taxonID	type	description	language	source
038B879CFF8CC577FF0CF98294CBFC07.taxon	description	(Figs 1 – 7) Material examined. Armenia, Kotayk Prov., nr. pre-Christian pagan temple of Garni in Garni Vill., 2 females and 1 male reared 31. III – 13. IV. 2020 from capitula of Centaurea solstitialis collected on 9. VIII. 2019 (coll. D. A. Evstigneev). Distribution. Turkey, Greece, Moldova (White & Clement, 1987; White & V. Korneyev, 1989), Ukraine (Kameneva & V. Korneyev, 1987; White & V. Korneyev, 1989; V. Korneyev & White, 1993), Israel (White & V. Korneyev, 1989), Azerbaijan (V. Korneyev & White, 1993) and Iran (Karimpour, 2011; Mohamadzade Namin & V. Korneyev, 2018). Comments. This species is recorded from Armenia for the first time. It was known from Turkey, Azerbaijan and Iran neighbouring with Armenia. The morphological details of U. sirunaseva are illustrated in Figs 1 – 7. This species is characterised by distinct subbasal, discal, preapical and apical crossbands (crossbands are listed from wing base to wing apex), distinct preapical primary and secondary steps and truncate apex of the aculeus. The species belongs to the Urophora solstitialis species – group (V. Korneyev & White, 1993) and differs from the other members of the group in the entirely yellow femur without any black spots (Figs 1, 2) and the short distance between the primary and secondary steps of the aculeus (Figs 6, 7).	en	Evstigneev, D. A., Glukhova, N. V. (2022): Tephritid flies (Diptera: Tephritidae) of the Caucasus and Transcaucasia: new records and new host plants. Zoosystematica Rossica (China) 31 (1): 118-129, DOI: 10.31610/zsr/2022.31.1.118, URL: https://doi.org/10.31610/zsr/2022.31.1.118
038B879CFF8CC574FCABFB9B9654FF48.taxon	description	(Figs 8 – 14) Material examined. Armenia, Ararat Prov., nr. Khor Virap monastery, stretch of rocky desert in hills, 2 females and 2 males reared 12. VIII. 2018 from capitula of Lactuca orientalis collected on 3. VIII. 2018 (coll. D. A. Evstigneev). Distribution. China (Becker, 1907), Armenia (Zaitzev, 1947), Kazakhstan (Richter, 1965), Syria, Iraq, Jordan (V. Korneyev & Dirlbek, 2000), Iran (Gilasian & Merz, 2008), Azerbaijan, Turkmenistan, Uzbekistan, Kyrgyzstan, Tajikistan (V. Korneyev et al., 2011), Turkey (Koçak & Kemal, 2013; Yaran & Kütük, 2015). Comments. Lactuca orientalis (Boiss.) Boiss. [syn. Scariola orientalis (Boiss.) Soják] is recorded for the first time as a host plant of H. roborowskii. Previously Lactuca sp. (Mohamadzade Namin et al., 2010) and L. serriola L. (Pourhaji et al., 2016) were recorded as host plants of this species. The morphological details of H. roborowskii are illustrated in Figs 8 – 14. There is a hyaline area in cell r 4 + 5: it is a continuation of broad hyaline area in cell m and it extends towards crossvein R-M (Figs 8, 9, 13). Tribe Tephritini	en	Evstigneev, D. A., Glukhova, N. V. (2022): Tephritid flies (Diptera: Tephritidae) of the Caucasus and Transcaucasia: new records and new host plants. Zoosystematica Rossica (China) 31 (1): 118-129, DOI: 10.31610/zsr/2022.31.1.118, URL: https://doi.org/10.31610/zsr/2022.31.1.118
038B879CFF8FC574FF03FF6F96D7FB6D.taxon	description	(Fig. 15) Material examined. Russia: Kabardino – Balkarian Republic, Elbrus Distr., nr. Terskol Vill., meadow on mountainside, 2 females and 1 male reared 28. VIII. 2014 from capitula of Psephellus sp. collected on 20 – 24. VIII. 2014 (coll. D. A. Evstigneev); Republic of Adygea, Maykop Distr., Lagonaki Plateau, mountain meadow, 1 male reared 14. VIII. 2014 from capitula of Centaurea polyphylla collected on 5. VIII. 2014 (coll. D. A. Evstigneev); Republic of North Ossetia – Alania, Western Digoria, Tana glade bog, 1 km W of Kubus Mt., 42.89350 ° N 43.57733 ° E, 2077 m, 12. IX. 2018, 2 females and 2 males, sweep-netting (coll. A. A. Przhiboro). Georgia, Tbilisi, 2 males reared 31. VII. 2019 from capitula of Amberboa glauca collected on 29. VII. 2019 (coll. D. A. Evstigneev). Distribution. Eurasia and North Africa to Ethiopia (Morgulis et al., 2015). Comments. The above specimens were reared from the capitula of several plant species. Centaurea polyphylla Ledeb. ex Nordm. [syn. Grosshei mia polyphylla (Ledeb.) Holub, G. ossica (C. Koch) Sosn. et Takht.] and Amberboa glauca (Willd.) Grossh. are recorded for the first time as host plants of A. helianthi. The habitus of A. helianthi is illustrated in Fig. 15. The morphological details of specimens from the Caucasus and Transcaucasia agree with those of specimens from European Russia (Evstigneev, 2016).	en	Evstigneev, D. A., Glukhova, N. V. (2022): Tephritid flies (Diptera: Tephritidae) of the Caucasus and Transcaucasia: new records and new host plants. Zoosystematica Rossica (China) 31 (1): 118-129, DOI: 10.31610/zsr/2022.31.1.118, URL: https://doi.org/10.31610/zsr/2022.31.1.118
038B879CFF8FC574FF03FAB594F6FE51.taxon	description	(Figs 16 – 22) Material examined. Armenia, Gegharkunik Prov., nr. Berdkunk Vill., shore of Lake Sevan contaminat- ed with excreta of farm animals, 6 females and 2 males reared 2. VIII. 2021 from capitula of Carduus nutans collected on 1. VIII. 2021 (coll. D. A. Evstigneev). Distribution. Kazakhstan, Mongolia (Richter, 1965, 1970), France (Foote, 1984), central European part of Russia (Richter, 1970), Moldova (V. Korneyev & Kameneva, 1992), Italy, Germany, Switzerland (Merz, 1994), Hungary (Merz, 2000), Andorra (Merz, 2001), Iran (Mohamadza- de Namin, 2011), Turkey (Koçak & Kemal, 2013), Albania, Ukraine (S. Korneyev, 2016). Comments. Tephritis hendeliana is recorded from Armenia and Transcaucasia for the first time. The larvae were found to develop in the capitula of Carduus nutans L. (syn. C. thoermeri Weinm.), a typical host plant of this species. The morphological details of T. hendeliana are illustrated in Figs 16 – 22; they are in agreement with the diagnosis of the species. The glans of the phallus of T. hendeliana (Fig. 20) is illustrated for the first time.	en	Evstigneev, D. A., Glukhova, N. V. (2022): Tephritid flies (Diptera: Tephritidae) of the Caucasus and Transcaucasia: new records and new host plants. Zoosystematica Rossica (China) 31 (1): 118-129, DOI: 10.31610/zsr/2022.31.1.118, URL: https://doi.org/10.31610/zsr/2022.31.1.118
038B879CFF8FC574FCBAFD9E9200FB39.taxon	description	(Figs 23 – 25) Material examined. Armenia, Ararat Prov., nr. Zod Vill., “ Salt marshes ” natural monument heavily pollut- ed by people and farm animals, tractor dug area with excreta of farm animals, 1 male reared 10. VIII. 2021 from capitulum of Lactuca tatarica collected on 7. VIII. 2021 (coll. D. A. Evstigneev). Distribution. Mongolia (Richter, 1975), China (Foote, 1984), Kyrgyzstan (V. Korneyev & Kameneva, 1993), Russia, Ukraine (S. Korneyev & Karpyuk, 2009), Iran (Gharajedaghi et al., 2011). Comments. This species is recorded from Armenia and Transcaucasia for the first time. The larvae develop in the capitula of Lactuca tatarica (L.) C. A. Mey. (V. Korneyev & Kameneva, 1993). The morphological details of T. oedipus male from Armenia are illustrated in Figs 23 – 25.	en	Evstigneev, D. A., Glukhova, N. V. (2022): Tephritid flies (Diptera: Tephritidae) of the Caucasus and Transcaucasia: new records and new host plants. Zoosystematica Rossica (China) 31 (1): 118-129, DOI: 10.31610/zsr/2022.31.1.118, URL: https://doi.org/10.31610/zsr/2022.31.1.118
038B879CFF8FC575FCBAFB469300FA66.taxon	description	(Figs 26 – 32) Material examined. Armenia, Vayots Dzor Prov., between Mozrov Vill. and Arpi Vill., crumbling mountainside along dirty road, 2 females reared 19. VII. 2018 from capitula of Reichardia dichotoma collected on 18. VII. 2018 (coll. D. A. Evstigneev). Russia, Republic of North Ossetia – Alania: nr. Vladikavkaz, ca. 2 km S of Chmi Vill., mountainside facing Suargom River, 4 females and 3 males reared 10. VIII. 2020 from capitula of Reichardia dichotoma collected on 9. VIII. 2020 (coll. D. A. Evstigneev); Western Digoria, Tana Glade bog, 1 km W of Kubus Mt., 42.89350 ° N 43.57733 ° E, 2077 m, 12. IX. 2018, 1 female, sweep-netting (coll. A. A. Przhiboro). Distribution. Transpalaearctic species (Merz, 1994). Comments. Larvae of this species live in the capitula of a wide range of Asteraceae (White, 1988; Merz, 1994; Freidberg & Kugler, 1989). Reichardia dichotoma (DC.) Freyn (syn. R. glauca Matthews) is recorded for the first time as a host plant of T. amoena. Previously, another species of the genus Reichardia, R. picroides (L.) Roth, was recorded as a host plant for T. amoena in Italy (Mazzon et al., 2008). The morphological details of T. amoena are illustrated in Figs 26 – 32. This species can be distinguished from the similar Trupanea stellata (Fuessly, 1775) by the following features of wing coloration (Fig. 26): (1) brown ray on DM - Cu crossvein reaching posterior margin of wing (brown ray enclosing DM-Cu crossvein and apical part of Cu 1 vein); (2) narrow brown band extending from pterostigma to subapical area of cell br; (3) brown ray extending from subapical star-shaped spot towards vein C, and reaching vein C at level of crossvein R-M (it is necessary to draw a line through crossvein R-M and continue it towards vein C).	en	Evstigneev, D. A., Glukhova, N. V. (2022): Tephritid flies (Diptera: Tephritidae) of the Caucasus and Transcaucasia: new records and new host plants. Zoosystematica Rossica (China) 31 (1): 118-129, DOI: 10.31610/zsr/2022.31.1.118, URL: https://doi.org/10.31610/zsr/2022.31.1.118
038B879CFF8EC572FCABF9AA9529F81C.taxon	description	(Figs 33 – 36) Material examined. Russia, Republic of North Ossetia – Alania, nr. Vladikavkaz, ca. 2 km S of Chmi Vill., Suargom River bank, 3 females reared 16. VIII. 2020 from capitula of Cladochaeta candidissima collected on 9. VIII. 2020 (coll. D. A. Evstigneev). Distribution. Transpalaearctic species (Merz, 1994). Comments. Larvae of this species live in the capitula of a wide range of Asteraceae (White, 1988; Merz, 1994; Freidberg & Kugler, 1989). Cladochaeta candidissima (M. Bieb.) DC. (syn. Helichrysum candidissimum Loudon) is recorded for the first time as a host plant of T. stellata. The morphological details of T. stellata are illustrated in Figs 33 – 36. This species can be identified by the following combination of wing coloration features: (1) brown ray on DM-Cu crossvein not reaching hind posterior margin of wing (brown ray located only on crossvein DM-Cu; vein Cu 1 entirely within a hyaline area); (2) no brown band extending from pterostigma to subapical area of cell br; (3) brown ray extending from subapical star-shaped spot to vein C and reaching it before level of crossvein R-M (Fig. 33).	en	Evstigneev, D. A., Glukhova, N. V. (2022): Tephritid flies (Diptera: Tephritidae) of the Caucasus and Transcaucasia: new records and new host plants. Zoosystematica Rossica (China) 31 (1): 118-129, DOI: 10.31610/zsr/2022.31.1.118, URL: https://doi.org/10.31610/zsr/2022.31.1.118
038B879CFF89C570FCBAFAF79399F81D.taxon	description	(Figs 37 – 43) Material examined. Armenia: Erevan, vicinity of Erebuni fortress, 1 female reared 18. VII. 2017 from capitulum of Centaurea solstitialis collected on 7. VII. 2017 (coll. D. A. Evstigneev); Kotayk Prov., near pre-Christian pagan temple of Garni in Garni Vill., 1 female and 3 males reared 31. III – 12. IV. 2020 from capitula of Centaurea solstitialis collected on 9. VIII. 2019 (coll. D. A. Evstigneev). Distribution. Bulgaria, Greece, Italy, Turkey (White, 1989) and Iran (Karimpour, 2011; V. Korneyev et al., 2013). Comments. This species is recorded from Armenia and Transcaucasia for the first time. The morphological details of T. uncinata are illustrated in Figs 37 – 43. The members of Terellia virens species – group reliably differ from each other in the shape of glans of the phallus. White (1989) described T. uncinata, recorded Centaurea solstitialis L. as its host plant, and designated a male reared from C. solstitialis in Italy (Puglia, San Severo) as the holotype. Terellia uncinata became the third species in the Terellia virens species – group specified by V. Korneyev (1985), in addition to T. virens (Loew, 1846) and T. zerovae Korneyev, 1985. The latter species was described from specimens collected in Dushanbe (Tajikistan); a male reared from the capitulum of Centaurea iberica Trevir. ex Spreng was designated as the holotype. Subsequently, three more species from T. virens species – group were described (V. Korneyev et al., 2013): T. freidbergi V. Korneyev, 2013, associated with capitula of C. behen L., T. ivannikovi V. Korneyev et Evstigneev, 2013, associated with capitula of C. chartolepis Greuter (syn. Chartolepis intermedia Boiss), and T. whitei V. Korneyev et Mohamadzade Namin, 2013, associated with capitula of various species of the genus Cousinia.	en	Evstigneev, D. A., Glukhova, N. V. (2022): Tephritid flies (Diptera: Tephritidae) of the Caucasus and Transcaucasia: new records and new host plants. Zoosystematica Rossica (China) 31 (1): 118-129, DOI: 10.31610/zsr/2022.31.1.118, URL: https://doi.org/10.31610/zsr/2022.31.1.118
038B879CFF8AC57EFF0CF9649325F81D.taxon	description	(Figs 44 – 51) Material examined. Armenia, Aragatsotn Prov., Arailer Mt., mountainside facing Yeghvard town, mountain meadow, 1 female reared 30. III. 2019 from capitulum of Centaurea sp. collected on 24. VII. 2018 (coll. D. A. Evstigneev). Russia, Kabardino-Balkarian Republic, Elbrus Distr., Cheget Mt., ca. 2100 m, 1 male reared 21. II. 2015 from capitulum of Centaurea cheiranthifolia collected on 18 – 25. VII. 2014 (coll. D. A. Evstigneev). Distribution. Austria, France (Hendel, 1927), Armenia (V. Korneyev, 1983), Switzerland (Merz, 1994), Hungary (Mihályi, 1959), Ukraine (Merz, 1994; V. Korneyev, 2004), Czechoslovakia [without further details] (Merz, 1994), Slovakia (Dirlbek, 1999), North Caucasus in Russia (V. Korneyev, 2004), Italy (Mazzon et al., 2021). Comments. Centaurea cheiranthifolia Willd. is recorded for the first time as a host plant of X. laticauda. The morphological details of X. laticauda are illustrated in Figs 44 – 51. This species can be distinguished from the widespread X. miliaria (Schrank, 1781) by the following characters: thorax and abdomen black (Figs 44, 47, 50), and margins of preapical narrowed part of aculeus smooth, not serrate (Fig. 46).	en	Evstigneev, D. A., Glukhova, N. V. (2022): Tephritid flies (Diptera: Tephritidae) of the Caucasus and Transcaucasia: new records and new host plants. Zoosystematica Rossica (China) 31 (1): 118-129, DOI: 10.31610/zsr/2022.31.1.118, URL: https://doi.org/10.31610/zsr/2022.31.1.118
