taxonID	type	description	language	source
37E38F0A0C8850B289B92C44B39ED79D.taxon	diagnosis	Diagnosis. Boulenophrys yezhongensis sp. nov. is assigned to the genus Boulenophrys based on molecular phylogenetic analyses and the following generic diagnostic characters: snout shield-like; projecting beyond the lower jaw; canthus rostralis distinct; chest glands small and round, closer to the axilla than to midventral line; femoral glands on rear part of thigh; vertical pupils; and the presence of subarticular tubercles at the base of each finger (Fei et al. 2016; Lyu et al. 2023). Boulenophrys yezhongensis sp. nov. can be distinguished from its congeners by a combination of the following morphological characters: body size moderate (SVL 41.2 – 46.2 mm in adult males and 51.8 – 58.6 mm in adult females); a small horn-like tubercle at edge of each upper eyelid; vomerine teeth absent; tongue not notched posteriorly; tympanum distinctly visible, round; two metacarpal tubercles on hand; toes with rudimentary webbing and narrow lateral fringes; heels overlapping when thighs are positioned at right angles to the body; tibiotarsal articulation reaching the level to the middle of eye when leg stretched forward; a single internal subgular vocal sac in adult males; in breeding males, nuptial pads with black nuptial spines on the dorsal bases of the first and second fingers.	en	Liu, Jing, Feng, Chao-Bo, Shen, Tuo, Li, Shi-Ze, Cheng, Yanlin, Wei, Gang, Wang, Bin, Su, Haijun (2025): A new species of the genus Boulenophrys (Anura, Megophryidae) from Guizhou, China. Herpetozoa 38: 117-136, DOI: 10.3897/herpetozoa.38.e155430
37E38F0A0C8850B289B92C44B39ED79D.taxon	description	Description of holotype. CIB SC 20240531005 (Figs 4, 5). SVL 44.4 mm; head width greater than head length (HDW / HDL ratio about 1.2); snout obtusely pointed, protruding well beyond the margin of the lower jaw in ventral view; loreal region vertical and concave; canthus rostralis well-developed; top of head flat in dorsal view; eye large, eye diameter 40.0 % of head length; pupils vertical; nostril orientated laterally, closer to snout than eye; tympanum distinct, TYP / EYE ratio 0.6; vomerine ridges present and vomerine teeth absent; margin of tongue smooth, not notched behind. Forelimbs slender and comparatively short relative to those of congeneric species; length of lower arm and hand 44.4 % of SVL; fingers robust; relative finger lengths: II <IV <I <III; digit tips globular, without lateral fringes; distinct subarticular tubercles at finger bases; two prominent metacarpal tubercles, oval-shaped, the inner larger than the outer. Hindlimbs slender (HLL / SVL 1.47); heels overlapping when thighs at right angles to body, tibiotarsal articulation reaching mid-eye when leg stretched forward; tibia longer than thigh; relative toe lengths: I <II <V <III <IV; toes round, slightly dilated; subarticular tubercles at toe base; toes with rudimentary webbing, narrow and barely visible lateral fringe; inner metatarsal tubercle oval-shaped; outer metatarsal tubercle absent. Dorsal skin rough, with numerous brownish-yellow granules; several large warts scattered on flanks; tubercles on the dorsum forming a weak V-shaped ridge; two discontinuous dorsolateral parallel ridges on either side of the V-shaped ridges; tubercles on flanks, dorsal thighs, and tibias; supratympanic fold distinct; white granules with black hard spines on temporal fold region and adjacent areas. Ventral surface smooth with numerous white granules; pectoral glands indistinct; femoral glands on rear of thighs; inner thighs and shanks with white granules, each tipped with single black hard spine.	en	Liu, Jing, Feng, Chao-Bo, Shen, Tuo, Li, Shi-Ze, Cheng, Yanlin, Wei, Gang, Wang, Bin, Su, Haijun (2025): A new species of the genus Boulenophrys (Anura, Megophryidae) from Guizhou, China. Herpetozoa 38: 117-136, DOI: 10.3897/herpetozoa.38.e155430
37E38F0A0C8850B289B92C44B39ED79D.taxon	description	Secondary sexual characters. Adult males (SVL 41.2 – 46.2 mm) are smaller than adult females (SVL 51.8 – 58.6 mm). Adult males have a single subgular vocal sac. In breeding males, there are dark black nuptial pads at the bases of the first and second fingers, and these pads are covered with fine, dense black nuptial spines. Moreover, white granules are present on the inner thighs, around the anus, the temporal fold region, and these granules are adorned with black hard spines. In contrast, females possess only white granules.	en	Liu, Jing, Feng, Chao-Bo, Shen, Tuo, Li, Shi-Ze, Cheng, Yanlin, Wei, Gang, Wang, Bin, Su, Haijun (2025): A new species of the genus Boulenophrys (Anura, Megophryidae) from Guizhou, China. Herpetozoa 38: 117-136, DOI: 10.3897/herpetozoa.38.e155430
37E38F0A0C8850B289B92C44B39ED79D.taxon	distribution	Distribution and habitats. Boulenophrys yezhongensis sp. nov. is known from the type locality, Yezhong Nature Reserve, Shuicheng County, Guizhou Province, China, at elevations of approximately 2022 m. The individuals of the new species were frequently found on stones in the streams or gorges surrounded by evergreen broadleaved forest (Fig. 8), and two sympatric amphibian species, i. e., Rana chaochiaoensis and Quasipaa boulengeri, were found.	en	Liu, Jing, Feng, Chao-Bo, Shen, Tuo, Li, Shi-Ze, Cheng, Yanlin, Wei, Gang, Wang, Bin, Su, Haijun (2025): A new species of the genus Boulenophrys (Anura, Megophryidae) from Guizhou, China. Herpetozoa 38: 117-136, DOI: 10.3897/herpetozoa.38.e155430
37E38F0A0C8850B289B92C44B39ED79D.taxon	etymology	Etymology. The specific epithet yezhongensis refers to the type locality of the species. We propose the common English name “ Yezhong Horned Toad ” and the Chinese name “ Yě Zhōng Jiǎo Chán (野钟角蟾) ” for this species.	en	Liu, Jing, Feng, Chao-Bo, Shen, Tuo, Li, Shi-Ze, Cheng, Yanlin, Wei, Gang, Wang, Bin, Su, Haijun (2025): A new species of the genus Boulenophrys (Anura, Megophryidae) from Guizhou, China. Herpetozoa 38: 117-136, DOI: 10.3897/herpetozoa.38.e155430
