taxonID	type	description	language	source
03D487FDFF9A8B6E1BBA20CD1BC4FF17.taxon	type_taxon	Type species: Bassus elegans Gravenhorst, 1829. Designated by Viereck (1912).	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF9A8B6E1BBA20CD1BC4FF17.taxon	diagnosis	Diagnosis. According to Klopfstein (2014 b), this genus can be distinguished from other genera by the following combination of character states. Face coriaceous and matt; without vertical impressions; entirely black or black with yellow central marking in female (Figs 2 B, 3 B, 4 B, 5 B, 6 B, 7 B, 9 B, 10 B, 11 B, 12 B, 13 B); entirely yellow or black with yellow inner orbits and yellow central marking in male (Figs 2 E, 3 E, 4 D, 6 F, 8 B, 10 E, 11 F, 12 F, 13 D). Clypeus usually with thin apical margin; impressed along apical margin; resulting in central area being convex. Antenna with apical flagellomeres usually longer than wide. Tyloids always present in male; linear and narrow and without long setae in most species (exceptions: H. tauriscorium and H. venustus). Mesoscutum without notaulus; variously sculptured, ranging from entirely smooth and shining, with or without punctures, to being strongly coriaceous and matt; yellow shoulder markings present or absent; inner corners of markings (if present) sometimes extended into two parallel lines on mesoscutum. Scutellum only carinate basally. Mesopleuron sometimes entirely smooth and shining; often punctate and / or coriaceous at least on lower half; epicnemial carina complete ventrally. Propodeum sometimes with full set of carinae enclosing basal, petiolar, and lateral areas; carinae usually partly or fully reduced; spiracle inconspicuous. Areolet present or absent, if present, vein 3 rs-m often interrupted or unpigmented. Hind wing with 2 – 4 basal hamuli. Hind tibia with various colouration, i. e., orange, yellow, or white with dark apex and subbasal spot, very rarely (H. venustus) even black-white-black banded. Female metasoma usually evenly tapered to apex (Figs 3 D, 6 D); sometimes strongly compressed posterior to third segment (Fig. 12 D); posterior margins never concave; without transverse impressions. T I usually without latero-median carina, but if carina present, its two carinae converge over basal half, parallel, and widely separated on apical half (Fig. 1 B). T II with spiracle above lateral fold. T III with spiracle usually above (but rarely below) or behind lateral fold. Metasoma black (Fig. 3 D); sometimes with yellow markings, or marked with orange (Figs 6 D, G, 11 G, 12 D). Ovipositor sheaths 0.3 × as long as hind tibia; either parallel-sided and fully enclosing ovipositor or tapered and diagonally truncate; with inconspicuous setae ventrally and apically. T VIII and T IX of male present as separate sclerites; hypopygium about 2.0 × wider than long, emarginated apically and forming two lobes; outer corners of lobes rounded.	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF9A8B6E1BBA20CD1BC4FF17.taxon	distribution	Distribution. Holarctic, Neotropical, and Oriental regions (Yu et al., 2016).	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF9B8B691BBA26E11B4EFE01.taxon	description	(Figs 2 A – E)	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF9B8B691BBA26E11B4EFE01.taxon	materials_examined	Specimens examined. JAPAN: [Hokkaido] 1 F (holotype), Sapporo, 5. VII. 1951, T. Uchida leg. (SEHU); 1 M, Zenibako, 24. VI. 1968, K. Kusigemati leg. (SEHU); 1 F, Akkeshi Town, Bekanbeushi marsh, 31. VIII. 2003, R. Matsumoto leg. (OMNH); 1 F, ditto, 12 – 31 VII 2003, R. Matsumoto leg. (MT) (OMNH); 1 F, ditto, 1 – 31 VIII 2003, R. Matsumoto leg. (MsT) (OMNH); 1 F, Hidaka Town, Uenzaru-gawa, 1 – 28 VIII 2007, A. Ueda leg. (MsT) (KPMNH); 1 F, ditto, 10 VII – 1 VIII 2007, A. Ueda leg. (MsT) (KPNMH); 1 F, Kamikawa Town, Kamikoshi, 1. VIII. 2021, S. Morishita leg. (TMNH); [Honshu] 1 F, Tochigi Pref., Kuriyama Vil., Kinunuma, 19 VII – 1 VIII 2004, H. Makihara leg. (MsT) (KPNMH); 2 M, Tochigi Pref., Nasushiobara, 15. V. 2011, T. Matsumura leg. (KPNMH); 1 F, Yamanashi Pref., Koshu City, Mt. Daibosatsu, Kaminikkawatoge, 16. VI. 2007, T. Ban leg. (KPNMH); 1 M, Yamanashi Pref., Hokuto City, Masutomi, Biwakubozawa, 27. IX. 2008, K. Watanabe leg. (KPNMH); 1 F, Yamanashi Pref., Narusawa Vil., Kouyoudai, 15. V. 2021, K. Watanabe leg. (KPNMH); 1 F, Kanagawa Pref., Miura City, 29 – 30 IV 1976, N. Obayashi leg. (EUM); 1 F, Kanagawa Pref., Kamakura City, 27. VI. 2000, H. Nagase leg. (KPNMH); 2 M, Kanagawa Pref., Hiratsuka City, Mt. Komayama, Shonandaira, 21. IV. 2007, K. Watanabe leg. (KPNMH); 3 M, Kanagawa Pref., Hadano City, Mt. Koubouyama, 4. IV. 2007, K. Watanabe leg. (KPNMH); 1 F, ditto, 27. IX. 2008, K. Watanabe leg. (KPNMH); 1 F, Kanagawa Pref., Odawara City, Kamisoga, 29. IV. 2017, K. Watanabe leg. (KPNMH); 1 F, Kanagawa Pref., Hakone Town, Mt. Komagatake, 20. IV. 2000, H. Nagase leg. (KPNMH); 1 F, Toyama Pref., Toyama City, Arimine, Inonedani, 14 – 21 VII 2009, M. Watanabe leg. (MT) (KPMNH); 2 F, ditto, 28 VII – 4 VIII 2009, M. Watanabe leg. (MsT) (KPMNH); 1 F, ditto, 4 – 11 VIII 2009, M. Watanabe leg. (MT) (KPMNH); 1 F, ditto, 11 – 16 VIII 2009, M. Watanabe leg. (MT) (KPMNH); 1 F, ditto, 25 VIII – 1 IX 2009, M. Watanabe leg. (MT) (KPMNH); 1 F, Toyama Pref., Toyama City, Arimine, Jurodani, 4 – 11 VIII 2009, M. Watanabe leg. (MT) (KPMNH); 1 F, ditto, 16 – 25 VIII 2009, M. Watanabe leg. (MsT) (KPMNH); 1 F, Toyama Pref., Nanto City, Togamura, Kamimomose, 25 VIII – 1 IX 2009, M. Watanabe leg. (MsT) (KPMNH); 1 M, ditto, 15 – 29 IX 2009, M. Watanabe leg. (MT) (KPMNH); 2 F, Nagano Pref., Otaki Vil., Mt. Ontakesan, Hakkaisan, 30. VII. 2012, S. Fujie leg. (KPMNH); 1 F, ditto, 13. VI. 2015, K. Watanabe leg. (KPMNH); 1 F, ditto, 4. VIII. 2017, K. Watanabe leg. (KPMNH); 1 F, Nagano Pref., Otaki Vil., Mt. Ontakesan, Tanohara, 13. VI. 2015, K. Watanabe leg. (KPMNH); 1 F, Aichi Pref., Toyohashi City, Imure-cho, Takayama, 27 V 2019, S. Morishita leg. (YPT) (TMNH); 1 F, Aichi Pref., Toyohashi City, Ishimaki-cho, Ege, 15 X 2018, S. Morishita leg. (TMNH); 1 F 1 M, Aichi Pref., Toyohashi City, Oiwa-cho, Taimatsutoge, 7 IV 2021, S. Morishita leg. (TMNH); 2 F 2 M, Aichi Pref., Toyokawa City, Mikami-cho, 15 IX 2019, S. Morishita leg. (TMNH); 1 M, Aichi Pref., Toyokawa City, Mt. Hongusan, 17 V 2019, S. Morishita leg. (TMNH).	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF9B8B691BBA26E11B4EFE01.taxon	diagnosis	Diagnosis. Antenna with 20 – 22 (in female) or 21 – 22 (in male) flagellomeres. Tyloids on FL VII to FL XIV in male. Face coriaceous and matt; densely punctate. Clypeus depressed apically; robes rounded at outer corners. Mesoscutum smooth and polished; densely punctate; often with coriaceous area medially. Mesopleuron smooth and polished; often with coriaceous area below speculum; densely punctate except for on and below speculum. Propodeum finely rugulose; posterior part of lateromedian longitudinal carina and pleural carina present. Fore wing length 5.5 – 8.5 mm. Areolet present. Dorsal part of hind coxa coriaceous basally, smooth apically. Metasoma dorsoventrally depressed in both sexes. Latero-median carina usually indistinct, at most present on basal 0.3 of T I. T I 1.0 – 1.1 (in female) or 1.1 – 1.5 (in male) × as long as maximum width; rugulose punctate laterally; finely striate medially; otherwise coriaceous. T II 0.7 – 0.85 (in female) or 0.8 – 0.9 (in male) × as long as maximum width. Face without pair of yellow markings along each inner orbit (Figs 2 B, E). Face entirely black or black with yellow median marking which separated from clypeus in female (Fig. 2 B); entirely yellow in male (Fig. 2 E). Clypeus entirely yellow in both sexes (Figs 2 B, E). Malar space entirely black in females (Fig. 2 B); entirely yellow in male (Fig. 2 E). Yellow shoulder markings of mesoscutum present (Figs 2 A, D). Scutellum black with yellow apical marking (Fig. 2 C). Fore and mid coxae entirely yellow to yellowish-brown in female (Fig. 2 A); entirely yellow in male (Fig. 2 D). Hind coxa entirely orange or orange with yellow apex in female (Fig. 2 A); yellow with dorsal orange to black stripe or orange to black with yellow apex in male (Fig. 2 D). Hind tibia white; darkened basally and apically; often with black subbasal marking (Figs 2 A, D). Metasoma entirely black in female. T III with pair of yellow to brown basal markings in male. T IV and sometimes T V each with transverse yellow apical band in male.	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF9B8B691BBA26E11B4EFE01.taxon	distribution	Distribution. Japan (Hokkaido, Honshu, and Shikoku). Outside Japan, this species is distributed in Korea (Yu et al. 2016).	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF9B8B691BBA26E11B4EFE01.taxon	biology_ecology	Bionomics. Host is unknown. In Japan, adults were collected in broadleaved forests and wetlands.	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF9E8B6A1BBA20CD1DBAFBBD.taxon	description	(Figs 3 A – F)	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF9E8B6A1BBA20CD1DBAFBBD.taxon	materials_examined	Specimens examined. JAPAN: [Honshu] 1 F, Yamagata Pref., Mamurogawa Town, 14. VII. 2012, Y. Matsubara & K. Fukuda leg. (MsT) (KPMNH); 1 F, Mie Pref., Inabe City, Mt. Fujiwaradake, 1. VI. 2021, S. Morishita leg. (KPMNH); 2 F, ditto, (TMNH). GERMANY: 1 F (det. E. Diller), Gottingen, 6. VII. 1946, (ZSM). ITALY: 1 M (det. E. Diller), Catania, Tremestieri Etneo, 4. V. 1994, Turrisi leg. (ZSM).	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF9E8B6A1BBA20CD1DBAFBBD.taxon	diagnosis	Diagnosis. See Klopfstein (2014 b).	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF9E8B6A1BBA20CD1DBAFBBD.taxon	distribution	Distribution. Japan (Honshu). Outside Japan, this species is widely distributed in Western Palearctic region (Yu et al. 2016).	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF9E8B6A1BBA20CD1DBAFBBD.taxon	biology_ecology	Bionomics. Host is unknown. In Japan, adults were collected from the canopies of broad‐leaved trees at altitudes of ca. 1,000 meters.	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF9E8B6A1BBA20CD1DBAFBBD.taxon	discussion	Remarks. This is the first record of this species from Japan. Japanese specimens share the character states with the diagnosis of European specimens proposed by Klopfstein (2014 b) well, while Japanese specimens differs from European specimens in the following character states: fore wing length 6.5 – 8.0 mm (5.1 – 6.5 mm in European specimens); hind trochantellus black with reddish-brown apex (entirely orange in European specimens); hind femur black with reddish-brown base (entirely orange in European specimens); hind tibia entirely black (orange with apex dark in European specimens). However, as no significant differences were found in other characters, we conclude that the observed variation is intraspecific variation within a single species.	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF9F8B6A1BBA23451D28FC9F.taxon	description	(Figs 4 A – D)	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF9F8B6A1BBA23451D28FC9F.taxon	diagnosis	Diagnosis. See Klopfstein (2014 b).	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF9F8B6A1BBA23451D28FC9F.taxon	distribution	Distribution. Japan (Kunashiri Is.) (Klopfstein 2014 a). Outside Japan, this species is widely distributed in Holarctic region (Yu et al. 2016). Bionomics. Host is unknown in Japan. Outside of Japan, Episyrphus balteatus (De Geer 1776) (Diptera, Syrphidae) has been recorded as a host (Hedwig 1958; Thirion 1994). Calybites phasianipennellus (Hubner 1813) and Lasiocampa quercus (Linnaeus 1758) (Lepidoptera, Lasiocampidae) have also been recorded as hosts (Morley 1906), but are considered doubtful records.	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF9F8B6A1BBA23451D28FC9F.taxon	discussion	Remarks. We could not find additional material from Japan.	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF9F8B671BBA261A1C5EF8DD.taxon	description	(Figs 5 A – C)	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF9F8B671BBA261A1C5EF8DD.taxon	materials_examined	Specimens examined. JAPAN: [Hokkaido] 1 F, Kamikawa Town, Ginsendaisen, 1. VIII. 2021, K. Watanabe leg. (KPMNH); 1 F, Chitose City, Bifue, Kusabue-rindo, 8 – 21 VII 2012, N. Kuhara leg. (MsT) (EUM). [Honshu] 1 F, Gunma Pref., Tsumagoi Vil., Kanbara, Takamine-kogen, 3. IX. 2015, K. Watanabe leg. (KPMNH); 1 F, ditto, 23. VIII. 2016, S. Shimizu leg. (KPMNH); 1 F, Kanagawa Pref., Mt. Hinokiboramaru, 23. VI. 2013, (FIT) (KPMNH); 1 F, Kanagawa Pref., Odawara City, Kamisoga, 29. IV. 2017, K. Watanabe leg. (KPMNH); 1 F, Toyama Pref., Toyama City, Arimine, Inonedani, 25 VIII – 1 IX 2009, M. Watanabe leg. (MsT) (KPMNH); 1 F, ditto, 15 – 22 IX 2009, M. Watanabe leg. (MsT) (KPMNH); 4 F, Nagano Pref., Otaki Vil., Mt. Ontakesan, Hakkaisan, 7. VIII. 2010, K. Watanabe leg. (KPMNH); 2 F, ditto, 4. VIII. 2017, K. Watanabe leg. (KPMNH); 2 F, Nagano Pref., Otaki Vil., Mt. Ontakesan, 25 VI – 15 VII 2015, S. Shimizu leg. (MsT) (KPMNH); 1 F, Aichi Pref., Shitara Town, Tsuguhonsawa, 26 V – 23 VI 2020, S. Morishita leg. (MsT) (TMNH). [Kyushu] 1 F, Kagoshima Pref., Takachihonomine, 27. VII. 1973, K. Kusigemati leg. (SEHU).	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF9F8B671BBA261A1C5EF8DD.taxon	diagnosis	Diagnosis. Antenna with 21 – 22 (in female) or 23 (in male) flagellomeres. Tyloids on FL VII to FL XV in male. Face coriaceous and matt; densely punctate. Clypeus depressed apically; robes rounded at outer corners. Mesoscutum coriaceous and matt in female; coriaceous and polished in male; densely punctate in both sexes. Mesopleuron coriaceous and matt; densely punctate except for small smooth area around the speculum (Fig. 5 C). Propodeum finely rugulose; posterior part of lateromedian longitudinal carina and pleural carina present. Fore wing length 5.0 – 7.0 mm. Areolet present. Dorsal part of hind coxa coriaceous and matt. Metasoma dorsoventrally depressed in both sexes. Latero-median carina usually indistinct; at most present on basal 0.4 of T I (Fig. 1 A). T I 1.1 – 1.3 × as long as maximum width; rugulose laterally, otherwise coriaceous. T II 0.8 – 0.9 × as long as maximum width. Face without pair of yellow markings along each inner orbits (Fig. 5 B). Face black with yellow median marking which separated from clypeus in female (Fig. 5 B); entirely yellow in male. Clypeus entirely yellow to brown in both sexes (Fig. 5 B). Malar space entirely black in female (Fig. 5 B); entirely yellow in male. Yellow shoulder markings of mesoscutum present (Fig. 5 A). Scutellum entirely black. All coxae entirely black in female (Fig. 5 A). Fore and mid coxae entirely yellow; hind coxa black with yellow apices in male. Hind tibia white; darkened basally and apically; often with a black subbasal marking. Metasoma entirely black in female. T III sometimes with pair of yellow to brown basal marking in male. T IV sometimes each with transverse yellow basal band in male.	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF9F8B671BBA261A1C5EF8DD.taxon	distribution	Distribution. Japan (Hokkaido, Honshu, and Kyushu). Outside Japan, this species is distributed in Russia (Yu et al. 2016). Bionomics. Host is unknown. In Japan, adults were collected in broad-leaved forest.	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF9F8B671BBA261A1C5EF8DD.taxon	discussion	Remarks. This is the first record of this species from Honshu and Kyushu. Japanese specimens share the character states defined in the original description (Klopfstein 2014 a), while female Japanese specimens collected in Honshu differ from the original description in the following character states: clypeus yellowish-brown to brown (Fig. 5 B) (entirely yellow in the original description); fore and mid femora, tibiae, and tarsomeres brown to blackish-brown (Fig. 5 A) (entirely orange in the original description); hind femur entirely black (Fig. 5 A) (entirely orange in the original description); fore wing length 5.0 – 7.0 mm (5.1 – 5.5 mm in the original description). However, as no significant differences were found in other character states, we conclude that the observed variation is intraspecific variation within a single species.	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF928B671BBA21A51DC1FE4D.taxon	description	(Figs 6 A – G)	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF928B671BBA21A51DC1FE4D.taxon	materials_examined	Specimens examined. JAPAN: [Hokkaido] 1 F, Hidaka Town, Uenzaru-gawa, 10 VII – 1 VIII 2007, A. Ueda leg. (MT) (KPMNH). [Honshu] 1 F, Yamagata Pref., Mamurogawa Town, 3. X. 2009, Y. Matsubara & K. Fukuda leg. (KPMNH); 1 F, Fukushima Pref., Kitakata City, Yamato, 11. VII. 2014, (KPMNH); 1 M, Gunma Pref., Tsumagoi Vil., Kanbara, Takamine-kogen, 3. IX. 2015, K. Watanabe leg. (KPMNH); 1 F, Toyama Pref., Toyama City, Arimine, Inonedani, 15 – 22 IX 2009, M. Watanabe leg. (MT) (KPMNH); 1 M, Toyama Pref., Toyama City, Arimine, Jurodani, 15 – 22 IX 2009, M. Watanabe leg. (MT) (KPMNH); 1 M, Toyama Pref., Toyama City, Kamegai, 1 – 8 IX 2009, M. Watanabe leg. (MT) (KPMNH); 1 F, ditto, 15 – 26 IX 2009, M. Watanabe leg. (MT) (KPMNH); 1 F, Nagano Pref., Otaki Vil., Mt. Ontakesan, Tanohara, 17. VII. 2007, K. Watanabe leg. (KPMNH); 1 M (holotype), Gifu Pref., Mt. Norikura, 29. VII. 1954, Townes family leg. (AEI); 1 F, (paratype), Nagano Pref., Kamikochi, 22. VII. 1954, Townes family leg. (SEHU).	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF928B671BBA21A51DC1FE4D.taxon	diagnosis	Diagnosis. Antenna with 20 – 21 (in female) or 23 (in male) flagellomeres. Tyloids on FL VII to FL XII or FL XIII in male. Face coriaceous and matt; densely punctate. Clypeus depressed apically; robes rounded at outer corners. Mesoscutum coriaceous and matt in female; coriaceous and polished in male; densely punctate in both sexes. Mesopleuron coriaceous and matt; at most with very sparse punctures anteriorly and ventrally in female (Fig. 6 C); smooth and polished; with coriaceous area below episternal scrobe; at most with very sparse punctures anteriorly and ventrally in male. Propodeum coriaceous; posterior part of lateromedian longitudinal carina and pleural carina present. Fore wing length 4.5 – 5.5 mm. Areolet present. Dorsal part of hind coxa coriaceous and matt. Metasoma dorsoventrally depressed in both sexes. Latero-median carina usually indistinct; at most present on basal 0.4 of T I. T I 1.4 – 1.5 × as long as maximum width; strigose laterally; otherwise coriaceous. T II 0.85 – 0.9 × as long as maximum width. Face without pair of yellow markings along each inner orbits (Figs 6 B, F). Face entirely black or black with a brown median marking which separated from clypeus in female (Fig. 6 B); entirely yellow in male (Fig. 6 F). Clypeus yellow with black base in both sexes (Figs 6 B, F). Malar space entirely black in female (Fig. 6 B), entirely yellow in male (Fig. 6 F). Yellow shoulder markings of mesoscutum absent in female (Fig. 6 A); present in male (Fig. 6 E). Scutellum entirely black. All coxae entirely black in female (Fig. 6 A). Fore and mid coxae entirely yellow; hind coxa entirely black in male (Fig. 6 E). Hind tibia whitish-yellow to white; darkened basally and apically; often with a black dorsal stripe and a black subbasal marking. T III with a brown apical marking and T II and T IV often with brown marking in female (Fig. 6 D). T III with pair of yellow to brown basal marking in male (Fig. 6 G). T III and T VI often each with a yellow to brown basal or apical band in male (Fig. 6 G).	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF928B671BBA21A51DC1FE4D.taxon	distribution	Distribution. Japan (Hokkaido and Honshu).	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF928B671BBA21A51DC1FE4D.taxon	biology_ecology	Bionomics. Unknown.	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF928B671BBA21A51DC1FE4D.taxon	discussion	Remarks. This is the first record of this species from Hokkaido.	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF948B601BBA20CD1D2DFD45.taxon	description	(Figs 7 A – C)	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF948B601BBA20CD1D2DFD45.taxon	materials_examined	Type series. Holotype: F, JAPAN, Honshu, Nagano Pref., Otaki Vil., Mt. Ontakesan, 25 VI – 15 VII 2015, S. Shimizu leg. (MT) (KPMNH). Paratypes: JAPAN: [Honshu] 1 F, Toyama Pref., Toyama City, Arimine, Inonedani, 14 – 21 IX 2009, M. Watanabe leg. (MT) (KPMNH); 1 F, Toyama Pref., Nanto City, Togamura, Kamimomose, 15 – 29 IX 2009, M. Watanabe leg. (MT) (KPMNH); 1 F, Ishikawa Pref., Hakusan City, Yahata, 7 – 24 X 2009, H. Fukutomi leg. (MT) (MU).	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF948B601BBA20CD1D2DFD45.taxon	diagnosis	Diagnosis. Antenna with 21 – 23 flagellomeres. Face coriaceous and matt; densely punctate. Clypeus depressed apically; robes weakly angulated at outer corners. Mesoscutum coriaceous and polished; densely punctate. Mesopleuron smooth and polished; densely punctate except for on and below speculum. Propodeum finely rugulose; posterior part of lateromedian longitudinal carina and pleural carina present. Fore wing length 6.5 – 8.5 mm. Areolet present. Dorsal part of hind coxa coriaceous and polished. Metasoma dorsoventrally depressed. Latero-median carina absent. T I 0.9 – 1.1 × as long as maximum width; finely rugulose laterally; otherwise coriaceous. T II 0.6 – 0.7 × as long as maximum width. Face without pair of yellow markings along each inner orbits (Fig. 7 B). Face black with a small yellow median marking which separated from clypeus (Fig. 7 B). Clypeus entirely yellow to brown (Fig. 7 B). Malar space entirely black (Fig. 7 B). Yellow shoulder markings of mesoscutum present (Fig. 7 A). Scutellum black with yellow apical marking. All coxae entirely black (Fig. 7 A). Hind tibia black with white subbasal marking or band (Fig. 7 C). Metasoma entirely black.	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF948B601BBA20CD1D2DFD45.taxon	description	Description. Female (n = 4). Body covered with silverly setae; length 7.5 – 10.5 (holotype: 10.5) mm. Head 0.5 – 0.53 (holotype: 0.52) × as long as wide in dorsal view; coriaceous and densely punctate. Clypeus 1.8 – 2.1 (holotype: 2.1) × as maximum width as high; sparsely punctate; depressed apically; robes weakly angulate at outer corners. Face 1.9 – 2.1 (holotype: 2.0) × as maximum width as high; weakly convex medially in lateral view; separated from clypeus by shallow clypeal sulcus. Malar space 0.66 – 0.75 (holotype: 0.72) × as long as basal mandibular width. POL 1.9 – 2.1 (holotype: 1.9) × as OD. OOL 1.4 – 1.6 (holotype: 1.4) × as OD. POL 1.36 – 1.45 (holotype: 1.36) × as long as OOL. Antenna with 21 – 23 (holotype: 21) flagellomeres. FL I 1.33 – 1.5 (holotype: 1.4) × as long as FL II. Maxillary palp IV 1.0 – 1.1 (holotype: 1.1) × as long as Maxillary palp V. Mesosoma densely punctate. Lateral aspect of pronotum rugulose anteriorly. Mesopleuron smooth below speculum. Sternaulus weakly impressed. Propodeum finely rugulose; posterior part of lateromedian longitudinal carina and pleural carina present. Fore wing length 6.5 – 8.5 (holotype: 8.5) mm. Areolet present. Vein 1 cu-a of fore wing postfurcal or interstitial to vein M & RS. Hind wing with nervellus intercepted below middle. Hind wing with 8 – 10 (holotype: 8) distal hamuli. Dorsal part of hind coxa coriaceous and polished. Hind femur 3.7 – 4.0 (holotype: 4.0) × as long as maximum depth in lateral view. Hind tibia 6.2 – 6.6 (holotype: 6.6) × as long as maximum depth in lateral view. Ratio of length of hind first to fifth tarsomeres 1.0: 0.55 – 0.6 (holotype: 0.6): 0.4: 0.2 – 0.25 (holotype: 0.24): 0.3. Metasoma coriaceous; dorsoventrally depressed. T I rectangular in dorsal view; 0.9 – 1.1 (holotype: 1.05) × as long as maximum width; finely rugulose laterally; latero-median carina absent. T II 0.6 – 0.7 (holotype: 0.66) × as long as maximum width; striate anteriorly. Coloration (Figs 7 A – C). Body (excluding wings and legs) black. Face with small yellow marking medially. Mandible except for apex and base yellow. Clypeus yellow to brown. Antenna blackish-brown. Tegula brown to blackish-brown. Yellow shoulder markings of mesoscutum present. Legs black. Apex of fore and mid trochanters yellow. Fore and mid trochantelli, femora, tibiae, and tarsomeres yellowish-brown to brown. Hind trochanter yellowish-brown apically. Apex of hind trochantellus and base of hind femur reddish-brown. Hind tibia with white subbasal marking or band. Male. Unknown.	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF948B601BBA20CD1D2DFD45.taxon	distribution	Distribution. Japan (Honshu). Bionomics. Unknown.	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF948B601BBA20CD1D2DFD45.taxon	etymology	Etymology. The species name is from Japan.	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF948B601BBA20CD1D2DFD45.taxon	discussion	Remarks. This species resembles H. sundevalli in the mesopleuron with dense punctures except for on and below speculum, the yellow shoulder markings present, the scutellum with yellow marking, and the entirely black metasoma, but can be distinguished from the latter by the following combination of character states: propodeum finely rugulose (coarsely rugulose in H. sundevalli); latero-median carina of T I absent (present in H. sundevalli); T I 0.9 – 1.1 × as long as maximum width (1.1 – 1.4 in H. sundevalli).	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF958B621BBA24DD1B28F815.taxon	description	(Figs 8 A, B)	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF958B621BBA24DD1B28F815.taxon	materials_examined	Specimens examined. JAPAN: [Hokkaido] 1 M, Kamikawa Town, Sounkyo, Ginsendai 8 – 9 IX 2014, S. Shimizu leg. (KPMNH).	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF958B621BBA24DD1B28F815.taxon	diagnosis	Diagnosis. Antenna with 23 (in female) or 25 (in male) flagellomeres. Tyloids absent in male. Face coriaceous and matt; sparsely punctate. Clypeus depressed apically; robes rounded at outer corners. Mesoscutum coriaceous and matt; densely punctate. Mesopleuron coriaceous and matt; densely punctate anteriorly and ventrally. Propodeum finely rugulose; lateral longitudinal and pleural carina present; petiolar area surrounded by carina (Fig. 1 C). Fore wing length 7.8 (in female) or 8.0 (in male) mm. Areolet present. Dorsal part of hind coxa coriaceous and matt. Metasoma dorsoventrally depressed in both sexes. Latero-median carina present on basal 0.8 of T I. T I 2.0 (in female) or 1.8 (in male) × as long as maximum width; strigose anteriorly; otherwise coriaceous. T II 1.3 (in female) or 1.0 (in male) × as long as maximum width. Face black with pair of yellow markings along each inner orbit (Fig. 12 B). Clypeus entirely yellow in both sexes (Fig. 8 B). Malar space entirely black in both sexes (Fig. 8 B). Yellow shoulder markings of mesoscutum present (Fig. 8 A). Scutellum entirely yellow. All coxae entirely black in females. Fore coxa black with yellow apex, mid and hind coxae entirely black in males (Fig. 8 A). Hind tibia yellow, darkened apically (Fig. 8 A). Metasoma entirely black in both sexes.	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF958B621BBA24DD1B28F815.taxon	description	Description based on Japanese specimen. Male (n = 1). Body covered with silverly setae; length 10.0 mm. Head 0.51 × as long as wide in dorsal view; coriaceous; sparsely punctate. Clypeus 2.0 × as maximum width as high; depressed apically; robes rounded at outer corners. Face 2.3 × as maximum width as high; weakly convex medially in lateral view; separated from clypeus by shallow clypeal sulcus. Malar space 1.0 × as long as basal mandibular width. POL 1.55 × as OD. OOL 1.15 × as OD. POL 1.40 × as long as OOL. Antenna with 25 flagellomeres. Tyloids absent. FL I 1.2 × as long as FL II. Maxillary palp IV 1.3 × as long as Maxillary palp V. Mesosoma densely punctate. Lateral aspect of pronotum coarsely rugulose anteriorly and ventrally. Mesopleuron smooth below speculum. Sternaulus weakly impressed. Propodeum finely rugulose; lateral longitudinal and pleural carina present; petiolar area surrounded by carina (Fig. 1 C). Fore wing length 8.0 mm. Areolet present. Vein 1 cu-a of fore wing postfurcal to vein M & RS. Hind wing with nervellus intercepted below middle. Hind wing with 9 distal hamuli. Dorsal part of hind coxa coriaceous and matt. Hind femur 6.25 × as long as maximum depth in lateral view. Hind tibia 8.3 × as long as maximum depth in lateral view. Ratio of length of hind first to fifth tarsomeres 1.0: 0.7: 0.4: 0.2: 0.3. Metasoma coriaceous; dorsoventrally depressed. T I rectangular in dorsal view; 1.8 × as long as maximum width; strigose anteriorly; otherwise coriaceous; latero-median carina present on basal 0.8. T II 0.95 × as long as maximum width; strigose anteriorly. Coloration (Figs 8 A, B). Body (excluding wings and legs) black. Face with pair of yellow markings along each inner orbits. Palpi, mandible except for apex, subtegular ridge, tegula, upper mesepisternum, shoulder markings of mesoscutum, and scutellum yellow. Legs black. Apex of fore coxa, trochanters, trochantelli, and base of hind tibia yellow. Fore and mid femora, tibiae, and tarsomeres yellowish-brown to brown. Base and apex of hind femur brown.	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF958B621BBA24DD1B28F815.taxon	distribution	Distribution. Japan (Hokkaido). Outside Japan, this species is distributed in Sweden (Johansson 2020).	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF958B621BBA24DD1B28F815.taxon	biology_ecology	Bionomics. In Japan, adults were collected in broad-leaved forest at altitudes of ca. 1,000 meters.	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF958B621BBA24DD1B28F815.taxon	discussion	Remarks. This is the first record of the male of this species and the first record of this species from Japan.	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF978B621BBA21ED1F90FC19.taxon	description	(Figs 9 A – D)	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF978B621BBA21ED1F90FC19.taxon	materials_examined	Specimens examined. JAPAN: [Honshu] 3 F, Toyama Pref., Toyama City, Arimine, Inonedani, 15 – 22 IX 2009, M. Watanabe leg. (MsT) (KPMNH); 1 F, ditto, 1 – 8 IX 2009, M. Watanabe leg. (MsT) (KPMNH); 1 F (holotype), Nagano Pref., Kamikochi, 5. VII. 1954, Townes Family leg. (SEHU); 1 F, Nagano Pref., Otaki Vil., Mt. Ontakesan, Hakkaisan, 6. VIII. 2010, K. Watanabe leg. (KPNMH); 1 F, Nagano Pref., Otaki Vil., Mt. Ontakesan, Tanohara, 9. VIII. 2007, K. Watanabe leg. (KPNMH); 1 F, Nagano Pref., Otaki Vil., Mt. Ontakesan, 25 VI – 15 VII 2015, S. Shimizu leg. (MsT) (KPMNH).	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF978B621BBA21ED1F90FC19.taxon	diagnosis	Diagnosis. Antenna with 19 – 20 flagellomeres. Face coriaceous and matt; weakly punctate medially. Clypeus depressed apically; robes rounded at outer corners. Mesoscutum coriaceous and polished; densely punctate. Mesopleuron smooth and polished; often with coriaceous area below speculum; densely punctate except for on and below speculum (Fig. 9 C). Propodeum finely rugulose; posterior part of lateral longitudinal and lateromedian longitudinal carina and pleural carina present. Fore wing length 4.5 – 6.0 mm. Areolet present. Dorsal part of hind coxa coriaceous and matt. Metasoma dorsoventrally depressed in both sexes. T I 1.05 – 1.25 × as long as maximum width; strigose anteriorly and laterally; otherwise coriaceous. Latero-median carina usually indistinct; at most present on basal 0.3 of T I. T II 0.85 – 1.1 × as long as maximum width. Face without pair of yellow markings along each inner orbits (Fig. 9 B). Face black with yellow median marking which separated from clypeus (Fig. 9 B). Clypeus entirely yellow (Fig. 9 B). Malar space entirely black (Fig. 9 B). Yellow shoulder markings of mesoscutum present (Fig. 9 A). Scutellum yellow with black basomedian marking. All coxae entirely black (Fig. 9 A). Hind tibia white, darkened basally and apically, often with a black subbasal marking (Fig. 9 D). Metasoma entirely black.	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF978B621BBA21ED1F90FC19.taxon	distribution	Distribution. Japan (Honshu).	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF978B621BBA21ED1F90FC19.taxon	biology_ecology	Bionomics. Unknown.	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF978B7F1BBA25EE1D32FBD9.taxon	description	(Figs 10 A – E)	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF978B7F1BBA25EE1D32FBD9.taxon	materials_examined	Specimens examined. JAPAN: [Hokkaido] 1 F, Hidaka Town, Uenzaru-gawa, 10 VII – 1 VIII 2007, A. Ueda leg. (MsT) (KPMNH); 1 F, Kamishihoro Town, Nukabira, 22. VI. 1980, H. Takemoto leg. (NARO); 1 M (holotype of S. tenuitibialis), Sapporo, 30. V. 1956, K. Kamijo leg. (SEHU); 1 F, Soranuma, 17. VIII. 1967, K. Kusigemati leg. (SEHU); [Honshu] 2 M, Yamagata Pref., Mamurogawa Town, 22. VI. 2012, Y. Matsubara & K. Fukuda leg. (KPMNH); 1 F, Yamagata Pref., Mt. Asahidake, 27. VII. 1969, H. Takizawa leg. (SEHU); 1 F, Tochigi Pref., Nasushiobara City, Kotaki, 24 – 30 VII 2008, T. Matsumura leg. (MsT) (KPMNH); 1 M, Gunma Pref., Katashina Vil., Mt. Hotakasan, 2. VIII. 2007, K. Watanabe leg. (KPNMH); 2 M, Niigata Pref., Nagaoka City, Suyoshi, Mt. Nokogiriyama, 7. VI. 2014, S. Shimizu leg. (KPNMH); 2 F, Niigata Pref., Sado Is., Kanaishinbo, Hakuundai, Mt. Myokenzan, 4. VIII. 2009, K. Watanabe leg. (KPNMH); 1 F, Yamanashi Pref., Koshu City, Yanagisawa-toge, 5. VIII. 2008, K. Watanabe leg. (KPNMH); 1 F, Yamanashi Pref., Hokuto City, Masutomi, Biwakubosawa, 24. VI. 2007, K. Watanabe leg. (KPNMH); 1 F, ditto, 24. VI. 2007, T. Ban leg. (KPNMH); 1 M, Yamanashi Pref., Koshu City, Sagashio, 17. V. 2008, T. Muraki leg. (KPNMH); 1 M, Kanagawa Pref., Mt. Omuroyama, 7. VI. 2013, (FIT) (KPMNH); 1 M, Kanagawa Pref., Hadano City, Tanzawaterayama, Yabitsu-toge, Mt. Oyama, I. Waki leg. (KPNMH); 1 M, Kanagawa Pref., Hadano City, Mt. Koubouyama, 5. IV. 2007, K. Watanabe leg. (KPNMH); 1 F 1 M, ditto, 6. IV. 2008, K. Watanabe leg. (KPMNH); 1 F 1 M, ditto, 12. IV. 2008, K. Watanabe leg. (KPMNH); 1 M, Kanagawa Pref., Odawara City, Kayama, Sakawagawa, 25. III. 2017, K. Watanabe leg. (KPNMH); 1 M, Kanagawa Pref., Hinatayakushi, 29. VI. 1974, H. Takizawa leg. (SEHU); 4 M, Shizuoka Pref., Honkawane Town, Mt. Yamainudan, 14. VI. 2008, K. Watanabe leg. (KPNMH); 1 F, Nagano Pref., Kawakami Vil., Azusayama, 14. VI. 2015, K. Watanabe leg. (KPNMH); 1 M, Nagano Pref., Otaki Vil., Mt. Ontakesan, Hakkaisan, 5. VIII. 2010, K. Watanabe leg. (KPNMH); 2 M, ditto, 6. VIII. 2010, K. Watanabe leg. (KPNMH); 1 M, ditto, 7. VIII. 2010, K. Watanabe leg. (KPNMH); 1 F, ditto, 8. VIII. 2010, K. Watanabe leg. (KPNMH); 1 F, Nagano Pref., Otaki Vil., Mt. Ontakesan, Tanohara, 17. VII. 2007, K. Watanabe leg. (KPNMH); 2 M, ditto, 9. VIII. 2007, K. Watanabe leg. (KPNMH); 1 F, Toyama Pref., Toyama City, Arimine, jurodani, 7 – 14 VII 2009, M. Watanabe leg. (MsT) (KPMNH); 1 F, Toyama Pref., Toyama City, Arimine, Inonedani, 25 VIII – 1 IX 2009, M. Watanabe leg. (MsT) (KPMNH); 1 F, Toyama Pref., Toyama City, Kamegai, 15 – 26 IX 2009, M. Watanabe leg. (MsT) (KPMNH); 1 F, Aichi Pref., Shitara Town, Tsuguhonsawa, 26 V – 23 VI 2020, S. Morishita leg. (TMNH); 1 F, Aichi Pref., Toyohashi City, Imure-cho, Takayama, 21. V. 2019, S. Morishita leg. (TMNH); 1 F, Aichi Pref., Toyohashi City, Suse-cho, Kanbata, 23. X. 2018, S. Morishita leg. (TMNH); 1 M, Fukui Pref., Ikeda Town, Mizuumi, Mt. Hekosan, 18. VI. 2016, S. Shimizu leg. (KPNMH); 1 F, Hyogo Pref., Sayo Town, Funakoshi, 28 III – 4 IV 2010, (KPNMH); [Izu Isis.] 1 F (holotype of H. momoii), Tokyo, Izuoshima Is., 25. V. 1955, S. Momoi leg. (SEHU); [Shikoku] 1 M, Kochi Pref., Umaji Vil., 24. VI. 2010, Y. Matsubara & K. Fukuda leg. (MsT) (KPNMH); [Kyushu] 1 F, Kagoshima Pref., Mt. Kirishima, 27. V. 1966, K. Kusigemati leg. (SEHU); 1 F, Kagoshima Pref., Mt. Kurinodake, 10. V. 1967, K. Kusigemati leg. (SEHU). GERMANY: 1 M (det. S. Klopfstein), Oberhartz, Clausthal, 26. XI. 1928, G. Meyer leg. (ZSM).	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF978B7F1BBA25EE1D32FBD9.taxon	diagnosis	Diagnosis. See Klopfstein (2014 b).	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF978B7F1BBA25EE1D32FBD9.taxon	distribution	Distribution. Japan (Hokkaido, Honshu, Izuoshima Is., Shikoku, and Kyushu). Outside Japan, this species is widely distributed in Palearctic region (Yu et al. 2016).	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF978B7F1BBA25EE1D32FBD9.taxon	biology_ecology	Bionomics. Host is unknown. In Japan, adults were collected in broad-leaved forest.	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF978B7F1BBA25EE1D32FBD9.taxon	discussion	Remarks. This is the first record of this species from Japan.	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF8A8B7F1BBA22A11A7FFFFD.taxon	description	(Figs 11 A – G)	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF8A8B7F1BBA22A11A7FFFFD.taxon	materials_examined	Specimens examined. JAPAN: [Hokkaido] 2 F, Horokanai Town, Uryu-kenkyurin, Butokamanbetsu-path, 11 – 17 VII 2012, K. Watanabe leg. (MT) (KPNMH); 1 F, dotto, 16. VII. 2012, M. Ito leg. (KPNMH); 1 F, Chitose City, Kusabue-rindo, 8 – 21 VII 2012, N. Kuhara leg. (MsT) (EUM); 1 F, Nissho-toge, Sarugawa-river, 31 VII – 14 VIII 2010, N. Kuhara leg. (MsT) (EUM). [Honshu] 1 F, Toyama Pref., Toyama City, Arimine, Inonedani, 15 – 22 IX 2009, M. Watanabe leg. (MsT) (KPMNH); 1 F, Shizuoka Pref., Umegashima, 3 VI 2001, T. Sugiyama leg. (MU); 1 F, Aichi Pref., Shitara Town, Uradani, 27 VI – 3 VII 1994, K. Yamagishi leg. (MU). GERMANY: 1 M (det. E. Diller), Celle, 7. VI. 1964, (ZSM). NETHERLANDS: 1 F (det. C. J. Zwakhals), Tongeren, 16. VI. 1986, C. J. Zwakhals leg. (ZSM).	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF8A8B7F1BBA22A11A7FFFFD.taxon	diagnosis	Diagnosis. See Klopfstein (2014 b).	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF8A8B7F1BBA22A11A7FFFFD.taxon	distribution	Distribution. Japan (Etorofu Is., Shikotan Is., Hokkaido, and Honshu). Outside Japan, this species is widely distributed in Holarctic region and also recorded from Mexico (Yu et al. 2016).	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF8A8B7F1BBA22A11A7FFFFD.taxon	biology_ecology	Bionomics. Host unknown in Japan. Outside of Japan, Platycheirus scutatus (Meigen 1822) (Diptera, Syrphidae) has been recorded as a host (Thirion 1994).	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF8A8B7F1BBA22A11A7FFFFD.taxon	discussion	Remarks. This is the first record of this species from Hokkaido and Honshu.	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF8A8B781BBA26851DDAFD5D.taxon	description	(Figs 12 A – F)	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF8A8B781BBA26851DDAFD5D.taxon	materials_examined	Specimens examined. JAPAN: [Hokkaido] 7 F 10 M, Akkeshi Town, Bekanbeushi marsh, 14. VII. 2003, R. Matsumoto leg. (OMNH); 5 F, ditto, 12 – 31 VII 2003, R. Matsumoto leg. (MsT) (OMNH); 1 M, ditto, 1. VIII. 2003, R. Matsumoto leg. (OMNH); 5 F, ditto, 1 – 30 VIII 2003, R. Matsumoto leg. (MsT) (OMNH); 11 F, ditto, 1 – 31 VIII 2003, R. Matsumoto leg. (MsT) (OMNH); 2 F 10 M, ditto, 30. VIII. 2003, R. Matsumoto leg. (OMNH); 1 F, Nukabira, 1. VII. 1980, K. Maeto leg. (NARO); 1 M, Toyokoro Town, Horooka, 17. VI. 2006, K. Watanabe leg. (KPNMH); 1 F, Nissho-toge, Sarugawa-river, 31 VII – 14 VIII 2010, N. Kuhara leg. (MsT) (EUM); [Honshu] 1 M, Nagano Pref., Otaki Vil., Mt. Ontakesan, 13 – 25 VI 2015, S. Shimizu leg. (KPNMH). GERMANY: 1 F (det. E. Diller), Northeim, 8. VIII. 1945, (ZSM); 1 M (det. E. Diller), Stolzenau, 8. VIII. 1945, (ZSM). NETHERLANDS: 1 F (det. C. J. Zwakhals), St. Pietersberg, 15. VIII. 1986, C. J. Zwakhals leg. (KPNMH); 1 F (det. C. J. Zwakhals), Asperen, 12. VIII. 1973, C. J. Zwakhals leg. (KPNMH).	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF8A8B781BBA26851DDAFD5D.taxon	diagnosis	Diagnosis. See Klopfstein (2014 b).	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF8A8B781BBA26851DDAFD5D.taxon	distribution	Distribution. Japan (Etorofu Is., Kunashiri Is., Hokkaido, and Honshu). Outside Japan, this species is widely distributed in Holarctic region (Yu et al. 2016).	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF8A8B781BBA26851DDAFD5D.taxon	biology_ecology	Bionomics. In Japan, Syrphus vitripennis Meigen, 1822 (Diptera, Syrphidae) have been recorded as host (Uchida 1930, 1957). Outside Japan, nine hoverfly species, Epistrophe eligans (Harris, 1780), Neocnemodon vitripennis (Meigen, 1822), Platycheirus fulviventris (Macquart, 1829), Scaeva pyrastri (Linnaeus, 1758), Sphaerophoria cylindrica Say, 1824, Sphaerophoria rueppellii (Wiedemann, 1820), Sphaerophoria scripta (Linnaeus, 1758), Syrphus ornatus Meigen, 1822, Syrphus torvus Osten Sacken, 1875 have been recorded as hosts (Rudow 1908; Rudow 1919; Scott 1939; Dasch 1964; Thirion 1994; Vidal 1997). Adoxophyes orana (Fischer von Röslerstamm, 1834) (Lepidoptera, Tortricidae) and Delia radicum (Linnaeus, 1758) (Diptera, Anthomyiidae) have also been recorded as hosts (Janssen 1958; Constantineanu & Mustata 1973; Mustata 1978; Pisica 1983), but are considered doubtful records.	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF8A8B781BBA26851DDAFD5D.taxon	discussion	Remarks. This is the first record of this species from Honshu.	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF8D8B7A1BBA24DA1DD0F8F9.taxon	description	(Figs 13 A – E)	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF8D8B7A1BBA24DA1DD0F8F9.taxon	materials_examined	Specimens examined. JAPAN: [Honshu] 1 M, Gunma Pref., Tsumagoi Vil., Kanbara, Takamine-kogen, 3. IX. 2015, K. Watanabe leg. (KPMNH); 1 M, Niigata Pref., Nagaoka City, Suyoshi, Mt. Nokogiriyama, 7. VI. 2014, S. Shimizu leg. (KPMNH); 1 M, Toyama Pref., Toyama City, Arimine, Inonedani, 15 – 22 IX 2009, M. Watanabe leg. (MT) (KPMNH); 1 M, Nagano Pref., Otaki Vil., Mt. Ontakesan, Hakkaisan, 5 IX 2022, S. Morishita leg. (TMNH); 1 F (holotype), Gifu Pref., Mt. Norikura, 29. VII. 1954, Townes family leg. (AEI); 1 M, Gifu Pref., Hirayu, 12. X. 1982, K. Konishi leg. (NARO). AUSTRIA: 1 F (det. E. Diller), Oberosterreich, Stallburgalm, 4. VIII. 1986, M. Schwarz leg. (ZSM). GERMANY: 1 M (det. E. Diller), Goslar, 18. VIII. 1948, E. Bauer leg. (ZSM).	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF8D8B7A1BBA24DA1DD0F8F9.taxon	diagnosis	Diagnosis. See Klopfstein (2014 b).	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF8D8B7A1BBA24DA1DD0F8F9.taxon	distribution	Distribution. Japan (Honshu). Outside Japan, this species is widely distributed in Palearctic region (Yu et al. 2016).	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF8D8B7A1BBA24DA1DD0F8F9.taxon	biology_ecology	Bionomics. Host unknown in Japan. Outside of Japan, Epistrophe grossulariae (Meigen 1822) (Diptera, Syrphidae) has been recorded as a host (Rotheray 1986; Thirion 1994). In Japan, adults were collected in broadleaved forests.	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF8D8B7A1BBA24DA1DD0F8F9.taxon	discussion	Remarks. This is the first record of this species from Japan. Japanese specimens share similar character states with the diagnosis of European specimens proposed by Klopfstein (2014 b), while female Japanese specimen (holotype of H. enizemopsis) differs from European specimens in the following character states: fore coxa entirely black (black with yellow apex in European specimens); mid and hind trochanters, trochantelli, and femora entirely black (orange in European specimens); hind tibia entirely black (yellow or orange with darknened apex in European specimens). However, as no significant differences were found in other character states, we conclude that the observed variation is intraspecific variation within a single species.	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF8F8B751BBA25451D6FF86D.taxon	type_taxon	Type species: Bassus exsultans Gravenhorst, 1829. Designated by Viereck (1914).	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF8F8B751BBA25451D6FF86D.taxon	diagnosis	Diagnosis. According to Klopfstein (2014 b), this genus can be distinguished from other genera by the following combination of character states. Face coriaceous and matt; without vertical impressions; entirely black or with yellow central marking in female; entirely yellow in male. Clypeus with thin apical margin; with basal elevation and impressed only laterally; resulting in central area being convex and sides being concave. Antenna with apical flagellomeres longer than wide; with tyloids in male; tyloids linear and narrow and without long setae. Mesoscutum without notaulus; coriaceous and matt with inconspicuous but sometimes dense punctures; yellow shoulder markings usually present. Scutellum only carinate basally. Mesopleuron either entirely coriaceous and matt or rather smooth, especially on upper half; never strongly punctate. Epicnemial carina interrupted behind fore coxa. Propodeum with carinae reduced; at most with some traces of lateral carinae. Propodeal spiracle inconspicuous. Areolet absent; very rarely present as aberration in S. tarsatorius. Hind wing with 2 – 3 basal hamuli. Hind tibia white, yellow or orange with apex dark. Female metasoma evenly tapered to apex; without transverse impressions. T I without latero-median carina. T II with spiracle dorsal, above lateral fold. T III with spiracle below or behind the fold. Metasoma black; often with yellow apical bands on most tergites (Fig. 14 B), or orange on median tergites. Ovipositor sheaths 0.3 × as long as hind tibia; rather stout; laterally compressed; slightly tapered and fully enclosing ovipositor or diagonally truncate; with inconspicuous setae ventrally and apically. T VIII and T IX of males present as separate sclerites. Hypopygium about 2.0 × wider than long, emarginated apically and forming two lobes; outer corners of lobes rounded.	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF8F8B751BBA25451D6FF86D.taxon	distribution	Distribution. Afrotropical, Australasian, Holarctic, Neotropical, and Oriental regions (Yu et al., 2016).	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF8F8B751BBA25451D6FF86D.taxon	discussion	Remarks. Two species, H. atamiensis and H. tarsatorius, are known in Japan. The former is described based on incomplete descriptions based on damaged holotype. Both species are currently unseparable, and they are likely to be the same species, but taxonomic treatment is reserved.	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF808B751BBA21341C78FBCD.taxon	discussion	Remarks. The holotype of Syrphoctonus atamiensis Ashmead, 1906 has no mesosoma or wings. The original description only provides information on body length and colouration. Thus, generic identification of this species is impossible. Therefore, we treat this taxon as incertae sedis.	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF808B751BBA21341C78FBCD.taxon	distribution	Distribution. Japan (Honshu).	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF808B741BBA23541A56FB11.taxon	description	(Figs 22 A, B)	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF808B741BBA23541A56FB11.taxon	materials_examined	Specimens examined. JAPAN: [Honshu] 1 F, Ibaraki Pref., Goka, 12. IV. 1999, M. Uchida leg. (KPMNH); 1 F 1 M, Saitama Pref., Urawa, Tajima, 9. IV. 1999, T. Nambu leg. (KPMNH); 2 M, Tokyo, Akiruno City, Ninomiya, Tamagawariverside, 3. III. 2010, K. Watanabe leg. (KPMNH); 1 M, Tokyo, Takao, Hikagesawa, Kokesawa, 3. III. 2007, K. Watanabe leg. (KPMNH); 1 F, Kanagawa Pref., Ayase City, Yoshioka, Irigaya, 8. IV. 2000, I. Waki leg. (KPMNH); 1 F, Kanagawa Pref., Ebina City, Sagamigawa, 30. IV. 2006, M. Oishi & R. Watanabe leg. (KPMNH); 1 M, Kanagawa Pref., Atsugi City, 25. IV. 2008, A. Sakai leg. (KPMNH); 1 F, Kanagawa Pref., Atsugi City, Nakaogino, 26. IV. 2008, M. Gunji leg. (KPMNH); 1 M, Kanagawa Pref., Atsugi City, Sanda, 26. IV. 2008, H. Katahira leg. (KPMNH); 2 M, Kanagawa Pref., Hiratsuka City, Mt. Komayama, Shonandaira, 21. IV. 2007, K. Watanabe leg. (KPMNH); 1 M, Kanagawa Pref., Miyamae-ku, Sugao-kouen, 26. V. 2001, T. Ishizaki leg. (KPMNH); 1 F, Kanagawa Pref., Hadano City, Mt. Koubouyama, 5. IV. 2007, K. Watanabe leg. (KPMNH); 1 M, ditto, 15. IV. 2007, K. Watanabe leg. (KPMNH); 1 F, ditto, 29. IV. 2007, K. Watanabe leg. (KPMNH); 1 M, ditto, 6. IV. 2008, K. Watanabe leg. (KPMNH); 1 F 1 M, ditto, 12. IV. 2008, K. Watanabe leg. (KPMNH); 1 F 4 M, ditto, 18. IV. 2010, K. Watanabe leg. (KPMNH); 1 M, Shizuoka Pref., Higashiizu Town, Inatori, 19. III. 2009, T. Muraki leg. (KPMNH); 3 F, Nagano Pref., Otaki Vil., Mt. Ontakesan, Hakkaisan, 5. VIII. 2010, K. Watanabe leg. (KPNMH); 9 F, ditto, 6. VIII. 2010, K. Watanabe leg. (KPNMH); 1 F, ditto, 7. VIII. 2010, K. Watanabe leg. (KPNMH); 6 F, ditto, 8. VIII. 2010, K. Watanabe leg. (KPNMH); 33 F 2 M, ditto, 5 – 9 VIII 2010, K. Watanabe leg. (KPNMH); [Amami Isls.] 1 M, Kagoshima Pref., Tokunoshima Is., Isen-cho, Nakayama, 21. III. 2008, K. Watanabe leg. (KPNMH).	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF808B741BBA23541A56FB11.taxon	diagnosis	Diagnosis. See Klopfstein (2014 b).	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF808B741BBA23541A56FB11.taxon	distribution	Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu, Hachijojima Is., Izuoshima Is., Amamioshima Is., and Tokunoshima Is.). Outside Japan, this species is widely distributed in Holarctic region (Yu et al. 2016).	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF808B741BBA23541A56FB11.taxon	biology_ecology	Bionomics. This species was collected by light trap (Yang et al. 2006). In Japan, adults were collected in broadleaved forests and grasslands. In Japan, three hoverfly species, Epistrophe omogensis (Siraki & Edashige, 1953), Metasyrphus frequens Matsumura, 1917, Syrphus vitripennis Meigen, 1822 have been recorded as host (Uchida 1957; Minamikawa 1969). Outside Japan, 15 hoverfly species, Epistrophe euchroma (Kowarz, 1885), Episyrphus balteatus (De Geer, 1776), Eupeodes corollae (Fabricius, 1794), Eupeodes luniger (Meigen, 1822), Eupeodes nitens (Zetterstedt, 1843), Melanostoma mellinum (Linnaeus, 1758), Neocnemodon vitripennis (Meigen, 1822), Paragus quadrifasciatus Meigen, 1822, Platycheirus albimanus (Fabricius, 1781), Platycheirus scutatus (Meigen, 1822), Scaeva pyrastri (Linnaeus, 1758), Sphaerophoria scripta (Linnaeus, 1758), Dasysyrphus venustus (Meigen, 1822), Syrphus ribesii (Linnaeus, 1758), Syrphus torvus Osten Sacken, 1875 have been recorded as hosts (Cameron 1913; Coulon 1934; Dusek, Laska & Sedivy 1979; Goidanich 1928; Hedwig 1950; Horstmann 1970; Kettner 1970; Ngamo, Delabye, Stilmant & Hance 1998; Pisica & Petcu 1972; Pisica 1983; Pschorn-Walcher & Zwölfer 1956; Salt 1936; Scott 1939; Thirion 1994; Wnuk 1974). Anthonomus pomorum (Linnaeus, 1758) (Coleoptera, Curculionidae) and Coccinella septempunctata Linnaeus, 1758 (Coleoptera, Coccinellinae) have also been recorded as hosts (Keler 1940; Schaefer & Semyanov 1992), but are considered doubtful records.	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
03D487FDFF808B741BBA23541A56FB11.taxon	discussion	Remarks. This is the first record of this species from Tokunoshima Island.	en	Morishita, Shunsuke, Watanabe, Kyohei (2025): Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan. Zootaxa 5588 (1): 49-76, DOI: 10.11646/zootaxa.5588.1.2, URL: https://doi.org/10.11646/zootaxa.5588.1.2
