20 (3): 15-33, DOI: 10.5252/cryptogamie-bryologie2025v46a3, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a3.pdf
