identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03DD9679FFDBFFA4FF18409A5B5AA43F.text	03DD9679FFDBFFA4FF18409A5B5AA43F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cleidogonidae Cook 1896	<div><p>Family Cleidogonidae Cook, 1896a</p><p>Diagnosis. In North America, chordeumatid millipedes with mentum divided, unlike Conotylidae Cook, 1896a and Adritylidae Shear, 1971 . Anterior gonopods with prominent angiocoxites, sometimes divided into two branches, but always lacking flagella, unlike families in Striariidea Cook, 1896b and Brannerioidea Cook, 1896b. Colpocoxites fused at their bases, unlike Trichopetalidae Verhoeff, 1914 .</p><p>Remarks. The interpretation of cleidogonid gonopods by Shear (1972) seemed correct at the time, but more extensive experience with the family has resulted in changes. The structures previously called colpocoxites (derivatives of the sclerotized coxal glands) are now considered angiocoxites (formed from the rims of coxal glands), and the previous “telopodites” are actually the colpocoxites (Shear 2024). Cleidogonid gonopods have entirely lost telopodites. The ninth legpair, formerly referred to as posterior gonopods, are now best called ninth legs since, though modified, they play no role in spermatophore transfer. Further description of the gonopods, specific to Cleidogona, are in the discussion below.</p></div>	https://treatment.plazi.org/id/03DD9679FFDBFFA4FF18409A5B5AA43F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Snyder, Bruce A.;Shear, William A.	Snyder, Bruce A., Shear, William A. (2025): Four new species of Cleidogona Cook & Collins, 1895 (Diplopoda: Chordeumatida: Cleidogonidae) from the Great Smoky Mountains National Park, USA. Zootaxa 5722 (2): 249-264, DOI: 10.11646/zootaxa.5722.2.5, URL: https://doi.org/10.11646/zootaxa.5722.2.5
03DD9679FFDBFFA5FF1846FE5870A2AB.text	03DD9679FFDBFFA5FF1846FE5870A2AB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cleidogona Cook & Collins 1895	<div><p>Genus Cleidogona Cook &amp; Collins, 1895</p><p>Diagnosis. Cleidogonidae with undivided gonopod angiocoxites, unlike Psuedotremia Cope, 1869 and Solaenogona Hoffman, 1950b . Cleidogona lacks sternal projections anterior to the angiocoxites found in Dybasia Loomis, 1964 . Distinct from Tianella in lacking groups of flagelliform strips on the angiocoxites. Male ninth legs are only slightly reduced in size, consist of 4–6 articles, and femora are enlarged and triangular, unlike Cabraca Shear, 1982, which has a single article, Tiganogona Chamberlin, 1928 which has 2–3 articles, and Lineagona Shear, 2024 which lacks enlarged, triangular prefemora.</p><p>Remarks. Males of Cleidogona, in addition to the gonopods (as described above for the family), have secondary sexual modifications of some of the legs. Legpairs 3–7 are much enlarged, with mesally curved prefemora and femora. These have been observed to clasp the head and anterior segments of the female during mating (pers. obs., WAS). The seventh legpair usually has prominent posterior knobs. These help perhaps in stabilizing the gonopods when retracted. The ninth legpair is variously reduced, but in North American species retains five or six podomeres; the prefemur and coxa are frequently fused. This has led to misinterpretations in some past studies, where the femora are referred to as prefemora. The ninth leg coxae usually have median knobs or processes which interlock with the gonopod colpocoxites at rest, holding the gonopods in the retracted position, and may also carry vestigial basal gland pores. The femora are elongate and distally enlarged, giving the impression of a triangular podomere. Coxae of the tenth and eleventh legpair are elongate and apically bear the openings of coxal glands; frequently the margins of these glands show modifications such as knobs or shelves. A prominent process extends anteriorly from the midline of sternum of the twelfth legs (Shear 1972).</p><p>The gonopods consist of four elements, a sternum, two coxae and paired angiocoxites and colpocoxites. The sternum is broad anteriorly, with a distinct median depression and lateral lobes; posteriorly the sternum is narrow and in some species may be incomplete. The coxae are pyramidal in shape and well separated. They bear from 3 to 12 or more setae and there may or may not be a triangular process extending posteriorly. The angiocoxites are generally the most complex and distinctive of the elements. They extend dorsally to separate the coxae and in most species appear to be bilaterally fused. Distally there are anterior and posterior elements, the anterior being relatively complex and the posterior with a vertical groove containing many fine cuticular fimbriae. The colpocoxites are almost always smaller and less complex than the angiocoxites, but like them are basally fused. They are evenly curved, with the anterior part of the arc usually visible between the angiocoxites. The posterior part is always longer and distally divided into short, blunt, finger-like processes that at rest clasp the tips of the colpocoxites. The coxal knobs of the ninth legs interlock with the basal, posterior part of the posterior arc. The colpocoxites are freely movable through the agency of muscles internal to the gonopods and concealed by the sternum; their attachment to the gonopod complex is membranous.</p><p>Unlike most higher taxa of choredumatidans, the female genitalia of many species of Cleidogona have characters that permit species identification. The anterior receptacle is usually reduced to a simple, rounded lobe. The lateral and medial vulval valves may be entirely separate, or anteriorly fused (as is the case with the four species described herein). The valves frequently bear paired processes which may have distinctive shapes. The vulvae are subtended posteriorly by a postgenital plate that also may be distinctive. The postgenital plate appears to be a development of the second sternum.</p></div>	https://treatment.plazi.org/id/03DD9679FFDBFFA5FF1846FE5870A2AB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Snyder, Bruce A.;Shear, William A.	Snyder, Bruce A., Shear, William A. (2025): Four new species of Cleidogona Cook & Collins, 1895 (Diplopoda: Chordeumatida: Cleidogonidae) from the Great Smoky Mountains National Park, USA. Zootaxa 5722 (2): 249-264, DOI: 10.11646/zootaxa.5722.2.5, URL: https://doi.org/10.11646/zootaxa.5722.2.5
03DD9679FFDAFFA5FF18470A5B11A4FD.text	03DD9679FFDAFFA5FF18470A5B11A4FD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cleidogona jocassee Hoffman 1950	<div><p>Cleidogona jocassee Hoffman, 1950a NEW PARK RECORD</p><p>Cleidogona jocassee Hoffman, 1950a: 19–21, pl. 7, fig. 15–18— Shear (1972: 229, figs 316–319), Snyder (2008: 29)</p><p>Material Examined: NORTH CAROLINA: Haywood Co., Purchase Knob ATBI <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-83.06028&amp;materialsCitation.latitude=35.59194" title="Search Plazi for locations around (long -83.06028/lat 35.59194)">Plot</a>, 35° 35’ 31” N 83° 3’ 37” W: pitfall trap 35, 22 May–8 Jun 2001, 1 male ; pitfall trap 36, 18 Jun–3 Jul 2002, 1 male, 1 female, 1 immature; pitfall trap 40, 23 Apr–15 May 2001, I. C. Stocks, 1 male; pitfall trap 38, 8 May–4 Jun 2002, P. E. Super &amp; J. Lowe, 1 male; pitfall trap 37, 26 Apr–8 May 2002, 1 male; pitfall trap 38, 18 Jun–3 Jul 2002, J. &amp; D. Lowe, 2 males.</p><p>Remarks. Cleidogona jocassee was recorded as likely to be in GSMNP by Snyder (2008) since it was known from Haywood and Swain Cos., North Carolina, but outside the GSMNP boundary (Shear 1972). It is now documented from Purchase Knob.</p></div>	https://treatment.plazi.org/id/03DD9679FFDAFFA5FF18470A5B11A4FD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Snyder, Bruce A.;Shear, William A.	Snyder, Bruce A., Shear, William A. (2025): Four new species of Cleidogona Cook & Collins, 1895 (Diplopoda: Chordeumatida: Cleidogonidae) from the Great Smoky Mountains National Park, USA. Zootaxa 5722 (2): 249-264, DOI: 10.11646/zootaxa.5722.2.5, URL: https://doi.org/10.11646/zootaxa.5722.2.5
03DD9679FFDAFFA3FF1845395C7DA477.text	03DD9679FFDAFFA3FF1845395C7DA477.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cleidogona chilhoweensis Snyder & Shear 2025	<div><p>Cleidogona chilhoweensis Snyder &amp; Shear, sp. nov.</p><p>urn:lsid:zoobank.org:act: 63A63F36-124E-473A-96F9-07926EE1C77A</p><p>Figs 1–8</p><p>Material Examined. Types: Holotype: 1 male, TENNESSEE: Blount Co., along US Highway 129 adjacent to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-83.99167&amp;materialsCitation.latitude=35.55" title="Search Plazi for locations around (long -83.99167/lat 35.55)">Lake Chilhowee</a>, Plot C +15, 35° 33.0’ N 83° 59.5’ W, 269–291 m elevation, hand collection, 14 Oct 2006, B. A. Snyder et al. (USNMENT02180525) . Paratypes: 1 male and 1 female, TENNESSEE: Blount Co., along US Highway 129 adjacent to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-83.9975&amp;materialsCitation.latitude=35.555553" title="Search Plazi for locations around (long -83.9975/lat 35.555553)">Lake Chilhowee</a>, site 3, near 35° 33’ 20” N 83° 59’ 51” W, hand collection, 27 Oct 2004, B. A. Snyder (GCIZ 9000) ; 1 male, same collection (GRSM 228785) .</p><p>Non-types: TENNESSEE: Blount Co., along US Highway 129 adjacent to Lake Chilhowee, site 1, 5m from roadside, hand collection, 27 Oct 2004, B. A. Snyder, 1 male. Blount Co., along US Highway 129 adjacent to Lake Chilhowee, hand collection, 13 Apr 2007, B. A. Snyder, 1 female. Both specimens used for SEM .</p><p>Diagnosis. Distinct from other Cleidogona in the details of the gonopods. Cleidogona chilhoweensis sp. nov. is most similar to and co-occurs with C. nantahala; both lack a coxal process. The colpocoxite of C. nantahala has a small mid-length flange and lacks a large basal flange and distal thumb-like projections.</p><p>Etymology. The species name refers to the type locality, the westernmost area of GSMNP adjacent to Lake Chilhowee, an impoundment of the Little Tennessee River formed in the 1950s. The area was previously the Cherokee settlement of Chilhowee. Adjective.</p><p>Description. Length of holotype 14.5 mm, greatest width 1.6 mm at segment 7. Ommatidia 26 (left side) or 27 (right side) in 7 rows. Color faded in preserved specimens, but typical of the genus in life. Segments cylindrical. Somatic characters and secondary sexual modifications typical for the genus (see above; also Shear 1972).</p><p>Gonopods (Figs 1–3): In anterior view (Fig. 1), sternum (s) narrow and bandlike medially, expanded below coxae. Coxae (cx) elongate but mostly obscured by angiocoxites (ac) (Figs 1–3). Angiocoxites robust, smooth, expanded distally into shield-like anterior branches (ab) with sinuous distal margins. Posterior branches (pb) extending above concave sections of these margins. Angiocoxites narrowing basally, extending down to sternum. Laterally flattened flanges of colpocoxites (cc) visible between angiocoxites.</p><p>In lateral view (Fig. 3), coxa (cx) elongate, subtriangular, lacking a coxal process, with a small tooth on the posterior edge towards the base. Angiocoxite (ac) upright, with a gentle curve anterior to the coxa but continuing straight distally.Angiocoxite apically divided, the anterior branch (ab) shield-like, wrapping laterally to obscure the view of the division. The posterior (inner) branch (pb) begins as a flange and is only separated distally (this is best seen in posterior view, Fig. 2). The apex is blade-shaped with a slightly curved tip.</p><p>Proximally, the colpocoxite (cc) makes a sharp 90° bend dorsad and then curves back ventrad almost 90°. Proximal bend with a large flange extending anteriad and ventrad, and a corner pointed towards the angiocoxite. Distal tip of the colpocoxite bluntly rounded with a thumb-like process on the lateral side which also points towards the angiocoxite.</p><p>Ninth legs (Fig. 4) typical for the genus, coxae and prefemora fused, with constriction at one third the length from the base. Femora (f9) triangular, subequal in length to the coxae. Tenth legs (Fig. 5) typical for the genus, sternum (s10) expanded into a tooth at the base of each leg and a very low medial ridge, coxae (cx10) elongate, slightly curved, with prominent gland at base (cg).</p><p>Females similar to males in all nonsexual aspects. Postgenital plate (pgp), (Fig. 6) not divided, upright, distally expanded laterad with a small medial apical notch. Vulvae (Figs 7, 8) with an inner valve (iv), outer valve (ov) and receptacle (r). Outer valves with paired projections on the posterior edge, medial projections (mp) longer than lateral projections (lp) (right medial projection broken in Figs 7, 8).</p><p>Distribution. Only known from the type locality (Fig. 29; full site description in Snyder et al. 2011).</p><p>Remarks. The type locality, on the western edge of GSMNP, is the warmest and driest location in the park.</p></div>	https://treatment.plazi.org/id/03DD9679FFDAFFA3FF1845395C7DA477	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Snyder, Bruce A.;Shear, William A.	Snyder, Bruce A., Shear, William A. (2025): Four new species of Cleidogona Cook & Collins, 1895 (Diplopoda: Chordeumatida: Cleidogonidae) from the Great Smoky Mountains National Park, USA. Zootaxa 5722 (2): 249-264, DOI: 10.11646/zootaxa.5722.2.5, URL: https://doi.org/10.11646/zootaxa.5722.2.5
03DD9679FFDCFFAEFF1846B65879A4DB.text	03DD9679FFDCFFAEFF1846B65879A4DB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cleidogona columbicauda Snyder & Shear 2025	<div><p>Cleidogona columbicauda Snyder &amp; Shear, sp. nov.</p><p>urn:lsid:zoobank.org:act: 2AA08882-F203-4EB6-8C24-F126CEB2EB26</p><p>Figs 9–15</p><p>Material Examined. Types: Holotype: 1 male, TENNESSEE: Cocke Co., Snakeden Ridge ATBI <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-83.22&amp;materialsCitation.latitude=35.743332" title="Search Plazi for locations around (long -83.22/lat 35.743332)">Plot</a>, 35° 44’ 36” N 83° 13’ 12” W, pitfall trap 91, 10 Mar–6 Apr 2002 (USNMENT02180526) . Paratypes: 1 male, 2 females, TENNESSEE: Cocke Co., Snakeden Ridge ATBI <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-83.22&amp;materialsCitation.latitude=35.743332" title="Search Plazi for locations around (long -83.22/lat 35.743332)">Plot</a>, 35° 44’ 36” N 83° 13’ 12” W, pitfall trap 95, 1–13 Dec 2000 (GCIZ 9001) . 1 male, 1 female, TENNESSEE: Cocke Co., Snakeden Ridge ATBI <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-83.22&amp;materialsCitation.latitude=35.743332" title="Search Plazi for locations around (long -83.22/lat 35.743332)">Plot</a>, 35° 44’ 36” N 83° 13’ 12” W, pitfall trap 91, 16 Jan–13 Feb 2002, (VMNH112502.1, VMNH112502.2) . 1 male, 1 female, TENNESSEE: Cocke Co., Snakeden Ridge ATBI <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-83.22&amp;materialsCitation.latitude=35.743332" title="Search Plazi for locations around (long -83.22/lat 35.743332)">Plot</a>, 35° 44’ 36” N 83° 13’ 12” W, pitfall trap 93, 6 May–5 Jun 2002 (GRSM 228786) .</p><p>Non-types: TENNESSEE: Blount Co., Cades Cove ATBI <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-83.8375&amp;materialsCitation.latitude=35.59139" title="Search Plazi for locations around (long -83.8375/lat 35.59139)">Plot</a>, 35° 35’ 29” N 83° 50’ 15” W: pitfall trap 18, 2–16 Jul 2001, 1 male, 1 immature . Sevier Co., Goshen Prong ATBI <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-83.54278&amp;materialsCitation.latitude=35.610554" title="Search Plazi for locations around (long -83.54278/lat 35.610554)">Plot</a>, 35° 36’ 38” N 83° 32’ 34” W: pitfall trap 101, 19–26 Feb 2001, 1 male, 1 female ; pitfall trap 103, 10 Feb–7 Mar 2002, 1 male; pitfall trap 104, 17 Sep–22 Oct 2001, 1 male. Indian Gap ATBI <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-83.44361&amp;materialsCitation.latitude=35.610832" title="Search Plazi for locations around (long -83.44361/lat 35.610832)">Plot</a>, 35° 36’ 39” N 83° 26’ 37” W: pitfall trap 23, 10 May–19 Jun 2002, 1 male ; pitfall trap 25, 15–28 Mar 2002, 1 male; pitfall trap 25, 4 Apr–10 May 2003, 1 male (used for SEM); pitfall trap 27, 14–27 Apr 2002, 1 male; pitfall trap 27, 16–28 Mar 2002, 1 male (used for SEM); pitfall trap 28, 4 Apr–10 May 2003, 1 male. Twin Creeks ATBI <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-83.498886&amp;materialsCitation.latitude=35.685" title="Search Plazi for locations around (long -83.498886/lat 35.685)">Plot</a>, 35° 41’ 6” N 83° 29’ 56” W: pitfall trap 10, 27 Mar–11 Apr 2001, 1 male . Under bark of rotten logs, old growth forest nr. Rocky Spur Branch, 963m, 8 Apr 1995, C. R. Parker, 1 male, 2 females . Brushy Mountain ATBI <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-83.43083&amp;materialsCitation.latitude=35.676666" title="Search Plazi for locations around (long -83.43083/lat 35.676666)">Plot</a>, 35° 40’ 36” N 83° 25’ 51” W: pitfall trap 61, 6 Apr–23 Jun 2003, 1 male, 1 immature ; pitfall trap 61, 27 Feb–12 Mar 2003, 1 male; pitfall trap 63, 5–21 Jul 2001, 1 male, 1 immature; pitfall trap 63, 6 Apr–23 Jun 2003, 1 male; pitfall trap 63, 23 May–5 Jun 2001, 1 male; pitfall trap 64, 7–19 Dec 2001, 1 male; pitfall trap 64, 5–21 Jun 2001, 1 male, 1 female; pitfall trap 64, 30 Apr–12 May 2002, 1 male; pitfall trap 64, 5–21 Jul 2001, 1 male; pitfall trap 64, 13–27 Mar 2002, 1 male; pitfall trap 64, 18–30 Jun 2002, 1 male; pitfall trap 65, 13–27 Mar 2002, 1 male; pitfall trap 66, 12 Dec 2000 – 16 Jan 2001, 1 male; pitfall trap 66, 6 Apr–23 Jun 2003, 1 male; pitfall trap 68, 11–26 Apr 2001, 1 male; pitfall trap 69, 6 Apr–23 Jun 2003, 1 male; pitfall trap 70, 25 Apr–23 May 2003, M. J. Petersen, S. Higdon &amp; C. R. Parker, 1 male, 1 female (used for SEM); pitfall trap 70, 15 Jan–15 Feb 2002, 1 male. Cocke Co., Albright Grove ATBI <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-83.280556&amp;materialsCitation.latitude=35.733334" title="Search Plazi for locations around (long -83.280556/lat 35.733334)">Plot</a>, 35° 43’ 60” N 83° 16’ 50” W: pitfall trap 81, 9 Mar–4 May 2003, 1 male ; pitfall trap 82, 9 Mar–4 May 2003, 1 male, 1 female; pitfall trap 83, 8 Feb–9 Mar 2003, 1 male; pitfall trap 84, 8 Feb–9 Mar 2003, 2 males; pitfall trap 84, 9 Mar–5 Apr 2003, 1 male; pitfall trap 86, 14–29 Mar 2002, 1 male; pitfall trap 87, 21 Jul–1 Aug 2001, 1 male; pitfall trap 87, 9 Mar–5 Apr 2003, 1 male; pitfall trap 89, 12 May–15 Jun 2002, 1 male, 1 immature. Snakeden Ridge ATBI <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-83.22&amp;materialsCitation.latitude=35.743332" title="Search Plazi for locations around (long -83.22/lat 35.743332)">Plot</a>, 35° 44’ 36” N 83° 13’ 12” W: pitfall trap 91, 5–16 Jan 2002, 1 male ; pitfall trap 91, 3 Mar–30 Apr 2003, 1 male, 1 female; pitfall trap 91, 6 Apr–6 May 2002, 1 male; pitfall trap 91, 19 Nov–4 Dec 2001, 2 males; pitfall trap 91, 6–19 Nov 2001, 1 male; pitfall trap 92, 30 Jan–15 Feb 2001, 1 male (used for SEM); pitfall trap 94, 25 Apr–9 May 2001, 1 male, 1 immature; pitfall trap 94, 3–25 Apr 2001, I. C. Stocks &amp; J. Breeden, 1 male; pitfall trap 94, 6 Apr–6 May 2002, 1 male; pitfall trap 95, 4–21 Dec 2001, 1 male; pitfall trap 95, 22 May–4 Jun 2001, 1 male; pitfall trap 95, 17–30 Jan 2001, 1 male; pitfall trap 97, 10 Mar–6 Apr 2002, 2 males; pitfall trap 97, 16 Feb–5 Mar 2001, 2 males; pitfall trap 98, 15 Mar–3 Apr 2001, I. C. Stocks &amp; J. Breeden, 1 male, 2 females; pitfall trap 98, 3–25 Apr 2001, 1 male, 1 immature; pitfall trap 98, 16 Jan–13 Feb 2002, 1 male; pitfall trap 98, 13 Dec 2000 – 7 Jan 2001, 1 male; pitfall trap 99, 15 Mar–3 Apr 2001, I. C. Stocks &amp; J. Breeden, 2 males; pitfall trap 99, 4–21 Dec 2001, 1 male; pitfall trap 99, 18 Feb–3 Mar 2003, 1 male; pitfall trap 100, 10 Mar–6 Apr 2002, 1 male; pitfall trap 100, 3 Mar–30 Apr 2003, 1 male; Malaise trap 20, 16 Feb–5 Mar 2001, 1 male.</p><p>NORTH CAROLINA: Swain Co., Kuwohi (Clingmans Dome) ATBI <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-83.49528&amp;materialsCitation.latitude=35.560276" title="Search Plazi for locations around (long -83.49528/lat 35.560276)">Plot</a>, 35° 33’ 37” N 83° 29’ 43” W: pitfall trap 56, 18 Oct–13 Nov 2000, 1 male ; pitfall trap 71, 8–24 Nov 2001, 2 males; pitfall trap 76, 24 Oct–8 Nov 2001, 1 male, 1 female; pitfall trap 77, 6–25 Jun 2001, 1 male; pitfall trap 79, 27 Apr–10 May 2002, 1 male; pitfall trap 79, 25 Jun–3 Jul 2001, 1 male. Andrews Bald ATBI <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-83.49416&amp;materialsCitation.latitude=35.538887" title="Search Plazi for locations around (long -83.49416/lat 35.538887)">Plot</a>, 35° 32’ 20” N 83° 29’ 39” W: pitfall trap 54, 28 Mar–14 Apr 2002, 1 male, 1 female ; pitfall trap 54, 24 27 Apr–10 May 2002, 1 male (used for SEM); pitfall trap 54, 24 Oct–8 Nov 2001, 1 male; pitfall trap 55, 18 Jul–1 Aug 2002, 1 male; pitfall trap 56, 13–29 Nov 2000, 1 male; pitfall trap 56, 16 Mar–24 Apr 2001, 1 male; pitfall trap 57, 1 Feb–16 Mar 2001, 2 males; pitfall trap 57, 16 Mar–24 Apr 2001, 1 male; pitfall trap 58, 20 Oct–13 Nov 2000, 1 male, 1 female; pitfall trap 58, 28 Mar–14 Apr 2002, 1 male; pitfall trap 60, 24 Mar–16 Apr 2001, 1 male; pitfall trap 60, 24 Oct–8 Nov 2001, 1 male; 9721, App Tr 350m E Road Prong Trailhead, hog exclosure, 5400’ elevation, beech gap forest, 1m 2 litter #2, 10 Jun 1997, Coyle, Stocks, Aiken, Davis, 1 male .</p><p>Diagnosis. Cleidogona columbicauda sp. nov. is most similar to C. medialis in the overall shape of the gonopods, but they differ in the shape of the angiocoxite processes. Cleidogona medialis has medially directed processes which are separated from the angiocoxite apices, whereas C. columbicauda sp. nov. has lamellae that point mediad proximally and continue distad on the angiocoxite apices.Additional differences include the coxae of the male tenth legs (lateral process in C. columbicauda sp. nov., medial process in C. medialis) and the female postgenital plate (low, undivided in C. columbicauda sp. nov., divided in C. medialis).</p><p>Etymology. The species name refers to the interleaving of the distal ends of the angiocoxites and colpocoxites, which is reminiscent of the close fit of a dovetail joint in woodworking. The name is a combination of the Latin for dove (columba) and the Latin for tail (cauda). Noun in apposition.</p><p>Description. Length of holotype 11.9 mm, greatest width 1.3 mm at segment 7. Ommatidia 26 in 7 rows. Color faded in holotype and most preserved specimens, but typical of the genus in some specimens. Segments cylindrical, with low shoulders. Somatic characters and secondary sexual modifications are typical for the genus (see above; also Shear 1972).</p><p>Gonopods (Figs 9–11): In anterior view (Fig. 9), sternum (s) a narrow band, expanded laterally. Coxae (cx) subtriangular. Angiocoxites (ac) linear, stout proximally, with sinuous edges. Middle of proximal region wrinkled. Distally, angiocoxites curve posteriad, narrow, and bear a flange on their medial surfaces. Each angiocoxite ends in a broad, rugose, medially curving lamella that wraps around the distal end of the colpocoxite; the distal colpocoxite process (dcp) is lateral to the lamellae. The edges of the lamellae are toothed and a small hook (h) is present distally. A process (p) arises between the angiocoxites, variable in length (see remarks). The proximal ends of the colpocoxites (cc) are visible on either side of this process.</p><p>In lateral view (Fig. 11), coxae (cx) elongate, lacking a coxal process, base somewhat expanded laterad. Angiocoxite (ac) gently curving posteriad, wrinkled proximally, small dorsal notch just distal to coxa, expanding distally into a large, posteriodorsally directed laciniate lobe (ll) and a thin, ventrally located lamella (vl). The lamella curves medially, with a toothed edge and a distal hook (h). The distal end of the angiocoxite almost completely covers the distal end of the colpocoxite (cc), except for the distal colpocoxite process (dcp) that is lateral to the angiocoxite. The colpocoxite is curved with two large ridges. The inner ridge continues between the middle of the angiocoxites and becomes a laterally flattened, bluntly pointed proximal process. The outer ridge ends in a large triangular tooth (dcp). The main stem of the colpocoxite is greatly expanded distally and nests medial to the angiocoxite lobe and flange.</p><p>Ninth legs (Fig. 12) typical for the genus, sternum with a small medial bump, fused prefemora and coxae sinuous, vestigial glands produced on knobs. Femora (f9) triangular, slightly longer than the fused prefemora and coxae. Tenth legs (Fig. 13) typical for the genus, sternum (s10) expanded into a tooth medially at the base of each leg and a pronounced medial ridge. Coxal glands (cg) produced on knobs giving the appearance of a short branch. Coxae (cx10) with distal lateral knobs and an abrupt widening at the distal ends.</p><p>Females similar to males in all nonsexual aspects. Postgenital plate (pgp) not divided, low, expanded ventrally into three weakly divided lobes (Figs 14, 15). Vulvae with an inner valve (iv), outer valve (ov), and paired projections on the posterior edge (Figs 14, 15). Medial projections (mp) pointed, longer; lateral projections (lp) narrow distally before widening into two teeth.</p><p>Distribution. Cleidogona columbicauda sp. nov. is widely distributed in GSMNP, having been collected at 9 of 19 ATBI plots and a few additional sites (Fig. 29). Its range stretches from Cades Cove east to Snakeden Ridge, and south to Andrews Bald, with elevations from 522 m at Cades Cove to 1954 m at Kuwohi (formerly Clingmans Dome). It appears to be more common at higher elevation sites; 94% of specimens collected were from sites above 950 m, and 45% over 1450 m. Habitats include open and forested sites: old field, heath bald, grassy bald; cove forests (second-growth and old-growth), hemlock forest, high-elevation beech forest, and spruce-fir forest.</p><p>Remarks. The process between the angiocoxites is variable in length. In most specimens it is prominent, extending about halfway to the lamellae, but it some it is short, and in others it is lacking. Specimens with the process exist at all sites where C. columbicauda sp. nov. has been collected, but specimens lacking the process cooccur at all high-elevation sites (above 1450 m). Specimens with and without the process have even been found in the same pitfall trap.</p></div>	https://treatment.plazi.org/id/03DD9679FFDCFFAEFF1846B65879A4DB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Snyder, Bruce A.;Shear, William A.	Snyder, Bruce A., Shear, William A. (2025): Four new species of Cleidogona Cook & Collins, 1895 (Diplopoda: Chordeumatida: Cleidogonidae) from the Great Smoky Mountains National Park, USA. Zootaxa 5722 (2): 249-264, DOI: 10.11646/zootaxa.5722.2.5, URL: https://doi.org/10.11646/zootaxa.5722.2.5
03DD9679FFD1FFACFF1846DA5A8CA51F.text	03DD9679FFD1FFACFF1846DA5A8CA51F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cleidogona hendrixi Snyder & Shear 2025	<div><p>Cleidogona hendrixi Snyder &amp; Shear, sp. nov.</p><p>urn:lsid:zoobank.org:act: DEAB45DD-F2D4-4409-B327-85FE8FC0A135</p><p>Figs 16–20</p><p>Material Examined. Types: Holotype: 1 male, NORTH CAROLINA: Haywood Co., Purchase Knob ATBI <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-83.06028&amp;materialsCitation.latitude=35.59194" title="Search Plazi for locations around (long -83.06028/lat 35.59194)">Plot</a>, 35° 35’ 31” N 83° 3’ 37” W, pitfall trap 35, 7–21 Dec 2001 (USNMENT02180527) . Paratypes: 1 male, same location, pitfall trap 35, 9–23 Nov 2002 (GCIZ 9002); 1 male, same location, pitfall trap 37, 27 Apr–15 May 2001 (GRSM 228787) .</p><p>Non-types: In addition to the types, 1 male was collected from pitfall trap 35 on each of the following dates: 2 Oct–9 Nov 2002 (used for SEM), 9–23 Nov 2002, and 15 Mar–10 May 2003 (used for SEM).</p><p>Diagnosis. Cleidogona hendrixi sp. nov. is unique in that no other Cleidogona has both dorsal and ventral lamellae on the angiocoxites of the gonopods. Several Mexican Cleidogona species (e.g., C. tequila Shear 1972 and C. totonaca Shear 1972) have generally similar curved, rod-like angiocoxites with dorsal lamellae, but these all lack ventral lamellae and the colpocoxites are not reflexed.</p><p>Etymology. This species is named for Dr. Paul F. Hendrix, the first author’s PhD advisor, who initially sent him to the Smokies, setting him on a path to describe these species. Noun in the genitive case.</p><p>Description. Length of holotype 15.1 mm, greatest width 1.8 mm at segment 7. Ommatidia 28 in 7 rows. Color faded in preserved specimens except for head, antennae, and tarsi, which are brown. Segments cylindrical. Somatic characters and secondary sexual modifications are typical for the genus (see above; also Shear 1972).</p><p>Gonopods (Figs 16–18): In anterior view (Fig. 17), sternum broad and flat. Coxae (cx) almost entirely obscured by angiocoxites. Angiocoxites (ac) curve strongly posteriorly (Figs 16, 17). They are widely spaced and parallel (or slightly converging) proximally and then gently curve apart (laterad) distally, while gradually narrowing. A thin mohawk-like ventral lamella is present for the distal 1/2 to 2/3 of the length. The entirety of the colpocoxites (cc) are visible between angiocoxites, extending posteriad and reflexing towards the base of the angiocoxites.</p><p>In lateral view (Fig. 18), coxa (cx) elongate, straight-sided where it meets the angiocoxite (ac), squared on the posterior corner, and rounded dorsally. Proximally, the angiocoxite broadens into an elongate shape that has a small bump adjacent to the rod-like distal portion but otherwise mirrors the shape of the coxa. The coxa and angiocoxite proximal area together form an almost symmetrical shield-like shape. The angiocoxite is rod-like distally, curving at midlength and tapering to a blunt point. Lamellae are present on both the ventral (vl) and dorsal (dl) sides; the ventral lamella is visible from the curve of the angiocoxite to just before the tip, where it is tallest (subequal to the width of the narrowing angiocoxite). The ventral lamella originates somewhat more proximally and on the ventromedial face of the angiocoxite. The dorsal lamella originates at the proximal end of the rod-like portion of the angiocoxite and is continuous to the distal end. The lamella is somewhat shorter at the bend of the angiocoxite and also very short in a distal region, which aligns with where the colpocoxite (cc) would contact the angiocoxite if flexed. The lamella is so small here that it appears to be absent in some specimens. Proximally, the colpocoxite (cc) curves posteriad and reflexes 180° to point anteriad, ending parallel and just dorsal of the angiocoxite. The distal tip of the colpocoxite is broadly expanded with a lateral thumb-like process, also directed anteriad.</p><p>Ninth legs (Fig. 19) typical for the genus, coxae with pores on large knobs, slightly constricted above, and bean-shaped distally. Femora (f9) subequal in length to the coxae, triangular, with a small knob on the ventral corner. Tenth legs (Fig. 20) typical for the genus, sternum (s10) expanded into a tooth at the base of each leg and a pronounced medial ridge. Coxal glands (cg) produced on knobs. Coxae (cx10) sublinear, widening distally.</p><p>Females unknown.</p><p>Distribution. Only known from the type locality, the Purchase Knob ATBI plot (Fig. 29), a second-growth southern Appalachian cove forest at 1346 m elevation (Parker &amp; Bernard 2006).</p><p>Remarks. Three Cleidogona are found at Purchase Knob: C. hendrixi sp. nov., C. jocassee, and C. margarita . The latter two species are described and illustrated in Shear (1972).</p></div>	https://treatment.plazi.org/id/03DD9679FFD1FFACFF1846DA5A8CA51F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Snyder, Bruce A.;Shear, William A.	Snyder, Bruce A., Shear, William A. (2025): Four new species of Cleidogona Cook & Collins, 1895 (Diplopoda: Chordeumatida: Cleidogonidae) from the Great Smoky Mountains National Park, USA. Zootaxa 5722 (2): 249-264, DOI: 10.11646/zootaxa.5722.2.5, URL: https://doi.org/10.11646/zootaxa.5722.2.5
03DD9679FFD3FFAAFF18479E5ADEA717.text	03DD9679FFD3FFAAFF18479E5ADEA717.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cleidogona sparta Snyder & Shear 2025	<div><p>Cleidogona sparta Snyder &amp; Shear, sp. nov.</p><p>urn:lsid:zoobank.org:act: 91FA18B2-69A1-4CBD-92AC-C2E6C5C19773</p><p>Figs 21–28</p><p>Material Examined. Types: Holotype: 1 male, TENNESSEE: Sevier Co., Twin Creeks ATBI <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-83.498886&amp;materialsCitation.latitude=35.685" title="Search Plazi for locations around (long -83.498886/lat 35.685)">Plot</a>, 35° 41’ 6” N 83° 29’ 56” W, pitfall trap 10, 16–31 Jul 2002 (USNMENT02180528) . Paratypes: 1 male, 3 females, TENNESSEE: Sevier Co., Twin Creeks ATBI <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-83.498886&amp;materialsCitation.latitude=35.685" title="Search Plazi for locations around (long -83.498886/lat 35.685)">Plot</a>, 35° 41’ 6” N 83° 29’ 56” W, pitfall trap 06, 6–21 June 2001 (GCIZ 9003) . 1 male, TENNESSEE: Sevier Co., Twin Creeks ATBI <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-83.498886&amp;materialsCitation.latitude=35.685" title="Search Plazi for locations around (long -83.498886/lat 35.685)">Plot</a>, 35° 41’ 6” N 83° 29’ 56” W, pitfall trap 07, 30 Jul–13 Aug 2001 (VMNH112503) . 1 male, 1 female, TENNESSEE: Sevier Co., Twin Creeks ATBI <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-83.498886&amp;materialsCitation.latitude=35.685" title="Search Plazi for locations around (long -83.498886/lat 35.685)">Plot</a>, 35° 41’ 6” N 83° 29’ 56” W, pitfall trap 03, 15 Aug–10 Sep 2002 (GRSM 228788) .</p><p>Non-types: TENNESSEE: Sevier Co., Goshen Prong ATBI <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-83.54278&amp;materialsCitation.latitude=35.610554" title="Search Plazi for locations around (long -83.54278/lat 35.610554)">Plot</a>, 35° 36’ 38” N 83° 32’ 34” W: pitfall trap 101, 17–30 July 2001, 1 male, 2 females ; pitfall trap 103, 17 July–5 Aug 2002, 2 males (1 used for SEM); pitfall trap 104, 18 Jun–2 Jul 2001, 1 male, 1 immature; pitfall trap 104, 17 July–5 Aug 2002, 1 male, 1 immature; pitfall trap 106, 13–27 Aug 2001, I. C. Stocks &amp; M. Williams, 1 male; pitfall trap 106, 17 July–5 Aug 2002, 1 male, 1 immature; pitfall trap 108, 2–17 July 2001, 1 male. Twin Creeks ATBI <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-83.498886&amp;materialsCitation.latitude=35.685" title="Search Plazi for locations around (long -83.498886/lat 35.685)">Plot</a>, 35° 41’ 6” N 83° 29’ 56” W: pitfall trap 02, 16–31 July 2002, 2 males ; pitfall trap 03, 6–16 July 2001, 1 male; pitfall trap 03, 16–31 Jul 2002, 1 male; pitfall trap 03, 26 Aug–10 Sep 2002, 1 male (used for SEM); pitfall trap 05, 15 Aug–10 Sep 2002, 1 male; pitfall trap 06, 6–21 June 2001, 1 female (used for SEM); pitfall trap 07, 5–21 June 2001, 1 male, 1 immature. Porter’s Flat, 9731 #1, hardwood cove forest, 737.6 m, 31 July 1997, Aiken / Coyle / Davis / Stocks, 1 male, 4 females . Off Cherokee Orchard Road, near Twin Creeks Science Center, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-83.50361&amp;materialsCitation.latitude=35.68722" title="Search Plazi for locations around (long -83.50361/lat 35.68722)">Baer</a> 4 pitfall, 35° 41’ 14” N 83° 30’ 13” W, 22 Jul 2010, D. Straube et al., 1 male . Off Cherokee Orchard Road, near Old Sugarlands Trailhead, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-83.49139&amp;materialsCitation.latitude=35.674446" title="Search Plazi for locations around (long -83.49139/lat 35.674446)">Old</a> trail 1 pitfall, 35° 40’ 28” N 83° 29’ 29” W, 20 Aug 2010, D. Straube et al., 1 male (used for SEM) . Near riverview quiet walkway east of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-83.52151&amp;materialsCitation.latitude=35.66367" title="Search Plazi for locations around (long -83.52151/lat 35.66367)">Huskey Gap Trailhead</a>, 35° 39’ 49.2” N, 83° 31’17.4” W, post-fire study plot UB2 litter, hand collected, 12 July 2018, C. M. Hawley, G. Chapman, M. A. Callaham Jr., M. K. Taylor, 1 male . Near riverview quiet walkway east of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-83.52151&amp;materialsCitation.latitude=35.66367" title="Search Plazi for locations around (long -83.52151/lat 35.66367)">Huskey Gap Trailhead</a>, 35° 39’ 49.2” N, 83° 31’17.4” W, post-fire study plot BP4 litter, hand collected, 12 July 2018, C. M. Hawley, G. Chapman, M. A. Callaham Jr., M. K. Taylor, 1 male . Cocke Co., Albright Grove ATBI <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-83.280556&amp;materialsCitation.latitude=35.733334" title="Search Plazi for locations around (long -83.280556/lat 35.733334)">Plot</a>, 35° 43’ 60” N 83° 16’ 50” W, pitfall trap 89, 6–12 May 2002, 1 male (used for SEM).</p><p>Diagnosis. The gonopods of Cleidogona sparta sp. nov. are similar in overall shape to those of C. major Cook &amp; Collins 1895, but have several distinctive differences. The colpocoxites differ, with C. sparta sp. nov. having a large, distinct tooth. The anterior angiocoxite branch is shieldlike in C. sparta sp. nov. but not in C. major . In C. sparta sp. nov., the coxal process is smaller and rounded, and the transition from coxal process to the posterior edge of angiocoxite is a smooth curve; in C. major the coxal process is longer and squared off, and is demarcated from the angiocoxite by a sharp angle. The shapes of the posterior angiocoxite branch differ greatly in lateral view, with C. sparta sp. nov. being much broader than C. major . The female postgential plate differs between C. sparta sp. nov. and C. major as well. The shape of the C. sparta sp. nov. posterior angiocoxite branch in lateral view is similar to that of C. atropos Shear 1972, however the C. atropos angiocoxite branches are greatly different lengths, rather than equal in C. sparta sp. nov. Cleidogona jocassee and C. margarita are also similar and geographically proximate, but C. jocassee lacks a coxal process and C. margarita has a large, quadrate coxal process.</p><p>Etymology. The species name refers to the lateral view of the colpocoxite apices, which are reminiscent of the profile of a Spartan/Trojan helmet with forward pointing plume. Noun in apposition.</p><p>Description. Length of holotype 24.9 mm, greatest width 2.2 mm at segment 7. Ommatidia 28 in 7 rows. Color faded in preserved specimens, except on the epicranium and frons. Segments cylindrical, with low shoulders. Somatic characters and secondary sexual modifications are typical for the genus (see above; also Shear 1972).</p><p>Gonopods (Figs 21–23): In anterior view (Fig. 21), sternum (s) broad, widest laterally. Coxae (cx) elongate, triangular. Angiocoxites (ac) close together, broad, robust, narrowing distally before dividing into two apical branches of equal length. Branches are about 1/4 the length of the angiocoxite. The anterior branch (ab) arises from the medial half of angiocoxite and has blunt apices, variably squared off or slightly rounded. The posterior branch (pb) arises laterally and the apices are almost squared off with a medial subapical notch. In anterior view, the anterior branch is wider than, medial to, and does not conceal the posterior branch. In some specimens, the anterior branch slightly covers part of the posterior branch. In posterior view (Fig. 22), colpocoxites (cc) appear lobe-like, somewhat flattened. They diverge laterally, forming a V shape.</p><p>In lateral view (Fig. 23), coxa (cx) elongate with a coxal process (cp) at the apex, ending in a blunt point. The space between the coxal process and the angiocoxite (ac) is a rounded notch. Anterior edge of angiocoxite (ac) gently curving posteriad until division. Anterior angiocoxite branch (ab) is shieldlike, anteriomedial to the posterior branch. Posterior edge of angiocoxite has a small tooth (t) just distal to notch above the coxal process. Posterior angiocoxite branch (pb) expands posteriad and narrows into an acuminate point; posterior edge is fringed in this region. Colpocoxite (cc) linear, rod-like proximally and expanded apically. The posterior edge continues in a semicircle through the distal edge, terminating in an anteriorly directed tooth as distal as the division between angiocoxite branches. The anterior side of the colpocoxite has a flange-like expansion on the distal half, ending at the base of the tooth..</p><p>Ninth legs (Fig. 24) typical for the genus, coxae with pores on wide knob-like projections, sublinear distally. Femora (f9) longer than coxae, expanded distally. Tenth legs (Fig. 25) typical for the genus, sternum with a low medial ridge. Coxae (cx10) wide proximally at gland (cg) locations, and abruptly widening at the distal ends.</p><p>Females similar to males in all nonsexual aspects. Postgenital plate (pgp) fused basally but otherwise divided into two projections, each with two branches that curve slightly towards each other, resembling a thumb and forefinger (Fig. 26). Lateral branches, shorter, wider, and textured with small bumps (Fig. 27). Vulvae do not have any projections on the posterior edge (Figs. 26, 28).</p><p>Distribution. Known from the eastern half of the Tennessee (northern) side of GSMNP, from Goshen Prong to Albright Grove (Fig. 29). Habitats include Southern Appalachian cove forests: second-growth, rich montane type (Goshen Prong); second-growth, typic montane type (Twin Creeks); and old-growth, Silverbell type (Albright Grove); elevations range from 617–1051 m (Parker &amp; Bernard 2006).</p><p>Remarks. Although having affinity with C. major, these species are geographically separated by what appears to be a long distance as most Cleidogona species are known to be short-range endemics. The closest published record of C. major is approximately 120 km to the northeast (Shear 1972).</p></div>	https://treatment.plazi.org/id/03DD9679FFD3FFAAFF18479E5ADEA717	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Snyder, Bruce A.;Shear, William A.	Snyder, Bruce A., Shear, William A. (2025): Four new species of Cleidogona Cook & Collins, 1895 (Diplopoda: Chordeumatida: Cleidogonidae) from the Great Smoky Mountains National Park, USA. Zootaxa 5722 (2): 249-264, DOI: 10.11646/zootaxa.5722.2.5, URL: https://doi.org/10.11646/zootaxa.5722.2.5
