taxonID	type	description	language	source
0395879FFF8CFF8EFF649360FBF7FD46.taxon	description	zoobank. org / urn: lsid: zoobank. org: act: 67 D 7 C 053 - 7016 - 4411 - B 335 - 6 F 9 EF 5872 D 48 (Figs. 1 – 5)	en	Cognato, Anthony I., Smith, Sarah M. (2020): Acanthotomicus diaboliculus and A. enzoi (Coleoptera: Curculionidae: Scolytinae: Ipini), New Species from Southeast Asia. The Coleopterists Bulletin 74 (3): 538-541, DOI: 10.1649/0010-065X-74.3.538, URL: http://dx.doi.org/10.1649/0010-065x-74.3.538
0395879FFF8CFF8EFF649360FBF7FD46.taxon	diagnosis	Diagnosis (Male). Spines on declivital interstriae 1 and 3 – 9, including larger spines on 3, 7 and 9 distinguishes A. diaboliculus from A. enzoi, which has spines on 2 – 6 and 9 with larger spines on 4 and 9, and from A. spinosus, which has spines on 2 – 9 with larger spines on 4 – 6.	en	Cognato, Anthony I., Smith, Sarah M. (2020): Acanthotomicus diaboliculus and A. enzoi (Coleoptera: Curculionidae: Scolytinae: Ipini), New Species from Southeast Asia. The Coleopterists Bulletin 74 (3): 538-541, DOI: 10.1649/0010-065X-74.3.538, URL: http://dx.doi.org/10.1649/0010-065x-74.3.538
0395879FFF8CFF8EFF649360FBF7FD46.taxon	description	Description. Holotype, male, total mean length 2.33 mm (paratypes 2.3 – 2.5 mm, n = 3), 3.33 × as long as wide; antennae, head, legs, thorax, and elytra reddish-brown (Figs. 1 – 4). Head. Epistomal setae present. Frons slightly convex, granulate, scattered punctures; scattered frontal tubercles approximately mid-frons, one larger median tubercle (Fig. 3). Scape longer than funicle; five funicular antennomeres including pedicel; antennal club flattened, longer than wide, with sensillae on anterior face, partitioned by two procurved sutures. Pronotum. Mean width 0.73 mm (0.7 – 0.8 mm, n = 3), 1.23 × as long as wide; summit approximately on anterior third, scalpriform rugosities with interspersed setae on anterior half; posterior half of disc shining, shallowly, minutely punctate, with scattered erect hair-like setae; base transverse, curved laterally (Figs. 1, 2). Scutellar shield. Triangular, flush with elytra (Fig. 1). Elytra. Together 1.75 × as long as wide, 1.56 × longer than pronotum. Elytral base transverse. Discal strial punctures uniseriate, nearly contiguous, shallowly impressed, 1.5 × width of interstriae; interstrial setae uniseriate, 1 – 1.5 × width of interstria (Fig. 2). Declivital margin of each elytron with four smaller and three larger subequal acute spines, the smallest as long as two strial punctures and the largest at least as long as three strial punctures; small first spine on interstria 1; large second spine on interstria 3; small third spine on interstria 4; small fourth spine on interstria 5; small fifth spine on interstria 6; large sixth spine on interstria 7; small seventh spine on interstria 8; large eighth spine on interstria 9 and comprising the apex of the declivity; apical third of interstriae 5 – 9 granulate / tuberculate (Figs. 1, 2, 4). Declivital face with striae 1 and 2 curving laterally, declivital suture setose. Female. Similar to male in most features except frons concave, glabrous, without median large granule; elytral spines and tubercles reduced in size (Fig. 5).	en	Cognato, Anthony I., Smith, Sarah M. (2020): Acanthotomicus diaboliculus and A. enzoi (Coleoptera: Curculionidae: Scolytinae: Ipini), New Species from Southeast Asia. The Coleopterists Bulletin 74 (3): 538-541, DOI: 10.1649/0010-065X-74.3.538, URL: http://dx.doi.org/10.1649/0010-065x-74.3.538
0395879FFF8CFF8EFF649360FBF7FD46.taxon	materials_examined	Type Material. Holotype: male, “ THAI [LAND]: [Chaiyaphum Province] Phu Khieo N [ational] P [ark] [Wildlife Sanctuary], vii 2005, Hulcr et al. coll; Aca. spTH 32, DNA Ex [tract] ” (MSUC). Paratypes (3): “ THAILAND: Chaiyaphum: [Province] Phu Khieo Wildl. [ife] Res. [ervation], 8. vii. 2005, Hulcr et al. coll; THA 099: T 32 ” (MSUC, 1 male). CHINA: Yunnan, Lijiang, host: Quercus acutissima 1962. VI. 2 (label translated by Dr. You Li) (NMNH, 1 male, 1 female).	en	Cognato, Anthony I., Smith, Sarah M. (2020): Acanthotomicus diaboliculus and A. enzoi (Coleoptera: Curculionidae: Scolytinae: Ipini), New Species from Southeast Asia. The Coleopterists Bulletin 74 (3): 538-541, DOI: 10.1649/0010-065X-74.3.538, URL: http://dx.doi.org/10.1649/0010-065x-74.3.538
0395879FFF8CFF8EFF649360FBF7FD46.taxon	etymology	Etymology. Latin diabolus = devil, - iculus = little. A noun in apposition.	en	Cognato, Anthony I., Smith, Sarah M. (2020): Acanthotomicus diaboliculus and A. enzoi (Coleoptera: Curculionidae: Scolytinae: Ipini), New Species from Southeast Asia. The Coleopterists Bulletin 74 (3): 538-541, DOI: 10.1649/0010-065X-74.3.538, URL: http://dx.doi.org/10.1649/0010-065x-74.3.538
0395879FFF8CFF8EFF649360FBF7FD46.taxon	biology_ecology	Hosts. This species is known from Quercus (Fagaceae).	en	Cognato, Anthony I., Smith, Sarah M. (2020): Acanthotomicus diaboliculus and A. enzoi (Coleoptera: Curculionidae: Scolytinae: Ipini), New Species from Southeast Asia. The Coleopterists Bulletin 74 (3): 538-541, DOI: 10.1649/0010-065X-74.3.538, URL: http://dx.doi.org/10.1649/0010-065x-74.3.538
0395879FFF8CFF8EFF649360FBF7FD46.taxon	discussion	Notes. Holotype dissected for DNA extraction; head, pronotum, and remaining body separated and glued to a paper point and pinned. Genbank accession numbers — COI: KF 862797, 16 S: KF 862718, 28 S: KF 862750, CAD: KF 862833 (Cognato 2013).	en	Cognato, Anthony I., Smith, Sarah M. (2020): Acanthotomicus diaboliculus and A. enzoi (Coleoptera: Curculionidae: Scolytinae: Ipini), New Species from Southeast Asia. The Coleopterists Bulletin 74 (3): 538-541, DOI: 10.1649/0010-065X-74.3.538, URL: http://dx.doi.org/10.1649/0010-065x-74.3.538
0395879FFF8CFF88FD7090C6FEC4FEFF.taxon	description	zoobank. org / urn: lsid: zoobank. org: act: EE 468736 - 993 C- 46 FC-A 322 - E 5 B 4642 A 8098 (Figs. 6 – 9)	en	Cognato, Anthony I., Smith, Sarah M. (2020): Acanthotomicus diaboliculus and A. enzoi (Coleoptera: Curculionidae: Scolytinae: Ipini), New Species from Southeast Asia. The Coleopterists Bulletin 74 (3): 538-541, DOI: 10.1649/0010-065X-74.3.538, URL: http://dx.doi.org/10.1649/0010-065x-74.3.538
0395879FFF8CFF88FD7090C6FEC4FEFF.taxon	diagnosis	Diagnosis (Male). Spines on declivital interstriae 2 – 6, 9, including larger spines on 4 and 9, distinguishes A. enzoi from A. diaboliculus, which has spines on 1, 3 – 9, with larger spines on 3, 7, 9, and from A. spinosus, which has spines on 2 – 9 with larger spines on 4 – 6.	en	Cognato, Anthony I., Smith, Sarah M. (2020): Acanthotomicus diaboliculus and A. enzoi (Coleoptera: Curculionidae: Scolytinae: Ipini), New Species from Southeast Asia. The Coleopterists Bulletin 74 (3): 538-541, DOI: 10.1649/0010-065X-74.3.538, URL: http://dx.doi.org/10.1649/0010-065x-74.3.538
0395879FFF8CFF88FD7090C6FEC4FEFF.taxon	description	Description. Holotype, male, total length ∼ 2.7 mm; antennae, head, legs, thorax, and elytra reddish-brown (Figs. 6 – 9). Head. Epistomal setae present. Frons slightly convex, shagreened, punctures scattered; scattered frontal tubercles approximately mid-frons, one larger median tubercle (Fig. 8). Scape longer than funicle; five funicular antennomeres including pedicel; antennal club flattened, longer than wide, with sensillae on anterior face, partitioned by two procurved sutures. Pronotum. 0.8 mm wide, 1.25 × as long as wide; summit approximately on anterior third, scalpriform rugosities with interspersed setae on anterior half; posterior half disc shining, shallowly, minutely punctate, with scattered erect hair-like setae; base transverse, curved laterally. Scutellar shield. Triangular, flush with elytra (Fig. 6). Elytra. Together 1.78 × as long as wide. Elytral base transverse. Discal strial punctures uniseriate, nearly contiguous, impressed, subequal to width of interstriae; interstriae nearly impunctate (occasional setose puncture) (Figs. 6, 7). Declivital margin of each elytron with 4 smaller and 2 larger subequal acute spines, the smallest as long as 3 strial punctures and the largest at least as long as 6 strial punctures; small first spine on interstria 2; small second spine on interstria 3; large third spine hooked inward on interstria 4; small fourth spine on interstria 5 connected by a low tumescence to the small fifth spine on interstria 6; the sixth large spine hooked upward, arising from interstria 9 and comprising the apex of the declivity (Figs. 6, 7, 9). Declivital face with striae 1 and 2 curving laterally, declivital suture setose. Female. Unknown.	en	Cognato, Anthony I., Smith, Sarah M. (2020): Acanthotomicus diaboliculus and A. enzoi (Coleoptera: Curculionidae: Scolytinae: Ipini), New Species from Southeast Asia. The Coleopterists Bulletin 74 (3): 538-541, DOI: 10.1649/0010-065X-74.3.538, URL: http://dx.doi.org/10.1649/0010-065x-74.3.538
0395879FFF8CFF88FD7090C6FEC4FEFF.taxon	materials_examined	Type Material. Holotype: male, “ THAI [LAND]: [Phetchabun Province] Nam Nao N [ational] P [ark], Castanopsis sp., vii 2005, Hulcr et al. coll; Aca. spi, DNA Ex [tract] ” (MSUC).	en	Cognato, Anthony I., Smith, Sarah M. (2020): Acanthotomicus diaboliculus and A. enzoi (Coleoptera: Curculionidae: Scolytinae: Ipini), New Species from Southeast Asia. The Coleopterists Bulletin 74 (3): 538-541, DOI: 10.1649/0010-065X-74.3.538, URL: http://dx.doi.org/10.1649/0010-065x-74.3.538
0395879FFF8CFF88FD7090C6FEC4FEFF.taxon	etymology	Etymology. Named after our son Lorenzo (Enzo) Cognato and his often-prickly personality. Noun in apposition.	en	Cognato, Anthony I., Smith, Sarah M. (2020): Acanthotomicus diaboliculus and A. enzoi (Coleoptera: Curculionidae: Scolytinae: Ipini), New Species from Southeast Asia. The Coleopterists Bulletin 74 (3): 538-541, DOI: 10.1649/0010-065X-74.3.538, URL: http://dx.doi.org/10.1649/0010-065x-74.3.538
0395879FFF8CFF88FD7090C6FEC4FEFF.taxon	biology_ecology	Hosts. This species is known from Castanopsis (Fagaceae).	en	Cognato, Anthony I., Smith, Sarah M. (2020): Acanthotomicus diaboliculus and A. enzoi (Coleoptera: Curculionidae: Scolytinae: Ipini), New Species from Southeast Asia. The Coleopterists Bulletin 74 (3): 538-541, DOI: 10.1649/0010-065X-74.3.538, URL: http://dx.doi.org/10.1649/0010-065x-74.3.538
0395879FFF8CFF88FD7090C6FEC4FEFF.taxon	discussion	Notes. Holotype dissected for DNA extraction; head, pronotum, and remaining body separated and glued to a paper point and pinned. Reported as A. spinosus in Cognato (2013) and Gao and Cognato (2018). Genbank accession numbers — COI: KF 862795, 16 S: KF 862717, 28 S: KF 862748, CAD: KF 862831 (Cognato 2013).	en	Cognato, Anthony I., Smith, Sarah M. (2020): Acanthotomicus diaboliculus and A. enzoi (Coleoptera: Curculionidae: Scolytinae: Ipini), New Species from Southeast Asia. The Coleopterists Bulletin 74 (3): 538-541, DOI: 10.1649/0010-065X-74.3.538, URL: http://dx.doi.org/10.1649/0010-065x-74.3.538
