taxonID	type	description	language	source
03D58C6FFF892B0EFF4D7ED5FD45FBBF.taxon	description	Fig. 2	en	Chouangthavy, Bounsanong, Yoshida, Takahiro, Eguchi, Katsuyuki (2025): First discovery of males of two ambrosia beetle species of Xylosandrus Reitter, 1913 (Coleoptera, Curculionidae, Scolytinae), with their first records from Lao P. D. R. Zootaxa 5604 (2): 118-130, DOI: 10.11646/zootaxa.5604.2.2, URL: https://doi.org/10.11646/zootaxa.5604.2.2
03D58C6FFF892B0EFF4D7ED5FD45FBBF.taxon	materials_examined	Material examined. 6 males: Lao P. D. R, Nam Ha National Biodiversity Conservation Area, Muang Sing, Luang Namtha province, Northern Lao P. D. R, 21.126178 N, 101.243714 E, 995 m alt., 14. ix. 2023, insecticide spray (B. Chouangthavy, leg; NUOL).	en	Chouangthavy, Bounsanong, Yoshida, Takahiro, Eguchi, Katsuyuki (2025): First discovery of males of two ambrosia beetle species of Xylosandrus Reitter, 1913 (Coleoptera, Curculionidae, Scolytinae), with their first records from Lao P. D. R. Zootaxa 5604 (2): 118-130, DOI: 10.11646/zootaxa.5604.2.2, URL: https://doi.org/10.11646/zootaxa.5604.2.2
03D58C6FFF892B0EFF4D7ED5FD45FBBF.taxon	diagnosis	Male diagnosis. The male is characterized by the following combination of characteristics: frons weakly convex; eyes upper part slightly smaller or equal to lower part; anterior margin of pronotum without serrations; declivital face with large punctures, posterolateral margins of elytra costate; disc and declivity with dense hair-like setae; declivity steep, beginning at basal third, then tapering toward rounded apex; aedeagus long and slender; penis body long and narrowed towards apex (Fig 6 D).	en	Chouangthavy, Bounsanong, Yoshida, Takahiro, Eguchi, Katsuyuki (2025): First discovery of males of two ambrosia beetle species of Xylosandrus Reitter, 1913 (Coleoptera, Curculionidae, Scolytinae), with their first records from Lao P. D. R. Zootaxa 5604 (2): 118-130, DOI: 10.11646/zootaxa.5604.2.2, URL: https://doi.org/10.11646/zootaxa.5604.2.2
03D58C6FFF892B0EFF4D7ED5FD45FBBF.taxon	description	Description of male. Body small, 1.1 – 1.5 mm long (mean = 1.3 mm; n = 6), 0.5 – 0.7 (mean = 0.6) time longer than wide. Pronotum, scutellum and elytra dark brown or black; antennae and legs yellowish brown to brown. Head. Epistoma undivided, transverse, with hair-like setae, with smooth surface. Frons weakly convex, weakly punctate; punctures each bearing an erect hair-like seta (0.09 - 0.15 mm), yellowish brown. Eyes small (approximately 0.09 – 0.1 mm from upper-most to lower-most), with anterior margin weakly emarginate; upper part slightly smaller or equal in size as lower part. Submentum distinctly triangular, large, and slightly impressed. Antennal scape long and uniformly thick, slightly longer than the club (3: 2); pedicel thick, as wide as the scape, varying in length, shorter than or same length as the funicle; funicle 2 - to 4 - segmented (4 - segmented in most cases), with segments thick and short, wider than long (3: 1), in individuals with 2 - segmented funicles (Fig. 4 A), segments very flat and wider than long (4: 1), in individuals with 4 - segmented funicles (Fig. 5 B); funicle sometimes incompletely fused with the club (Fig. 4 B): club obliquely truncate, longer than wide as in females; segment 1 well-defined, covering the entire posterior face, with the anterior apical margin either pointed at the first club (Fig. 5 A) or even (Fig. 5 B) on anterior face. Pronotum 0.3 – 0.6 mm (mean = 0.45 mm), as long as wide, rounded in dorsal view (type 1 defined in Smith et al. 2020); side slightly cured forward in basal and apical thirds; pronotal disc punctate, with summit situated at middle 1 / 3 from base, flat and smooth. Base bears weakly defined short hair-like setae. Lateral sides of the disc short or slightly equal to the anterior slope, basic type 0, flat and slightly curved toward the summit, summit at basal 1 / 2; anterior margin without serrations, anterior slope weakly asperate, asperities weakly spaced, flat, lower and rare transverse toward the summit. Scutellum rounded triangular or U-shaped, moderately sized, flattened, dark brown, more visible from above. Elytra 0.5 – 0.9 mm (mean = 0.7 mm) as long as wide, 0.4 – 0.8 mm (mean = 0.5 mm), as long as pronotum, proportion of elytral length to pronotal length (0.7 – 0.8 mm), slightly curved at bases; humeral angles rounded; lateral sides curving from humeral angle to apex. Disc short, basal area 1 / 3 of disc slightly rounded, apical 3 / 4 humped and connecting to declivital strongly curved toward apex; surface punctate, punctures fine and setose; striae and interstriae regular punctures, becoming finer towards apex. Declivital commencing from basal third, steep; margin rounded and costate; posterolateral margin costate in apical 1 / 3; striae and interstriae with confused punctures? on the posterior half of elytra and declivity, each puncture becoming obsolete on declivity towards apex, bearing long fine hairs. Legs. Procoxae widely separated, interspace between procoxae 0.03 times as wide as procoxa. Protibiae obliquely triangular, with 3 – 5 socketed teeth on lateral margin; posterior face smooth. Meso- and metatibiae with 6 – 8 socketed teeth. Genitalia. Aedeagus long and slender; penis body long and narrowly to apex; apodeme slenderly curves gradually towards the apex (Fig. 6 D)	en	Chouangthavy, Bounsanong, Yoshida, Takahiro, Eguchi, Katsuyuki (2025): First discovery of males of two ambrosia beetle species of Xylosandrus Reitter, 1913 (Coleoptera, Curculionidae, Scolytinae), with their first records from Lao P. D. R. Zootaxa 5604 (2): 118-130, DOI: 10.11646/zootaxa.5604.2.2, URL: https://doi.org/10.11646/zootaxa.5604.2.2
03D58C6FFF892B0EFF4D7ED5FD45FBBF.taxon	discussion	Remarks: The antennal structure of the population of male X. derupteterminatus is extremely variable. Initially, we classified the male specimens into three distinct morphospecies based on the distinctly varied characteristics of their antennae. This variation sometimes appears even on a singleton specimen, for instance, the apical margin of the first segment of the left club is medially pointed in anterior view, while one of the right club is even (Fig. 5 A and B). Additionally, in another population, the funicle segments are thick and composed of two or three distinct segments (Fig. 4 A and B). The color varies from black to yellowish brown.	en	Chouangthavy, Bounsanong, Yoshida, Takahiro, Eguchi, Katsuyuki (2025): First discovery of males of two ambrosia beetle species of Xylosandrus Reitter, 1913 (Coleoptera, Curculionidae, Scolytinae), with their first records from Lao P. D. R. Zootaxa 5604 (2): 118-130, DOI: 10.11646/zootaxa.5604.2.2, URL: https://doi.org/10.11646/zootaxa.5604.2.2
03D58C6FFF892B0EFF4D7ED5FD45FBBF.taxon	distribution	Distribution in Lao P. D. R: Luang Namtha province	en	Chouangthavy, Bounsanong, Yoshida, Takahiro, Eguchi, Katsuyuki (2025): First discovery of males of two ambrosia beetle species of Xylosandrus Reitter, 1913 (Coleoptera, Curculionidae, Scolytinae), with their first records from Lao P. D. R. Zootaxa 5604 (2): 118-130, DOI: 10.11646/zootaxa.5604.2.2, URL: https://doi.org/10.11646/zootaxa.5604.2.2
03D58C6FFF892B0EFF4D7ED5FD45FBBF.taxon	biology_ecology	Biological notes. We collected populations of this species from Pometia pinnata (Sapindaceae), found on small branches with a diameter ranging from 5 to 7 cm. These branches had fallen from a live tree, situated near a small river characterized by high humidity. The river maintained a continuous flow of water throughout the year. This primary forest habitat where samples were collected is located approximately 1 km from the main road. This ambrosia beetle species (X. derupteterminatus) was previously recorded only on Mangifera indica (Anacardiaceae) and Agathis (Araucariaceae) (Smith et al. 2020).	en	Chouangthavy, Bounsanong, Yoshida, Takahiro, Eguchi, Katsuyuki (2025): First discovery of males of two ambrosia beetle species of Xylosandrus Reitter, 1913 (Coleoptera, Curculionidae, Scolytinae), with their first records from Lao P. D. R. Zootaxa 5604 (2): 118-130, DOI: 10.11646/zootaxa.5604.2.2, URL: https://doi.org/10.11646/zootaxa.5604.2.2
03D58C6FFF8B2B01FF4D7DEDFC7EF854.taxon	description	Fig. 3	en	Chouangthavy, Bounsanong, Yoshida, Takahiro, Eguchi, Katsuyuki (2025): First discovery of males of two ambrosia beetle species of Xylosandrus Reitter, 1913 (Coleoptera, Curculionidae, Scolytinae), with their first records from Lao P. D. R. Zootaxa 5604 (2): 118-130, DOI: 10.11646/zootaxa.5604.2.2, URL: https://doi.org/10.11646/zootaxa.5604.2.2
03D58C6FFF8B2B01FF4D7DEDFC7EF854.taxon	materials_examined	Material examined. 1 male: Lao P. D. R, Nam Ha National Biodiversity Conservation Area, Muang Sing, Luang Namtha province, Northern Lao P. D. R, 21.126178 N, 101.243714 E, 995 m alt., 14. ix. 2023, insecticide spray (B. Chouangthavy, leg; NUOL).	en	Chouangthavy, Bounsanong, Yoshida, Takahiro, Eguchi, Katsuyuki (2025): First discovery of males of two ambrosia beetle species of Xylosandrus Reitter, 1913 (Coleoptera, Curculionidae, Scolytinae), with their first records from Lao P. D. R. Zootaxa 5604 (2): 118-130, DOI: 10.11646/zootaxa.5604.2.2, URL: https://doi.org/10.11646/zootaxa.5604.2.2
03D58C6FFF8B2B01FF4D7DEDFC7EF854.taxon	diagnosis	Male diagnosis. Frons strongly convex, weakly punctate; punctures each bearing erect hair-like seta; eyes with upper part smaller than lower part; anterior margin of pronotum without serrations; elytral disc humped, strongly curved toward declivity, without granulates, finely punctate; declivital face with large strial punctures; each of interstriae with erect hair-like setae; posterolateral margins of elytra carinate; aedeagus short and wide; penis body wide at the base, vase-shaped, gradually extending to a broader middle and narrowed towards apex (Fig 6 A).	en	Chouangthavy, Bounsanong, Yoshida, Takahiro, Eguchi, Katsuyuki (2025): First discovery of males of two ambrosia beetle species of Xylosandrus Reitter, 1913 (Coleoptera, Curculionidae, Scolytinae), with their first records from Lao P. D. R. Zootaxa 5604 (2): 118-130, DOI: 10.11646/zootaxa.5604.2.2, URL: https://doi.org/10.11646/zootaxa.5604.2.2
03D58C6FFF8B2B01FF4D7DEDFC7EF854.taxon	description	Description of male. body small, 1.4 mm long (n = 1), 0.7 times longer than wide. Pronotum and elytra light brown. Head. Epistoma not divided, transverse, with hair-like setae, with smooth surface. Frons strongly convex, weakly punctate; punctures each bearing an erect hair-like seta, both punctures and setae yellowish brown. Eyes somewhat strongly emarginate (approximately 0.09 - 0.1 mm), upper part smaller than lower part. Submentum large, distinctly triangular, slightly impressed. Antennal scape long, normal thick and bent, as long as club. Pedicel thick, as wide as scape. Antennal club as in female, combined length of the funicular segments about 2 times as long as scape, 4 th segment of funicle fused with antennal club. Pronotum 0.6 mm, as long as wide, rounded in dorsal view (type 1). Pronotal disc punctate, side slightly curved in basal towards apex, summit at middle, with asperities weak on anterior 1 / 2. Lateral sides of disc as short or slightly equal anterior slope, basic type 0, flat, slightly curved toward the summit, summit at basal 1 / 2; anterior margin without serrations, anterior slope weakly asperate, asperities widely spaced, flat, lower toward the summit. Scutellum triangular, large, U-shaped, at same level as surrounding elytral surface. Elytra 0.8 mm as long as wide 0.6; sides feebly convex on basal 1 / 3, then curving toward apex. Base slightly curved, humeral angles rounded, lateral side curving from humeral angle to apex. Disc short, in lateral view humped, apex rounded; striae and interstriae smooth, apical 3 / 4 humped and connecting to declivity, feebly convex toward apex, with few small punctures, punctures fine and rarely setose. Declivital commencing from basal third, steep; posterolateral margins strongly carinate to interstriae 7; striae and interstriae confused on the posterior half of elytra and declivity, each puncture becoming obsolete on declivity towards apex, rarely with short or long fine hair-like setae. Legs. Procoxae widely separated, interspace between procoxae 0.03 times as wide as procoxa. Protibiae obliquely triangular, with five socketed teeth on lateral margin; posterior face smooth; mesotibiae with eight socketed teeth; metatibiae with 10 socketed teeth. Genitalia. Aedeagus short and wide; penis body vase-shaped, wide at the base, gradually extending to a broader middle and narrowed towards apex; apodeme thick and curved, resembling the shape of forceps.	en	Chouangthavy, Bounsanong, Yoshida, Takahiro, Eguchi, Katsuyuki (2025): First discovery of males of two ambrosia beetle species of Xylosandrus Reitter, 1913 (Coleoptera, Curculionidae, Scolytinae), with their first records from Lao P. D. R. Zootaxa 5604 (2): 118-130, DOI: 10.11646/zootaxa.5604.2.2, URL: https://doi.org/10.11646/zootaxa.5604.2.2
03D58C6FFF8B2B01FF4D7DEDFC7EF854.taxon	distribution	Distribution in Lao P. D. R: Luang Namtha province	en	Chouangthavy, Bounsanong, Yoshida, Takahiro, Eguchi, Katsuyuki (2025): First discovery of males of two ambrosia beetle species of Xylosandrus Reitter, 1913 (Coleoptera, Curculionidae, Scolytinae), with their first records from Lao P. D. R. Zootaxa 5604 (2): 118-130, DOI: 10.11646/zootaxa.5604.2.2, URL: https://doi.org/10.11646/zootaxa.5604.2.2
03D58C6FFF8B2B01FF4D7DEDFC7EF854.taxon	biology_ecology	Biological notes: This species both male and female collected the same habitat environment and host plant species of X. derupteterminatus. Previously, female of this species had only been recorded on Eupatorium (Asteraceae) according to Eggers (1940). In the present study, we identified additional host plant species for X. derupteterminatus, collected from Pometia pinnata (Sapindaceae).	en	Chouangthavy, Bounsanong, Yoshida, Takahiro, Eguchi, Katsuyuki (2025): First discovery of males of two ambrosia beetle species of Xylosandrus Reitter, 1913 (Coleoptera, Curculionidae, Scolytinae), with their first records from Lao P. D. R. Zootaxa 5604 (2): 118-130, DOI: 10.11646/zootaxa.5604.2.2, URL: https://doi.org/10.11646/zootaxa.5604.2.2
