taxonID	type	description	language	source
0387A34AFFFCFFAB51CA48C305E0B980.taxon	type_taxon	Type species: Murex latifolius Bellardi, 1872	en	Kovács, Zoltán, Vicián, Zoltán (2019): Muricidae (Neogastropoda) from the Middle Eocene of the Hungarian Paleogene Basin. Fragmenta Palaeontologica Hungarica 36: 31-51, DOI: 10.17111/FragmPalHung.2019.36.31, URL: https://doi.org/10.17111/fragmpalhung.2019.36.31
0387A34AFFFCFFAC51B348AD025DBDA5.taxon	description	(Fig. 2)	en	Kovács, Zoltán, Vicián, Zoltán (2019): Muricidae (Neogastropoda) from the Middle Eocene of the Hungarian Paleogene Basin. Fragmenta Palaeontologica Hungarica 36: 31-51, DOI: 10.17111/FragmPalHung.2019.36.31, URL: https://doi.org/10.17111/fragmpalhung.2019.36.31
0387A34AFFFCFFAC51B348AD025DBDA5.taxon	materials_examined	Material – Coll. Gurdon (1 specimen).	en	Kovács, Zoltán, Vicián, Zoltán (2019): Muricidae (Neogastropoda) from the Middle Eocene of the Hungarian Paleogene Basin. Fragmenta Palaeontologica Hungarica 36: 31-51, DOI: 10.17111/FragmPalHung.2019.36.31, URL: https://doi.org/10.17111/fragmpalhung.2019.36.31
0387A34AFFFCFFAC51B348AD025DBDA5.taxon	discussion	Remarks – The specimen figured here is very close in morphology to the T. barattus material presented by MERLE et al. (2011, pl. 100, fig. 1). The species differs from other Middle Eocene Timbellus by rounded wide ovate aperture and straight siphonal canal. The presence in the Upper Lutetian deposits of Dudar extends both the geographic and the stratigraphic ranges of T. barattus. The species is a new record in the HPB.	en	Kovács, Zoltán, Vicián, Zoltán (2019): Muricidae (Neogastropoda) from the Middle Eocene of the Hungarian Paleogene Basin. Fragmenta Palaeontologica Hungarica 36: 31-51, DOI: 10.17111/FragmPalHung.2019.36.31, URL: https://doi.org/10.17111/fragmpalhung.2019.36.31
0387A34AFFFCFFAC51B348AD025DBDA5.taxon	distribution	Distribution – Lutetian, Hungarian Paleogene Basin: Dudar. Priabonian, NE Atlantic: Aquitanian Basin (France). Rupelian, Alpine Tethys: Bassano (N Italy).	en	Kovács, Zoltán, Vicián, Zoltán (2019): Muricidae (Neogastropoda) from the Middle Eocene of the Hungarian Paleogene Basin. Fragmenta Palaeontologica Hungarica 36: 31-51, DOI: 10.17111/FragmPalHung.2019.36.31, URL: https://doi.org/10.17111/fragmpalhung.2019.36.31
0387A34AFFFBFFAC51824F7D0595B999.taxon	description	(Figs 3 – 7)	en	Kovács, Zoltán, Vicián, Zoltán (2019): Muricidae (Neogastropoda) from the Middle Eocene of the Hungarian Paleogene Basin. Fragmenta Palaeontologica Hungarica 36: 31-51, DOI: 10.17111/FragmPalHung.2019.36.31, URL: https://doi.org/10.17111/fragmpalhung.2019.36.31
0387A34AFFFBFFAC51824F7D0595B999.taxon	materials_examined	Material – HNHM INV 2020.22. (1), Coll. Gurdon (1), Coll. Vicián (1), Coll. Zsoldos (1).	en	Kovács, Zoltán, Vicián, Zoltán (2019): Muricidae (Neogastropoda) from the Middle Eocene of the Hungarian Paleogene Basin. Fragmenta Palaeontologica Hungarica 36: 31-51, DOI: 10.17111/FragmPalHung.2019.36.31, URL: https://doi.org/10.17111/fragmpalhung.2019.36.31
0387A34AFFFBFFAC51824F7D0595B999.taxon	discussion	Remarks – The specimens are rather worn or fragmentary but the shell proportions and traces of spiral cords agree with the T. micropterus material illustrated by MERLE et al. (2011, pl. 102, figs 1 – 5). The type of Murex parvulmicropterus De Gregorio is a fragmentary juvenile specimen. It corresponds in size and morphology to T. micropterus but slightly differs in sculpture with weakly developed intervarical nodes. M. parvulmicropterus was proposed as a junior synonym of T. micropterus by MERLE et al. (2011), this arrangement is accepted herein. The small Late Eocene – Early Oligocene Timbellus rigidus (Oppenheim, 1901) (recorded from N Italy, S Czechia and S Poland) is a similar species but distinguishable by stratigraphic range and axial sculpture of marked, narrow intervarical ribs (see OPPENHEIM 1901, pl. 1, fig. 1; OPPENHEIM 1922, pl. 5, fig. 6; KRACH 1985, pl. 9, figs 8 – 10). T. micropterus is a new record in the HPB.	en	Kovács, Zoltán, Vicián, Zoltán (2019): Muricidae (Neogastropoda) from the Middle Eocene of the Hungarian Paleogene Basin. Fragmenta Palaeontologica Hungarica 36: 31-51, DOI: 10.17111/FragmPalHung.2019.36.31, URL: https://doi.org/10.17111/fragmpalhung.2019.36.31
0387A34AFFFBFFAC51824F7D0595B999.taxon	distribution	Distribution – Lutetian – Bartonian, NE Atlantic: Paris, Cotentin, and Loire Basin (France), Hampshire Basin (England), Alpine Tethys: San Giovanni Ilarione (N Italy), Hungarian Paleogene Basin: Dudar, Gánt, Tés.	en	Kovács, Zoltán, Vicián, Zoltán (2019): Muricidae (Neogastropoda) from the Middle Eocene of the Hungarian Paleogene Basin. Fragmenta Palaeontologica Hungarica 36: 31-51, DOI: 10.17111/FragmPalHung.2019.36.31, URL: https://doi.org/10.17111/fragmpalhung.2019.36.31
0387A34AFFFBFFAF519B48AE05C5BE19.taxon	description	(Figs 8 – 10)	en	Kovács, Zoltán, Vicián, Zoltán (2019): Muricidae (Neogastropoda) from the Middle Eocene of the Hungarian Paleogene Basin. Fragmenta Palaeontologica Hungarica 36: 31-51, DOI: 10.17111/FragmPalHung.2019.36.31, URL: https://doi.org/10.17111/fragmpalhung.2019.36.31
0387A34AFFFBFFAF519B48AE05C5BE19.taxon	materials_examined	Material – HNHM INV 2020.23.1 – 2. (2), Coll. Vicián (3).	en	Kovács, Zoltán, Vicián, Zoltán (2019): Muricidae (Neogastropoda) from the Middle Eocene of the Hungarian Paleogene Basin. Fragmenta Palaeontologica Hungarica 36: 31-51, DOI: 10.17111/FragmPalHung.2019.36.31, URL: https://doi.org/10.17111/fragmpalhung.2019.36.31
0387A34AFFFBFFAF519B48AE05C5BE19.taxon	discussion	Remarks – The morphology of the studied specimens (trivaricate shell, paucispiral protoconch, six rounded teleoconch whorls, long and straight siphonal canal, moderately developed spiral cords: 10 – 11 on the last whorl, wing-like varices, one fine intervarical rib between the varices) correspond to that of T. priabonicus. Other Timbellus species also appear in the Middle Eocene Alpine Tethys. From N Italy T. tripteroides (Lamarck, 1822) (= Murex stoppani De Gregorio, 1880) is known, while T. crenulatus tricarinatus (Lamarck, 1803) was recorded by DAINELLI (1915) and FABIANI (1915). T. crenulatus tricarinatus otherwise shows a wide geographic distribution: NE Atlantic (France, MERLE et al. 2011; England, TRACEY et al. 1996), North Sea Basin (Belgium, GLIBERT 1933), NE Alpine Tethys (Tatra Mts, Poland, KUŹNIAR 1910), Ukrainian Archipelago (MAKARENKO & ZELINSKAYA 1982). The specimen illustrated by ZELINSKAYA et al. (1968, pl. 15, figs 13 – 14) as Typhis schlotheimi from the Late Eocene of Ukraine is identical in morphology with the Murex trialatus specimen figured by KOENEN (1889, pl. 2, fig. 3), therefore it represents Timbellus priabonicus. The presence of T. priabonicus in the Late Lutetian HPB extends both the stratigraphic and the geographic ranges of the species.	en	Kovács, Zoltán, Vicián, Zoltán (2019): Muricidae (Neogastropoda) from the Middle Eocene of the Hungarian Paleogene Basin. Fragmenta Palaeontologica Hungarica 36: 31-51, DOI: 10.17111/FragmPalHung.2019.36.31, URL: https://doi.org/10.17111/fragmpalhung.2019.36.31
0387A34AFFFBFFAF519B48AE05C5BE19.taxon	distribution	Distribution – Lutetian, Alpine Tethys: Hungarian Paleogene Basin: Dudar. Priabonian, North Sea Basin (Germany), Ukrainian Archipelago.	en	Kovács, Zoltán, Vicián, Zoltán (2019): Muricidae (Neogastropoda) from the Middle Eocene of the Hungarian Paleogene Basin. Fragmenta Palaeontologica Hungarica 36: 31-51, DOI: 10.17111/FragmPalHung.2019.36.31, URL: https://doi.org/10.17111/fragmpalhung.2019.36.31
0387A34AFFF8FFAF51DF4B3905D1BAFA.taxon	type_taxon	Type species: Flexopteron philippinensis Shuto, 1969	en	Kovács, Zoltán, Vicián, Zoltán (2019): Muricidae (Neogastropoda) from the Middle Eocene of the Hungarian Paleogene Basin. Fragmenta Palaeontologica Hungarica 36: 31-51, DOI: 10.17111/FragmPalHung.2019.36.31, URL: https://doi.org/10.17111/fragmpalhung.2019.36.31
0387A34AFFF8FFAF51864F8F025CBA51.taxon	description	(Figs 11 – 16)	en	Kovács, Zoltán, Vicián, Zoltán (2019): Muricidae (Neogastropoda) from the Middle Eocene of the Hungarian Paleogene Basin. Fragmenta Palaeontologica Hungarica 36: 31-51, DOI: 10.17111/FragmPalHung.2019.36.31, URL: https://doi.org/10.17111/fragmpalhung.2019.36.31
0387A34AFFF8FFAF51864F8F025CBA51.taxon	materials_examined	Material – MGSH: E. 125 (holotype), E. 126 (2), E. 5194 (1), E. 5207 (1), E. 5270 (7), E. 5285 (6), E. 5514 (2), E. 5905 (1); HNHM: M. 59.7443 (> 70), M. 59.7444 (10), M. 61.2423 (2), INV 2020.24. (1); Coll. Berta (4), Coll. Evanics (1), Coll. Zsoldos (2), Coll. Vicián (1).	en	Kovács, Zoltán, Vicián, Zoltán (2019): Muricidae (Neogastropoda) from the Middle Eocene of the Hungarian Paleogene Basin. Fragmenta Palaeontologica Hungarica 36: 31-51, DOI: 10.17111/FragmPalHung.2019.36.31, URL: https://doi.org/10.17111/fragmpalhung.2019.36.31
0387A34AFFF8FFAF51864F8F025CBA51.taxon	discussion	Remarks – The “ exceptional ” similarity between Murex contabulatus Lamarck, 1803 and M. gantensis nov. sp. was already noted by SZŐTS (1953). Later STRAUSZ (1974) revised gantensis, and emended it as a subspecies of? P. contabulatus. Based on morphological differences (the teleoconch whorls of gantensis are more constricted at the base and the spines are slightly more curved than that of contabulatus, see MERLE et al. 2011, pl. 115, figs 5 – 9) the arrangement suggested by Strausz is accepted herein, and? P. contabulatus gantensis is regarded as a geographic subspecies. “ Murex ” contabulatiformis Schauroth, 1865 from the Italian Eocene is a similar form but differs by more angulate whorls.? P. contabulatus gantensis is widespread and relatively abundant in the HPB.	en	Kovács, Zoltán, Vicián, Zoltán (2019): Muricidae (Neogastropoda) from the Middle Eocene of the Hungarian Paleogene Basin. Fragmenta Palaeontologica Hungarica 36: 31-51, DOI: 10.17111/FragmPalHung.2019.36.31, URL: https://doi.org/10.17111/fragmpalhung.2019.36.31
0387A34AFFF8FFAF51864F8F025CBA51.taxon	distribution	Distribution – Lutetian – Bartonian, Hungarian Paleogene Basin: Dudar, Gánt, Csákvár, Tés, Balinka, Mesterberek, Neszmély, Tatabánya / Felsőgalla.	en	Kovács, Zoltán, Vicián, Zoltán (2019): Muricidae (Neogastropoda) from the Middle Eocene of the Hungarian Paleogene Basin. Fragmenta Palaeontologica Hungarica 36: 31-51, DOI: 10.17111/FragmPalHung.2019.36.31, URL: https://doi.org/10.17111/fragmpalhung.2019.36.31
0387A34AFFF6FFA351D64D9507DABDA5.taxon	description	(Figs 17 – 21)	en	Kovács, Zoltán, Vicián, Zoltán (2019): Muricidae (Neogastropoda) from the Middle Eocene of the Hungarian Paleogene Basin. Fragmenta Palaeontologica Hungarica 36: 31-51, DOI: 10.17111/FragmPalHung.2019.36.31, URL: https://doi.org/10.17111/fragmpalhung.2019.36.31
0387A34AFFF6FFA351D64D9507DABDA5.taxon	materials_examined	Holotype – HNHM PAL 2020.20.1., SL 13 (Figs 19 – 20). Type strata and locality – Upper Lutetian sandy clay, Tés, Hungary.	en	Kovács, Zoltán, Vicián, Zoltán (2019): Muricidae (Neogastropoda) from the Middle Eocene of the Hungarian Paleogene Basin. Fragmenta Palaeontologica Hungarica 36: 31-51, DOI: 10.17111/FragmPalHung.2019.36.31, URL: https://doi.org/10.17111/fragmpalhung.2019.36.31
0387A34AFFF6FFA351D64D9507DABDA5.taxon	etymology	Derivation of name – In honour of the collector: Márton Zsoldos (Bakonynána, Hungary).	en	Kovács, Zoltán, Vicián, Zoltán (2019): Muricidae (Neogastropoda) from the Middle Eocene of the Hungarian Paleogene Basin. Fragmenta Palaeontologica Hungarica 36: 31-51, DOI: 10.17111/FragmPalHung.2019.36.31, URL: https://doi.org/10.17111/fragmpalhung.2019.36.31
0387A34AFFF6FFA351D64D9507DABDA5.taxon	materials_examined	Material – Holotype and two additional specimens (Figs 17 – 18, 21).	en	Kovács, Zoltán, Vicián, Zoltán (2019): Muricidae (Neogastropoda) from the Middle Eocene of the Hungarian Paleogene Basin. Fragmenta Palaeontologica Hungarica 36: 31-51, DOI: 10.17111/FragmPalHung.2019.36.31, URL: https://doi.org/10.17111/fragmpalhung.2019.36.31
0387A34AFFF6FFA351D64D9507DABDA5.taxon	diagnosis	Diagnosis – Paziella (Flexopteron) species with multispiral protoconch and five teleoconch whorls, ovate aperture with strong denticles within, smooth columella, sculpture of strong spiral cords and numerous axial varices.	en	Kovács, Zoltán, Vicián, Zoltán (2019): Muricidae (Neogastropoda) from the Middle Eocene of the Hungarian Paleogene Basin. Fragmenta Palaeontologica Hungarica 36: 31-51, DOI: 10.17111/FragmPalHung.2019.36.31, URL: https://doi.org/10.17111/fragmpalhung.2019.36.31
0387A34AFFF6FFA351D64D9507DABDA5.taxon	description	Description – Multispiral protoconch of 3 1 / 4 smooth, rounded whorls, five shouldered teleoconch whorls, elongated spire, broad subsutural slope, ovate aperture, outer lip thickened by growth lamellae, strong denticles within (D 1 – D 5; D 1 – D 3 stronger than D 4 – D 5), weakly developed parietal callus, smooth columella, elongated and slightly curved siphonal canal, spiral sculpture of strong primary cords (P 1 – P 6 on the last whorl), axial sculpture of numerous sharp varices (eight on the last whorl).	en	Kovács, Zoltán, Vicián, Zoltán (2019): Muricidae (Neogastropoda) from the Middle Eocene of the Hungarian Paleogene Basin. Fragmenta Palaeontologica Hungarica 36: 31-51, DOI: 10.17111/FragmPalHung.2019.36.31, URL: https://doi.org/10.17111/fragmpalhung.2019.36.31
0387A34AFFF6FFA351D64D9507DABDA5.taxon	discussion	Remarks – Based on morphology the new species is assigned to subgenus Paziella (Flexopteron). The fragmentary Murex sp. from the shallow marine deposits of borehole Cs. 115 at Csabdi (KECSKEMÉTI-KÖRMENDY 1990, pl. 29, figs 5 – 6; refigured on Figs 17 – 18) is considered herein as a representative of P. (F.) zsoldosi n. sp. (unfortunately this specimen is not available in the mollusc collection of the MGSH). According to KECSKEMÉTI-KÖRMENDY (l. c.) the smooth, multispiral protoconch, the rounded early teleoconch whorls, as well as the weakly developed primary spiral cords and the sharp axial varices are closely allied to that of P. (F.) flexuosa (Deshayes). However, these features are common on different Eocene species of the subgenus, consequently the species level determination depends on the adult morphology. The adult shell of P. (F.) zsoldosi n. sp. is distinguishable from the Early Eocene P. (F.) flexuosa (Deshayes, 1865) and P. (F.) ogormani (Cossmann, 1923) by much stronger spiral cords and lack of spines; from the Middle Eocene P. (F.) foliacea fraterculus (Deshayes, 1865) by elongated spire, more widely spaced varices and lack of spines, and from the Middle Eocene P. (F.) elatior (Koenen, 1889) by less elongated shell and different sculpture (see MERLE et al. 2011, pls 135 – 138). The teleoconch whorls of Paziella (Flexopteron) sp. specimens from Tés (Figs 20 – 22) are broader, they represent another species. The material came from the sandy clay deposits of the Tés locality. It was associated by coastal to shallow water gastropods: Pseudobellardia auriculata (Schlotheim, 1820), Tympanotonos aculeatus (Schlotheim, 1820), Cerithium subcorvinum Oppenheim, 1894, Editharus roncanus (Brongniart, 1823), Ampullina perusta (Defrance in Brongniart, 1823), Deshayesia alpina (d’Orbigny, 1850), Stilospirula doroghensis (Oppenheim, 1892), Tectonatica pasinii (Bayan, 1870), Globularia incompleta (Zittel, 1862), Clavilithes noae Lamarck 1803, Dilatilabrum fortisi (Brongniart, 1823), Timbellus micropterus (Deshayes),? Pterochelus contabulatus gantensis (Szőts), Paziella (Flexopteron) sp., Janiopsis n. sp. and Hemiconus eszterhazyi (Papp, 1897).	en	Kovács, Zoltán, Vicián, Zoltán (2019): Muricidae (Neogastropoda) from the Middle Eocene of the Hungarian Paleogene Basin. Fragmenta Palaeontologica Hungarica 36: 31-51, DOI: 10.17111/FragmPalHung.2019.36.31, URL: https://doi.org/10.17111/fragmpalhung.2019.36.31
0387A34AFFF4FFA352104F7D0494BFED.taxon	description	(Figs 22 – 24)	en	Kovács, Zoltán, Vicián, Zoltán (2019): Muricidae (Neogastropoda) from the Middle Eocene of the Hungarian Paleogene Basin. Fragmenta Palaeontologica Hungarica 36: 31-51, DOI: 10.17111/FragmPalHung.2019.36.31, URL: https://doi.org/10.17111/fragmpalhung.2019.36.31
0387A34AFFF4FFA352104F7D0494BFED.taxon	materials_examined	Material – HNHM INV 2020.25., INV 2020.26. (2).	en	Kovács, Zoltán, Vicián, Zoltán (2019): Muricidae (Neogastropoda) from the Middle Eocene of the Hungarian Paleogene Basin. Fragmenta Palaeontologica Hungarica 36: 31-51, DOI: 10.17111/FragmPalHung.2019.36.31, URL: https://doi.org/10.17111/fragmpalhung.2019.36.31
0387A34AFFF4FFA352104F7D0494BFED.taxon	discussion	Remarks – The juvenile specimens from the Late Lutetian mollusc assemblage of Tés possess multispiral protoconch, sculpture of seven varices on the last whorl, and slightly spiny P 1. These features are typical of P. (Flexopteron). However, early whorls of different Eocene species of the subgenus are very similar (see above), and the species level identification seems impossible in lack of adult morphology. Nevertheless, the specimens indicate diverse occurrence of the subgenus in the Middle Eocene HPB.	en	Kovács, Zoltán, Vicián, Zoltán (2019): Muricidae (Neogastropoda) from the Middle Eocene of the Hungarian Paleogene Basin. Fragmenta Palaeontologica Hungarica 36: 31-51, DOI: 10.17111/FragmPalHung.2019.36.31, URL: https://doi.org/10.17111/fragmpalhung.2019.36.31
0387A34AFFF3FFA451BA4D8A05F4BD4F.taxon	type_taxon	Type species: Murex (Harmatia) stephani Noszky, 1940	en	Kovács, Zoltán, Vicián, Zoltán (2019): Muricidae (Neogastropoda) from the Middle Eocene of the Hungarian Paleogene Basin. Fragmenta Palaeontologica Hungarica 36: 31-51, DOI: 10.17111/FragmPalHung.2019.36.31, URL: https://doi.org/10.17111/fragmpalhung.2019.36.31
0387A34AFFF3FFA451D84C750584BAD2.taxon	description	(Fig. 25)	en	Kovács, Zoltán, Vicián, Zoltán (2019): Muricidae (Neogastropoda) from the Middle Eocene of the Hungarian Paleogene Basin. Fragmenta Palaeontologica Hungarica 36: 31-51, DOI: 10.17111/FragmPalHung.2019.36.31, URL: https://doi.org/10.17111/fragmpalhung.2019.36.31
0387A34AFFF3FFA451D84C750584BAD2.taxon	materials_examined	Holotype – HNHM M. 62.7135, SL 72 (P 1 spine not included) (Collected by Endre Szőts in 1956). Type strata and locality – Upper Lutetian – Lower Bartonian grey marl, Tokod, Hungary.	en	Kovács, Zoltán, Vicián, Zoltán (2019): Muricidae (Neogastropoda) from the Middle Eocene of the Hungarian Paleogene Basin. Fragmenta Palaeontologica Hungarica 36: 31-51, DOI: 10.17111/FragmPalHung.2019.36.31, URL: https://doi.org/10.17111/fragmpalhung.2019.36.31
0387A34AFFF3FFA451D84C750584BAD2.taxon	etymology	Derivation of name – The name refers to the type locality.	en	Kovács, Zoltán, Vicián, Zoltán (2019): Muricidae (Neogastropoda) from the Middle Eocene of the Hungarian Paleogene Basin. Fragmenta Palaeontologica Hungarica 36: 31-51, DOI: 10.17111/FragmPalHung.2019.36.31, URL: https://doi.org/10.17111/fragmpalhung.2019.36.31
0387A34AFFF3FFA451D84C750584BAD2.taxon	materials_examined	Material – Holotype.	en	Kovács, Zoltán, Vicián, Zoltán (2019): Muricidae (Neogastropoda) from the Middle Eocene of the Hungarian Paleogene Basin. Fragmenta Palaeontologica Hungarica 36: 31-51, DOI: 10.17111/FragmPalHung.2019.36.31, URL: https://doi.org/10.17111/fragmpalhung.2019.36.31
0387A34AFFF3FFA451D84C750584BAD2.taxon	diagnosis	Diagnosis – Trivaricate Harmatia species with rounded teleoconch whorls, ovate aperture, long siphonal canal, narrow primary spiral cords, and long primary cord spines.	en	Kovács, Zoltán, Vicián, Zoltán (2019): Muricidae (Neogastropoda) from the Middle Eocene of the Hungarian Paleogene Basin. Fragmenta Palaeontologica Hungarica 36: 31-51, DOI: 10.17111/FragmPalHung.2019.36.31, URL: https://doi.org/10.17111/fragmpalhung.2019.36.31
0387A34AFFF3FFA451D84C750584BAD2.taxon	description	Description – Medium-sized shell, protoconch missing, subfusiform teleoconch. Spire of three shouldered, rounded whorls. Last whorl 89 % of the total length of the preserved shell (without P 1 spine). Aperture ovate, internal part not seen. Siphonal canal without cord spines, long, narrow and slightly curved, penultimate siphonal canal present. Spiral sculpture of moderately developed, narrow primary cords (P 1 – P 4 visible on last whorl), axial sculpture of three spiny varices per whorl without intervarical ribs. Well-developed, long and open P 1 – P 6 spines on last whorl, P 1 spine slightly adapically curved, no intervarical ribs.	en	Kovács, Zoltán, Vicián, Zoltán (2019): Muricidae (Neogastropoda) from the Middle Eocene of the Hungarian Paleogene Basin. Fragmenta Palaeontologica Hungarica 36: 31-51, DOI: 10.17111/FragmPalHung.2019.36.31, URL: https://doi.org/10.17111/fragmpalhung.2019.36.31
0387A34AFFF3FFA451D84C750584BAD2.taxon	discussion	Remarks – The morphology of the new species corresponds to the diagnosis of genus Harmatia (see MERLE et al. 2011). The genus has been known from the Early Oligocene in Hungary and Austria with two species; the Middle Eocene occurrence of H. tokodensis n. sp. extends the stratigraphic range. Both H. stephani (NOSZKY 1940, pl. 2, fig. 4; refigured by MERLE et al. 2011, text-fig. 63 / 1), and H. guembeli longispina (NOSZKY 1940, pl. 1, fig. 31; refigured by MERLE et al. 2011, text-fig. 63 / 2) (Kiscellian, Budapest Újlak) are distinguishable by much lower spire, and H. stephani bears almost straight siphonal canal. The spire and sculpture of H. guembeli guembeli (Dreger) (Kiscellian, Bad Häring, Budapest Újlak) resemble that of H. tokodensis n. sp., the former, however, differs by strongly curved long siphonal canal. This feature cannot be seen on the fragmentary type specimen (DREGER 1892, pl. 4, fig. 5; refigured by LÖFFLER 1999, pl. 12, fig. 1, and MERLE et al. 2011, text-fig. 63 / 3), but clearly visible on other specimens illustrated by LÖFFLER (1999, pl. 12, figs 2 – 5).	en	Kovács, Zoltán, Vicián, Zoltán (2019): Muricidae (Neogastropoda) from the Middle Eocene of the Hungarian Paleogene Basin. Fragmenta Palaeontologica Hungarica 36: 31-51, DOI: 10.17111/FragmPalHung.2019.36.31, URL: https://doi.org/10.17111/fragmpalhung.2019.36.31
0387A34AFFF1FFA651744C7507B8B85A.taxon	description	(Figs 26 – 27)	en	Kovács, Zoltán, Vicián, Zoltán (2019): Muricidae (Neogastropoda) from the Middle Eocene of the Hungarian Paleogene Basin. Fragmenta Palaeontologica Hungarica 36: 31-51, DOI: 10.17111/FragmPalHung.2019.36.31, URL: https://doi.org/10.17111/fragmpalhung.2019.36.31
0387A34AFFF1FFA651744C7507B8B85A.taxon	materials_examined	Material – IFSM: 62756 (1).	en	Kovács, Zoltán, Vicián, Zoltán (2019): Muricidae (Neogastropoda) from the Middle Eocene of the Hungarian Paleogene Basin. Fragmenta Palaeontologica Hungarica 36: 31-51, DOI: 10.17111/FragmPalHung.2019.36.31, URL: https://doi.org/10.17111/fragmpalhung.2019.36.31
0387A34AFFF1FFA651744C7507B8B85A.taxon	discussion	Remarks – A fragmentary specimen is illustrated on the website of the International Fossil Shell Museum as Ponderia caillati (www. fossilshells. nl / hungeo 40. html; download: 02.10.2019), it was collected by Rudi Hessel on the mine dump of the old brown coal mine of Dudar. The size (SL 31) and the general morphology agree well with that of the P. bispinosa material presented by MERLE et al. (2001). Murex caillati Deshayes is a junior synonym of M. bispinosus J. de C. Sowerby.	en	Kovács, Zoltán, Vicián, Zoltán (2019): Muricidae (Neogastropoda) from the Middle Eocene of the Hungarian Paleogene Basin. Fragmenta Palaeontologica Hungarica 36: 31-51, DOI: 10.17111/FragmPalHung.2019.36.31, URL: https://doi.org/10.17111/fragmpalhung.2019.36.31
0387A34AFFF1FFA651744C7507B8B85A.taxon	distribution	Distribution – Lutetian – Priabonian, NE Atlantic: Paris Basin (France), Hampshire Basin (England), North Sea Basin (Germany), Alpine Tethys: Hungarian Paleogene Basin: Dudar.	en	Kovács, Zoltán, Vicián, Zoltán (2019): Muricidae (Neogastropoda) from the Middle Eocene of the Hungarian Paleogene Basin. Fragmenta Palaeontologica Hungarica 36: 31-51, DOI: 10.17111/FragmPalHung.2019.36.31, URL: https://doi.org/10.17111/fragmpalhung.2019.36.31
0387A34AFFF1FFA651C04D8A059FBD4F.taxon	type_taxon	Type species: Typhis zealandicus (Hutton, 1873)	en	Kovács, Zoltán, Vicián, Zoltán (2019): Muricidae (Neogastropoda) from the Middle Eocene of the Hungarian Paleogene Basin. Fragmenta Palaeontologica Hungarica 36: 31-51, DOI: 10.17111/FragmPalHung.2019.36.31, URL: https://doi.org/10.17111/fragmpalhung.2019.36.31
0387A34AFFF1FFA651A8496C05B2B825.taxon	type_taxon	Type species: Typhis belcheri Broderip, 1833	en	Kovács, Zoltán, Vicián, Zoltán (2019): Muricidae (Neogastropoda) from the Middle Eocene of the Hungarian Paleogene Basin. Fragmenta Palaeontologica Hungarica 36: 31-51, DOI: 10.17111/FragmPalHung.2019.36.31, URL: https://doi.org/10.17111/fragmpalhung.2019.36.31
0387A34AFFEFFFB8518B4C7506F7B87E.taxon	description	(Figs 30 – 31)	en	Kovács, Zoltán, Vicián, Zoltán (2019): Muricidae (Neogastropoda) from the Middle Eocene of the Hungarian Paleogene Basin. Fragmenta Palaeontologica Hungarica 36: 31-51, DOI: 10.17111/FragmPalHung.2019.36.31, URL: https://doi.org/10.17111/fragmpalhung.2019.36.31
0387A34AFFEFFFB8518B4C7506F7B87E.taxon	materials_examined	Material – MGSH: E. 467 (Holotype).	en	Kovács, Zoltán, Vicián, Zoltán (2019): Muricidae (Neogastropoda) from the Middle Eocene of the Hungarian Paleogene Basin. Fragmenta Palaeontologica Hungarica 36: 31-51, DOI: 10.17111/FragmPalHung.2019.36.31, URL: https://doi.org/10.17111/fragmpalhung.2019.36.31
0387A34AFFEFFFB8518B4C7506F7B87E.taxon	discussion	Remarks – Cantharus (Pollia) dudariensis Strausz was revised and reinterpreted as a representative of genus Nucellopsis by MERLE (2005 b). The genus is typical of coastal or lagoon environments. N. dudariensis is an endemic species in the HPB and very rare, only the holotype is known from the type locality, Dudar. The Bartonian specimens in the Paris Basin described by MERLE (2005 b) as N. dudariensis differ in morphology; they probably represent a new Nucellopsis species (D. Merle’s personal communication). Concerning another appearance of Nucellopsis in the Alpine Tethys, a specimen was recorded by LORENC (1950: 308, pl. 5, fig. 13) as Muricopsis plicatilis (Deshayes) from the Lutetian – Bartonian of the Vyškov Depression (Czechia). However, the fragmentary specimen differs from N. plicatilis by more slender shell, its determination needs to be revised.	en	Kovács, Zoltán, Vicián, Zoltán (2019): Muricidae (Neogastropoda) from the Middle Eocene of the Hungarian Paleogene Basin. Fragmenta Palaeontologica Hungarica 36: 31-51, DOI: 10.17111/FragmPalHung.2019.36.31, URL: https://doi.org/10.17111/fragmpalhung.2019.36.31
0387A34AFFEFFFB8518B4C7506F7B87E.taxon	distribution	Distribution – Late Lutetian, Alpine Tethys: Hungarian Paleogene Basin: Dudar.	en	Kovács, Zoltán, Vicián, Zoltán (2019): Muricidae (Neogastropoda) from the Middle Eocene of the Hungarian Paleogene Basin. Fragmenta Palaeontologica Hungarica 36: 31-51, DOI: 10.17111/FragmPalHung.2019.36.31, URL: https://doi.org/10.17111/fragmpalhung.2019.36.31
0387A34AFFEFFFB851C64D8A058BBD4F.taxon	type_taxon	Type species: Murex plicatilis Deshayes, 1835	en	Kovács, Zoltán, Vicián, Zoltán (2019): Muricidae (Neogastropoda) from the Middle Eocene of the Hungarian Paleogene Basin. Fragmenta Palaeontologica Hungarica 36: 31-51, DOI: 10.17111/FragmPalHung.2019.36.31, URL: https://doi.org/10.17111/fragmpalhung.2019.36.31
