taxonID	type	description	language	source
038950234342FF9FE1862F1BFDC4AF29.taxon	type_taxon	Type species: Arbela tetraonis Moore, 1879.	en	Yakovlev, Roman V., Hulsbosch, Ramon, Ahmad, Jalil, Singh, Navneet (2025): Revision of the family Metarbelidae (Lepidoptera) of the Oriental Region. XIII. Genus Indarbela Fletcher, 1922. Zootaxa 5661 (4): 553-560, DOI: 10.11646/zootaxa.5661.4.5, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
038950234342FF9FE1862F1BFDC4AF29.taxon	discussion	The replacement name for Arbela Moore, 1879.	en	Yakovlev, Roman V., Hulsbosch, Ramon, Ahmad, Jalil, Singh, Navneet (2025): Revision of the family Metarbelidae (Lepidoptera) of the Oriental Region. XIII. Genus Indarbela Fletcher, 1922. Zootaxa 5661 (4): 553-560, DOI: 10.11646/zootaxa.5661.4.5, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
038950234342FF9FE1862F1BFDC4AF29.taxon	description	Redescription. Male. Moths of medium size (length of forewing 9 – 13 mm). Antenna bipectinate, setae 4 times longer than antenna stem in diameter; antenna twice shorter than forewing. Forewing with pattern of transverse brown strokes of various widths on light brown or yellowish background, with expressed discal spot of medium size. In some specimens, brown stripes on the forewing almost fused. Hindwing brown, without pattern. Male genitalia. Uncus long, slightly extending distally, apically with deep semicircular bifurcation, lobes of uncus wide, semicircular; gnathos arms of medium length, ribbon-like; scaphium and subscaphium fused into long spindle-like tube; valve short, wide, semicircular, with smooth edges, saccular edge (in basal half) strongly sclerotized, spoon-like; juxta of medium size, heart-like; saccus almost reduced; phallus twice shorter than valve, with greatly extended bilobed basal process, vesica aperture about 1 / 3 of phallus in length, vesica without cornuti. Female. Moths of medium size (length of forewing 10 – 15 mm). Antenna bipectinate, setae twice longer than antenna stem in diameter; antenna three times shorter than forewing. Forewing with pattern of transverse brown strokes, partially fused into bands, background of fore wing light brown, with distinct small rounded discal spot. Hindwing light brown, with blurred rounded brown spots. Female genitalia. Ovipositor short, papillae anales semicircular, anterior apophyses almost equal to posterior apophyses in length, short, thick; antrum funnel-shaped; ductus of medium length, sclerotized, bursa of medium size, rounded, without signa.	en	Yakovlev, Roman V., Hulsbosch, Ramon, Ahmad, Jalil, Singh, Navneet (2025): Revision of the family Metarbelidae (Lepidoptera) of the Oriental Region. XIII. Genus Indarbela Fletcher, 1922. Zootaxa 5661 (4): 553-560, DOI: 10.11646/zootaxa.5661.4.5, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
038950234342FF9FE1862F1BFDC4AF29.taxon	diagnosis	Diagnosis. The genus Indarbela clearly differs from the close genera (with the forked uncus) of the Oriental Metarbelidae in the following characters: - the special pattern of stripes on the forewing, - the poorly extended apex of the uncus, - the specific spoon-like extension of the saccular edge of the valve, - the well-structured bursa.	en	Yakovlev, Roman V., Hulsbosch, Ramon, Ahmad, Jalil, Singh, Navneet (2025): Revision of the family Metarbelidae (Lepidoptera) of the Oriental Region. XIII. Genus Indarbela Fletcher, 1922. Zootaxa 5661 (4): 553-560, DOI: 10.11646/zootaxa.5661.4.5, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
038950234342FF98E1862B97FCD6A80B.taxon	description	Figs 1 – 20	en	Yakovlev, Roman V., Hulsbosch, Ramon, Ahmad, Jalil, Singh, Navneet (2025): Revision of the family Metarbelidae (Lepidoptera) of the Oriental Region. XIII. Genus Indarbela Fletcher, 1922. Zootaxa 5661 (4): 553-560, DOI: 10.11646/zootaxa.5661.4.5, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
038950234342FF98E1862B97FCD6A80B.taxon	materials_examined	Type locality: Ceylon. Type material: holotype (male) in NHMUK.	en	Yakovlev, Roman V., Hulsbosch, Ramon, Ahmad, Jalil, Singh, Navneet (2025): Revision of the family Metarbelidae (Lepidoptera) of the Oriental Region. XIII. Genus Indarbela Fletcher, 1922. Zootaxa 5661 (4): 553-560, DOI: 10.11646/zootaxa.5661.4.5, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
038950234342FF98E1862B97FCD6A80B.taxon	materials_examined	Material examined. Male (holotype of Cossus 4 - notatus Walker, 1856), Ceylon (NHMUK); male (holotype of Cossus abruptus Walker, 1865) E. Ind. (NHMUK); 1 male, 4 females (syntypes of Cossus tesselatus Moore, 1879), Calcutta, May [18] 61, May [18] 63, Apr. [18] 69 (NHMUK, MfN); 1 female (holotype of Arbela tetraonis Moore, 1879), Bombay (NHMUK); 1 female, Bangladesch, Tschittagong, iv – v. 1987 (MSW); 1 male, Ceylon (NHMUK, number specimen 012832510, slide 010315541); 1 female, Ceylon, Dec. [18] 93 (NHMUK, number specimen 012832497, slide 010315528); 1 male, Colombo, i. [18] 98, Mackwood coll. (NHMUK, number specimen 012832498, slide 010315529); 1 male, Calcutta, 15. iv. 1938, D. G. Sevastopulo (NHMUK, number specimen 012832499, slide 010315530); 1 male, Nepal, Rapti Tal, Monakhari Khola, Belwa, 350 m, 8. v. 1967, leg. Dierl, Forster & Schacht (ZSM); 1 male, 1 female, S. Indien, Madurai, 18. vii. 1985, leg. M. Eckrich (ZSM); 1 male, Himachal Pradesh, Hamirpur, 07. vi. [20] 08, leg. P. C. Pathania & A. Seni (NZCZSI); 1 male, Maharashtra, Amravati, 18. ix. 2022, leg. Jalil Ahmad (NZCZSI); 1 male, Tamil Nadu, Krishnagiri, Oppathavadi, 10. xii. 2021, leg. S. Chand & Party (NZCZSI).	en	Yakovlev, Roman V., Hulsbosch, Ramon, Ahmad, Jalil, Singh, Navneet (2025): Revision of the family Metarbelidae (Lepidoptera) of the Oriental Region. XIII. Genus Indarbela Fletcher, 1922. Zootaxa 5661 (4): 553-560, DOI: 10.11646/zootaxa.5661.4.5, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
038950234342FF98E1862B97FCD6A80B.taxon	description	External and morphological variability. Size (see the Redescription of the Genus). Colour: forewing background variable from light brown to chocolate brown (which in the collectible specimens may depend on the natural pattern fading), in some males line pattern on forewing very dense, almost merging. Similar variability in forewing background colour and density of spotted pattern on forewing in females. Minor variability in sacculus length, depth of incisura in uncus, width of valva, which we interpret as individual variability.	en	Yakovlev, Roman V., Hulsbosch, Ramon, Ahmad, Jalil, Singh, Navneet (2025): Revision of the family Metarbelidae (Lepidoptera) of the Oriental Region. XIII. Genus Indarbela Fletcher, 1922. Zootaxa 5661 (4): 553-560, DOI: 10.11646/zootaxa.5661.4.5, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
038950234342FF98E1862B97FCD6A80B.taxon	distribution	Distribution (Fig. 21): Bangladesh, India (Andhra Pradesh, Bihar, Bombay, Delhi, Himachal Pradesh, Karnataka, Kerala, Madhya Pradesh, Maharashtra, Orissa, Rajasthan, Tamil Nadu, Uttar Pradesh and West Bengal), Nepal, Sri Lanka (Walker 1856, 1865; Moore 1879 a, b, 1883; Gaede 1933; Sevastopulo 1962; Ali et al. 1996; Yakovlev & Zolotuhin 2020; Chandra et al. 2021; Ahmad et al. 2023; Anonymous 2025). Reported for Pakistan (Arshad & Hafiz 1983; Gul & Chaudhry 1992).	en	Yakovlev, Roman V., Hulsbosch, Ramon, Ahmad, Jalil, Singh, Navneet (2025): Revision of the family Metarbelidae (Lepidoptera) of the Oriental Region. XIII. Genus Indarbela Fletcher, 1922. Zootaxa 5661 (4): 553-560, DOI: 10.11646/zootaxa.5661.4.5, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
038950234342FF98E1862B97FCD6A80B.taxon	biology_ecology	Host plants. It is a known pest of woody vegetation in the region under the name “ Bark-Borer ”. It has been noted to feed on numerous species of trees: Moringa oleifera Lam. (Moringaceae), Syzygium cumini (L.) Skeels, Psidium L., Corymbia citriodora (Hook.) K. D. Hill & L. A. S. Johnson, Eucalyptus camaldulensis Dehnh. (Myrtaceae), Acacia lenticularis Buch. - Ham. ex Benth., A. nilotica (L.) Delile, A. catechu (L. f.) Willd., Albizia odoratissima (L. f.) Benth., A. lebbeck (L.) Benth., A. procera (Roxb.) Benth., Cassia fistula L., Falcataria falcata (L.) Greuter & R. Rankin, Sesbania cannabina (Retz.) Poir. (Fabaceae), Eriobotrya japonica (Thunb.) Lindl., Prunus domestica L., P. armeniaca L. (Rosaceae), Gmelina arborea Roxb., Tectona grandis L. f. (Lamiaceae), Ziziphus jujube Mill., Z. mauritiana Lamk. (Rhamnaceae), Punica granatum L. (Lythraceae), Artocarpus heterophyllus Lam., Morus nigra L., Ficus benghalensis L., F. carica L. (Moraceae), Mangifera L., Anacardium occidentale L. (Anancardiaceae), Citrus limon (L.) Osbeck, C. sinensis (L.) Osbeck, C. reticulata Blanco, 1837, C. × paradise Macfad., Chloroxylon swietenia DC. (Rutaceae), Litchi chinensis Sonn. (Sapindaceae), Phyllanthus emblica L. (Phyllanthaceae), Camellia sinensis (L.) Kuntze (Theaceae), Grewia asiatica L., Theobroma cacao L. (Malvaceae), Hevea brasiliensis Müll. Arg. (Euphorbiaceae), Manilkara zapota (L.) P. Royen, Mimusops elengi L. (Sapotaceae), Mitragyna parvifolia (Roxb.) Korth (Rubiaceae), Populus L. (Salicaceae), Terminalia arjuna (Roxb.) Wight & Arn. (Combretaceae) (Gardner 1945; Sandhu et al. 1979, 1997; Lal & Singh 1982; Arshad & Hafiz 1983; Teli et al. 1983; Das et al. 1985; Verma 1985; Remadevi 1989; Kumawat & Swaminathan 1990; Mote & Tambe 1990; Baksha 1991; Shevale 1991; Gul & Chaudhry 1992; Kumar 1994; Ali et al. 1996; Mathew 2002; Rao & Prasad 2004; Sasidharan et al. 2010; Plantwise Plus 2022).	en	Yakovlev, Roman V., Hulsbosch, Ramon, Ahmad, Jalil, Singh, Navneet (2025): Revision of the family Metarbelidae (Lepidoptera) of the Oriental Region. XIII. Genus Indarbela Fletcher, 1922. Zootaxa 5661 (4): 553-560, DOI: 10.11646/zootaxa.5661.4.5, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
