taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
03DD9679FFDAFFA3FF1845395C7DA477.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17892045/files/figure.png	https://doi.org/10.5281/zenodo.17892045	FIGURES 1–10. Cleidogona chilhoweensis sp. nov., and C. columbicauda sp. nov. Figs 1–8. C. chilhoweensis sp. nov. Fig. 1. Gonopods, anterior view. Fig. 2. Gonopods, posterior view. Fig. 3. Gonopods, lateral view. Fig. 4. Left leg 9, anterior view. Fig. 5. Left leg 10, anterior view. Fig. 6. Postgenital plate, posterior view. Fig. 7. Female genitalia, posterior view. Fig. 8. Female genitalia, ventral view. Figs 9, 10. C. columbicauda sp. nov. Fig. 9. Gonopods, anterior view. Fig. 10. Gonopods, posterior view. Abbreviations: ab, anterior branch of angiocoxite; ac, angiocoxite; cc, colpocoxite; cg, coxal gland; cx, coxa; cx10, coxa of leg 10; dcp, distal colpocoxite process; f9, femur of leg 9; h, hook; iv, inner valve; lp, lateral process; mp, medial process; ov, outer valve; p, process between angiocoxites; pb, posterior branch of angiocoxite; pgp, postgenital plate; r, receptacle; s, sternum; s10, sternum of legpair 10. Scale bars are 100 µm for Figs 1, 6–10; 200 µm for Figs 2, 3; 500 µm for Figs 4, 5. SEM stubs and associated specimens are retained by the Snyder Lab at Georgia College.	FIGURES 1–10. Cleidogona chilhoweensis sp. nov., and C. columbicauda sp. nov. Figs 1–8. C. chilhoweensis sp. nov. Fig. 1. Gonopods, anterior view. Fig. 2. Gonopods, posterior view. Fig. 3. Gonopods, lateral view. Fig. 4. Left leg 9, anterior view. Fig. 5. Left leg 10, anterior view. Fig. 6. Postgenital plate, posterior view. Fig. 7. Female genitalia, posterior view. Fig. 8. Female genitalia, ventral view. Figs 9, 10. C. columbicauda sp. nov. Fig. 9. Gonopods, anterior view. Fig. 10. Gonopods, posterior view. Abbreviations: ab, anterior branch of angiocoxite; ac, angiocoxite; cc, colpocoxite; cg, coxal gland; cx, coxa; cx10, coxa of leg 10; dcp, distal colpocoxite process; f9, femur of leg 9; h, hook; iv, inner valve; lp, lateral process; mp, medial process; ov, outer valve; p, process between angiocoxites; pb, posterior branch of angiocoxite; pgp, postgenital plate; r, receptacle; s, sternum; s10, sternum of legpair 10. Scale bars are 100 µm for Figs 1, 6–10; 200 µm for Figs 2, 3; 500 µm for Figs 4, 5. SEM stubs and associated specimens are retained by the Snyder Lab at Georgia College.	2025-11-19	Snyder, Bruce A.;Shear, William A.		Zenodo	biologists	Snyder, Bruce A.;Shear, William A.			
03DD9679FFDAFFA3FF1845395C7DA477.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17892061/files/figure.png	https://doi.org/10.5281/zenodo.17892061	FIGURE 29. Site locations of Cleidogona collections in the Great Smoky Mountains National Park. For species, see Table 1. Park property is shaded grey. Black lines denote state (thick) and county (thin) boundaries; grey lines are major roads.	FIGURE 29. Site locations of Cleidogona collections in the Great Smoky Mountains National Park. For species, see Table 1. Park property is shaded grey. Black lines denote state (thick) and county (thin) boundaries; grey lines are major roads.	2025-11-19	Snyder, Bruce A.;Shear, William A.		Zenodo	biologists	Snyder, Bruce A.;Shear, William A.			
03DD9679FFDCFFAEFF1846B65879A4DB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17892045/files/figure.png	https://doi.org/10.5281/zenodo.17892045	FIGURES 1–10. Cleidogona chilhoweensis sp. nov., and C. columbicauda sp. nov. Figs 1–8. C. chilhoweensis sp. nov. Fig. 1. Gonopods, anterior view. Fig. 2. Gonopods, posterior view. Fig. 3. Gonopods, lateral view. Fig. 4. Left leg 9, anterior view. Fig. 5. Left leg 10, anterior view. Fig. 6. Postgenital plate, posterior view. Fig. 7. Female genitalia, posterior view. Fig. 8. Female genitalia, ventral view. Figs 9, 10. C. columbicauda sp. nov. Fig. 9. Gonopods, anterior view. Fig. 10. Gonopods, posterior view. Abbreviations: ab, anterior branch of angiocoxite; ac, angiocoxite; cc, colpocoxite; cg, coxal gland; cx, coxa; cx10, coxa of leg 10; dcp, distal colpocoxite process; f9, femur of leg 9; h, hook; iv, inner valve; lp, lateral process; mp, medial process; ov, outer valve; p, process between angiocoxites; pb, posterior branch of angiocoxite; pgp, postgenital plate; r, receptacle; s, sternum; s10, sternum of legpair 10. Scale bars are 100 µm for Figs 1, 6–10; 200 µm for Figs 2, 3; 500 µm for Figs 4, 5. SEM stubs and associated specimens are retained by the Snyder Lab at Georgia College.	FIGURES 1–10. Cleidogona chilhoweensis sp. nov., and C. columbicauda sp. nov. Figs 1–8. C. chilhoweensis sp. nov. Fig. 1. Gonopods, anterior view. Fig. 2. Gonopods, posterior view. Fig. 3. Gonopods, lateral view. Fig. 4. Left leg 9, anterior view. Fig. 5. Left leg 10, anterior view. Fig. 6. Postgenital plate, posterior view. Fig. 7. Female genitalia, posterior view. Fig. 8. Female genitalia, ventral view. Figs 9, 10. C. columbicauda sp. nov. Fig. 9. Gonopods, anterior view. Fig. 10. Gonopods, posterior view. Abbreviations: ab, anterior branch of angiocoxite; ac, angiocoxite; cc, colpocoxite; cg, coxal gland; cx, coxa; cx10, coxa of leg 10; dcp, distal colpocoxite process; f9, femur of leg 9; h, hook; iv, inner valve; lp, lateral process; mp, medial process; ov, outer valve; p, process between angiocoxites; pb, posterior branch of angiocoxite; pgp, postgenital plate; r, receptacle; s, sternum; s10, sternum of legpair 10. Scale bars are 100 µm for Figs 1, 6–10; 200 µm for Figs 2, 3; 500 µm for Figs 4, 5. SEM stubs and associated specimens are retained by the Snyder Lab at Georgia College.	2025-11-19	Snyder, Bruce A.;Shear, William A.		Zenodo	biologists	Snyder, Bruce A.;Shear, William A.			
03DD9679FFDCFFAEFF1846B65879A4DB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17892051/files/figure.png	https://doi.org/10.5281/zenodo.17892051	FIGURES 11–16. Cleidogona columbicauda sp. nov., and C. hendrixi sp. nov. Figs 11–15. C. columbicauda sp. nov. Fig. 11. Gonopods, lateral view. Fig. 12. Right leg 9, anterior view. Fig. 13. Left leg 10, anterior view. Fig. 14. Female genitalia, posterior view. Fig. 15. Female genitalia, ventral view. Fig. 16. Gonopods of C. hendrixi sp. nov., posterior view.Abbreviations: ac, angiocoxite; cc, colpocoxite; cg, coxal gland; cx, coxa; cx10, coxa of leg 10; dcp, distal colpocoxite process; f9, femur of leg 9; h, hook; iv, inner valve; ll, laciniate lobe; lp, lateral process; mp, medial process; ov, outer valve; pgp, post genital plate; s10, sternum of legpair 10; vl, ventral lamella. Scale bars are 100 µm for Figs 11, 14–16; 200 µm for Fig. 12, 3; 500 µm for Fig. 13. SEM stubs and associated specimens are retained by the Snyder Lab at Georgia College.	FIGURES 11–16. Cleidogona columbicauda sp. nov., and C. hendrixi sp. nov. Figs 11–15. C. columbicauda sp. nov. Fig. 11. Gonopods, lateral view. Fig. 12. Right leg 9, anterior view. Fig. 13. Left leg 10, anterior view. Fig. 14. Female genitalia, posterior view. Fig. 15. Female genitalia, ventral view. Fig. 16. Gonopods of C. hendrixi sp. nov., posterior view.Abbreviations: ac, angiocoxite; cc, colpocoxite; cg, coxal gland; cx, coxa; cx10, coxa of leg 10; dcp, distal colpocoxite process; f9, femur of leg 9; h, hook; iv, inner valve; ll, laciniate lobe; lp, lateral process; mp, medial process; ov, outer valve; pgp, post genital plate; s10, sternum of legpair 10; vl, ventral lamella. Scale bars are 100 µm for Figs 11, 14–16; 200 µm for Fig. 12, 3; 500 µm for Fig. 13. SEM stubs and associated specimens are retained by the Snyder Lab at Georgia College.	2025-11-19	Snyder, Bruce A.;Shear, William A.		Zenodo	biologists	Snyder, Bruce A.;Shear, William A.			
03DD9679FFDCFFAEFF1846B65879A4DB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17892061/files/figure.png	https://doi.org/10.5281/zenodo.17892061	FIGURE 29. Site locations of Cleidogona collections in the Great Smoky Mountains National Park. For species, see Table 1. Park property is shaded grey. Black lines denote state (thick) and county (thin) boundaries; grey lines are major roads.	FIGURE 29. Site locations of Cleidogona collections in the Great Smoky Mountains National Park. For species, see Table 1. Park property is shaded grey. Black lines denote state (thick) and county (thin) boundaries; grey lines are major roads.	2025-11-19	Snyder, Bruce A.;Shear, William A.		Zenodo	biologists	Snyder, Bruce A.;Shear, William A.			
03DD9679FFD1FFACFF1846DA5A8CA51F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17892051/files/figure.png	https://doi.org/10.5281/zenodo.17892051	FIGURES 11–16. Cleidogona columbicauda sp. nov., and C. hendrixi sp. nov. Figs 11–15. C. columbicauda sp. nov. Fig. 11. Gonopods, lateral view. Fig. 12. Right leg 9, anterior view. Fig. 13. Left leg 10, anterior view. Fig. 14. Female genitalia, posterior view. Fig. 15. Female genitalia, ventral view. Fig. 16. Gonopods of C. hendrixi sp. nov., posterior view.Abbreviations: ac, angiocoxite; cc, colpocoxite; cg, coxal gland; cx, coxa; cx10, coxa of leg 10; dcp, distal colpocoxite process; f9, femur of leg 9; h, hook; iv, inner valve; ll, laciniate lobe; lp, lateral process; mp, medial process; ov, outer valve; pgp, post genital plate; s10, sternum of legpair 10; vl, ventral lamella. Scale bars are 100 µm for Figs 11, 14–16; 200 µm for Fig. 12, 3; 500 µm for Fig. 13. SEM stubs and associated specimens are retained by the Snyder Lab at Georgia College.	FIGURES 11–16. Cleidogona columbicauda sp. nov., and C. hendrixi sp. nov. Figs 11–15. C. columbicauda sp. nov. Fig. 11. Gonopods, lateral view. Fig. 12. Right leg 9, anterior view. Fig. 13. Left leg 10, anterior view. Fig. 14. Female genitalia, posterior view. Fig. 15. Female genitalia, ventral view. Fig. 16. Gonopods of C. hendrixi sp. nov., posterior view.Abbreviations: ac, angiocoxite; cc, colpocoxite; cg, coxal gland; cx, coxa; cx10, coxa of leg 10; dcp, distal colpocoxite process; f9, femur of leg 9; h, hook; iv, inner valve; ll, laciniate lobe; lp, lateral process; mp, medial process; ov, outer valve; pgp, post genital plate; s10, sternum of legpair 10; vl, ventral lamella. Scale bars are 100 µm for Figs 11, 14–16; 200 µm for Fig. 12, 3; 500 µm for Fig. 13. SEM stubs and associated specimens are retained by the Snyder Lab at Georgia College.	2025-11-19	Snyder, Bruce A.;Shear, William A.		Zenodo	biologists	Snyder, Bruce A.;Shear, William A.			
03DD9679FFD1FFACFF1846DA5A8CA51F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17892055/files/figure.png	https://doi.org/10.5281/zenodo.17892055	FIGURES 17–22. Cleidogona hendrixi sp. nov., and C. sparta sp. nov. Figs 17–20. C. hendrixi sp. nov. Fig. 17. Gonopods, anterior view. Fig. 18. Gonopods, lateral view. Fig. 19. Right leg 9, anterior view. Fig. 20. Left leg 10, anterior view. Figs 21, 22. C. sparta sp. nov. Fig. 21. Gonopods, anterior view. Fig. 22. Gonopods, posterior view. Abbreviations: ab, anterior branch of angiocoxite; ac, angiocoxite; cc, colpocoxite; cg, coxal gland; cx, coxa; cx10, coxa of leg 10; dl, dorsal lamella; f9, femur of leg 9; pb, posterior branch of angiocoxite; s, sternum; s10, sternum of legpair 10; vl, ventral lamella. Scale bars are 100 µm for Figs 17, 18; 500 µm for Figs 19, 20; 200 µm for Figs 21, 22. SEM stubs and associated specimens are retained by the Snyder Lab at Georgia College.	FIGURES 17–22. Cleidogona hendrixi sp. nov., and C. sparta sp. nov. Figs 17–20. C. hendrixi sp. nov. Fig. 17. Gonopods, anterior view. Fig. 18. Gonopods, lateral view. Fig. 19. Right leg 9, anterior view. Fig. 20. Left leg 10, anterior view. Figs 21, 22. C. sparta sp. nov. Fig. 21. Gonopods, anterior view. Fig. 22. Gonopods, posterior view. Abbreviations: ab, anterior branch of angiocoxite; ac, angiocoxite; cc, colpocoxite; cg, coxal gland; cx, coxa; cx10, coxa of leg 10; dl, dorsal lamella; f9, femur of leg 9; pb, posterior branch of angiocoxite; s, sternum; s10, sternum of legpair 10; vl, ventral lamella. Scale bars are 100 µm for Figs 17, 18; 500 µm for Figs 19, 20; 200 µm for Figs 21, 22. SEM stubs and associated specimens are retained by the Snyder Lab at Georgia College.	2025-11-19	Snyder, Bruce A.;Shear, William A.		Zenodo	biologists	Snyder, Bruce A.;Shear, William A.			
03DD9679FFD1FFACFF1846DA5A8CA51F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17892061/files/figure.png	https://doi.org/10.5281/zenodo.17892061	FIGURE 29. Site locations of Cleidogona collections in the Great Smoky Mountains National Park. For species, see Table 1. Park property is shaded grey. Black lines denote state (thick) and county (thin) boundaries; grey lines are major roads.	FIGURE 29. Site locations of Cleidogona collections in the Great Smoky Mountains National Park. For species, see Table 1. Park property is shaded grey. Black lines denote state (thick) and county (thin) boundaries; grey lines are major roads.	2025-11-19	Snyder, Bruce A.;Shear, William A.		Zenodo	biologists	Snyder, Bruce A.;Shear, William A.			
03DD9679FFD3FFAAFF18479E5ADEA717.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17892055/files/figure.png	https://doi.org/10.5281/zenodo.17892055	FIGURES 17–22. Cleidogona hendrixi sp. nov., and C. sparta sp. nov. Figs 17–20. C. hendrixi sp. nov. Fig. 17. Gonopods, anterior view. Fig. 18. Gonopods, lateral view. Fig. 19. Right leg 9, anterior view. Fig. 20. Left leg 10, anterior view. Figs 21, 22. C. sparta sp. nov. Fig. 21. Gonopods, anterior view. Fig. 22. Gonopods, posterior view. Abbreviations: ab, anterior branch of angiocoxite; ac, angiocoxite; cc, colpocoxite; cg, coxal gland; cx, coxa; cx10, coxa of leg 10; dl, dorsal lamella; f9, femur of leg 9; pb, posterior branch of angiocoxite; s, sternum; s10, sternum of legpair 10; vl, ventral lamella. Scale bars are 100 µm for Figs 17, 18; 500 µm for Figs 19, 20; 200 µm for Figs 21, 22. SEM stubs and associated specimens are retained by the Snyder Lab at Georgia College.	FIGURES 17–22. Cleidogona hendrixi sp. nov., and C. sparta sp. nov. Figs 17–20. C. hendrixi sp. nov. Fig. 17. Gonopods, anterior view. Fig. 18. Gonopods, lateral view. Fig. 19. Right leg 9, anterior view. Fig. 20. Left leg 10, anterior view. Figs 21, 22. C. sparta sp. nov. Fig. 21. Gonopods, anterior view. Fig. 22. Gonopods, posterior view. Abbreviations: ab, anterior branch of angiocoxite; ac, angiocoxite; cc, colpocoxite; cg, coxal gland; cx, coxa; cx10, coxa of leg 10; dl, dorsal lamella; f9, femur of leg 9; pb, posterior branch of angiocoxite; s, sternum; s10, sternum of legpair 10; vl, ventral lamella. Scale bars are 100 µm for Figs 17, 18; 500 µm for Figs 19, 20; 200 µm for Figs 21, 22. SEM stubs and associated specimens are retained by the Snyder Lab at Georgia College.	2025-11-19	Snyder, Bruce A.;Shear, William A.		Zenodo	biologists	Snyder, Bruce A.;Shear, William A.			
03DD9679FFD3FFAAFF18479E5ADEA717.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17892059/files/figure.png	https://doi.org/10.5281/zenodo.17892059	FIGURES 23–28. Cleidogona sparta sp. nov. Fig. 23. Gonopods, lateral view. Fig. 24. Right leg 9, anterior view. Fig. 25. Left leg 10, anterior view. Fig. 26. Female genitalia, posterior view. Fig. 27. Sculpture of postgenital plate. Fig. 28. Female genitalia, ventral view. Abbreviations: ab, anterior branch of angiocoxite; ac, angiocoxite; cc, colpocoxite; cg, coxal gland; cp, coxal process; cx, coxa; cx10, coxa of leg 10; f9, femur of leg 9; iv, inner valve; ov, outer valve; pb, posterior branch of angiocoxite; pgp, post genital plate; r, receptacle; s, sternum; t, tooth. Scale bars are 200 µm for Fig. 23; 500 µm for Figs 24, 25; 100 µm for Figs 26, 28; 10 µm for Fig. 27. SEM stubs and associated specimens are retained by the Snyder Lab at Georgia College.	FIGURES 23–28. Cleidogona sparta sp. nov. Fig. 23. Gonopods, lateral view. Fig. 24. Right leg 9, anterior view. Fig. 25. Left leg 10, anterior view. Fig. 26. Female genitalia, posterior view. Fig. 27. Sculpture of postgenital plate. Fig. 28. Female genitalia, ventral view. Abbreviations: ab, anterior branch of angiocoxite; ac, angiocoxite; cc, colpocoxite; cg, coxal gland; cp, coxal process; cx, coxa; cx10, coxa of leg 10; f9, femur of leg 9; iv, inner valve; ov, outer valve; pb, posterior branch of angiocoxite; pgp, post genital plate; r, receptacle; s, sternum; t, tooth. Scale bars are 200 µm for Fig. 23; 500 µm for Figs 24, 25; 100 µm for Figs 26, 28; 10 µm for Fig. 27. SEM stubs and associated specimens are retained by the Snyder Lab at Georgia College.	2025-11-19	Snyder, Bruce A.;Shear, William A.		Zenodo	biologists	Snyder, Bruce A.;Shear, William A.			
03DD9679FFD3FFAAFF18479E5ADEA717.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17892061/files/figure.png	https://doi.org/10.5281/zenodo.17892061	FIGURE 29. Site locations of Cleidogona collections in the Great Smoky Mountains National Park. For species, see Table 1. Park property is shaded grey. Black lines denote state (thick) and county (thin) boundaries; grey lines are major roads.	FIGURE 29. Site locations of Cleidogona collections in the Great Smoky Mountains National Park. For species, see Table 1. Park property is shaded grey. Black lines denote state (thick) and county (thin) boundaries; grey lines are major roads.	2025-11-19	Snyder, Bruce A.;Shear, William A.		Zenodo	biologists	Snyder, Bruce A.;Shear, William A.			
