identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03D6AC50FF8E8F6A0AF79D64CE53FD7C.text	03D6AC50FF8E8F6A0AF79D64CE53FD7C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Limnonectes maanyanorum Gonggoli & Shimada & Matsui & Nishikawa & Sidik & Kadafi & Farajallah & Hamidy 2025	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Limnonectes maanyanorum sp. nov.</p>
            <p>(Figs. 5–6)</p>
            <p>
                 Limnonectes kuhlii “Lineage 3”: Matsui et al. (2016)   Holotype.  
                <a title="Search Plazi for locations around (long 115.303406/lat -1.915357)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=115.303406&amp;materialsCitation.latitude=-1.915357">Adult</a>
                 female (MZB Amph 33530) from  
                <a title="Search Plazi for locations around (long 115.303406/lat -1.915357)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=115.303406&amp;materialsCitation.latitude=-1.915357">Mekang River</a>
                 ,  
                <a title="Search Plazi for locations around (long 115.303406/lat -1.915357)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=115.303406&amp;materialsCitation.latitude=-1.915357">Mt. Karasik</a>
                 ,  
                <a title="Search Plazi for locations around (long 115.303406/lat -1.915357)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=115.303406&amp;materialsCitation.latitude=-1.915357">Apar Batu Village</a>
                 , Awang District, Barito Timur Regency, Central Kalimantan Province (Borneo), Indonesia at - 1.915357°S, 115.303402°E, elevation 91 m a.s.l., collected by Ade Damara Gonggoli and Adventus Robertino Rangin on 12 January 2023. 
            </p>
            <p>
                  Paratypes (n = 10).  
                <a title="Search Plazi for locations around (long 115.308815/lat -1.918157)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=115.308815&amp;materialsCitation.latitude=-1.918157">Total</a>
                 of eight specimens from  
                <a title="Search Plazi for locations around (long 115.308815/lat -1.918157)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=115.308815&amp;materialsCitation.latitude=-1.918157">Mekang River</a>
                 ,  
                <a title="Search Plazi for locations around (long 115.308815/lat -1.918157)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=115.308815&amp;materialsCitation.latitude=-1.918157">Mt. Karasik</a>
                 ,  
                <a title="Search Plazi for locations around (long 115.308815/lat -1.918157)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=115.308815&amp;materialsCitation.latitude=-1.918157">Apar Batu Village</a>
                 , Awang District, Barito Timur Regency, Central Kalimantan Province (Borneo), Indonesia at - 1.918157°S, 115.308813°E, elevation 164 m a.s.l.: one adult male (MZB Amph 33514) collected with holotype; seven adult females (MZB Amph 33513, 33515, 33531–35) collected by Ade Damara Gonggoli, Adventus Robertino Rangin, Tendang, and Pa Nika on 22 February 2022  . 
            </p>
            <p>
                  Two specimens from  
                <a title="Search Plazi for locations around (long 116.9928/lat -1.05225)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=116.9928&amp;materialsCitation.latitude=-1.05225">Margomulyo Village</a>
                 , Samboja District, Kutai Kertanegara Regency, East Kalimantan Province (Borneo), Indonesia at - 1.052250°S, 116.992798°E, elevation 68.3 m a.s.l.  :  two juveniles (MZB Amph 15264–65) collected by Amir Hamidy and Abran Slamet on 18 February 2009.
            </p>
            <p> Etymology. The species epithet  maanyanorum refers to the name of the local ethnic, Dayak Maanyan, who live in Central Kalimantan, from where the species is found. The species epithet is in genitive plural, meaning of the “Maanyan”. </p>
            <p>Local common name. On Mt. Karasik, the local people of the Dayak Maanyan ethnic call this frog “Senteleng Watu”, which means “Katak Batu” in Indonesia and “Rock Frog” in English.</p>
            <p>Suggested English common name. Maanyan Creek Frog.</p>
            <p>Suggested Indonesian common name. Bangkong-tuli Maanyan.</p>
            <p> Diagnosis. The new species clustered within the  L. kuhlii complex in molecular analyses and can be distinguished from its congeners by the following combination of morphological characters: medium size (SVL 45.9 mm in the only adult male, 40.4–58.5 mm in adult females); toes fully webbed; tympanum hidden; dorsal skin of body rugose; dorsal skin of tibia densely covered by small warts and coarsely scattered with well-elevated large tubercles; flap of skin on outer side of third finger well-developed, moveable; inner and outer webbing of fourth toe fairly well excised; disk of third finger not much wider than basal phalanx; disk of fourth toe wider than basal phalanx; male nuptial pad on first finger without minute asperities; head longer than wide; throat covered with confluent irregular dark gray blotches. </p>
            <p>Description of the holotype (measurements in mm). Adult female (SVL 57.0), habitus stocky; head longer (HL 24.1, 42.3 % SVL) than wide (HW 23.0, 40.3 % SVL); rostrum longer (RL 9.3, 16.4 % SVL) than eye (ED 5.6, 9.9 % SVL); nostril dorsolateral on canthus, closer to snout tip (SL 4.0, 7.0 % SVL) than to eye (N-EL 5.4, 9.4 % SVL); internarial distance (IND 4.6, 8.0 % SVL) longer than interorbital distance (IOD 4.2, 7.3 % SVL); snout obtusely pointed in lateral view; canthus rounded; lore concave; pineal spot distinct; tympanum indistinct; pair of odontoid processes on lower jaw, obtuse in profile; vomerine teeth in oblique groups, groups separated from each other and from choana by half length of one group; vocal sac and vocal slits absent.</p>
            <p>Forelimb stout, relatively short (FLL 28.0, 49.2 % SVL); fingers moderately slender; finger length formula II &lt;I &lt;IV &lt;III; finger webbing absent; finger tips bluntly rounded, without circummarginal grooves; disk of third finger (3TDW 1.0, 1.7 % SVL) not much wider than basal phalanx; inner palmar tubercle oval and low (IPTL 3.3, 5.7 % SVL); middle palmar tubercle oval and low; outer palmar tubercle slightly oval and elongated, contacting with middle palmar tubercle; proximal subarticular tubercle slightly rounded, elevated; distal subarticular tubercle indistinct; skin flaps on inner and outer side of second and third fingers present, more developed on outer side of third finger than inner, freely moveable; supernumerary metacarpal tubercles absent.</p>
            <p>Hindlimb stout, relatively short (HLL 82.0, 143.9 % SVL), about three times length of forelimb, toe length formula I &lt;II &lt;V &lt;III &lt;IV; foot (FL 25.8, 45.3 % SVL) slightly longer than tibia (TL 25.5, 44.7 % SVL); tibiotarsal articulation reaching between eye and nostril in preserved condition; toes tips blunt, rounded; disk of fourth toe (4TDW 1.5, 2.6 % SVL) wider than basal phalanx; toes fully webbed, webbing formula I 1–1 II 1–1 III 1–1 IV 1–1 V; inner and outer webbing of fourth toe fairly well excised; skin flaps on outer side of fifth and inner side of first toe present, moveable; proximal subarticular tubercle distinct, oval elongated; middle subarticular tubercle distinct, oval elongated, longer than proximal subarticular tubercle; distal subarticular tubercle distinct, slightly rounded; inner metatarsal tubercle elongated (IMTL 4.2, 7.3 % SVL); outer metatarsal tubercle absent.</p>
            <p>Dorsal skin of head and anterior part of the trunk rugose, wrinkled, without warts; posteriorly rugose with small warts; few small warts and wrinkles on upper eyelids; transverse strip between posterior margins of eyes; dorsolateral fold absent; short dorsolateral ridge behind eyes; strong, curved, supratympanic fold from eye to above arm base; ventral skin of abdomen and thigh smooth; cloaca rugose with small warts; ulnar side of hand with scattered translucent spinules; dorsal skin of tibia densely covered with small warts and coarsely scattered with well-elevated large warts, each wart tipped with translucent spinule; tarsus covered with small warts dorsolaterally; skin flap along outer side of fifth toe with translucent spinules; ventral metatarsal skin of fourth toe with irregular translucent spinules.</p>
            <p>Coloration. Dorsal side greenish-brown with irregularly shaped, indistinct confluent dark spots; head with one dark brown interorbital bar; upper and lower lips with three dark brown bands; canthus and supratympanic fold covered by one dark gray band; throat white, with confluent irregular dark gray blotches; abdomen white; limbs marked dorsally with dark brown crossbars; thigh ventral side nearly translucent with thin yellowish blotches. In preservative, whole-body coloration turned olive-gray; dark gray blotches on throat remain unchanged; translucent spinules turned whitish.</p>
            <p>Variation. Individuals of the type series were very similar in proportions. Table 2 shows the individual variations in morphometric values. A single male was greater than females in RHW, RIOD, R1TOEL, ROSL, and RITOL, while females were larger than males in ROW. Male with obtuse tip of odontoid processes. Throat of one female (MZB Amph 33535) almost entirely covered by dark blotches. From base to level of subarticular tubercle of male first finger brownish, thickened, forming a nuptial pad without minute asperities.</p>
            <p> Comparisons.  Limnonectes maanyanorum sp. nov. differs from  L. barioensis and  L. tawauensis by webbing on inner and outer of the fourth toes fairly well excised, Fig. 9E [vs. little excised, Fig. 9G, H], first finger distinctly longer than second finger [vs. slightly longer: Matsui et al. (2024)], skin flap on outer side of third finger freely moveable [vs. weakly moveable, except  L. tawauensis : Matsui et al. (2024)], male having obtuse odontoid processes tip [vs. sharply pointed, except  L. barioensis : Matsui et al. (2024)]; and by female having longer first finger, RFFL 17.2–19.4 % SVL (vs. shorter, ≤ 15.6 % SVL: Matsui et al. 2024), and longer first toe, R1TOEL 19.4–21.9 % SVL (vs. shorter, ≤ 16.0 % SVL: Matsui et al. 2024). </p>
            <p> Limnonectes maanyanorum sp. nov. differs from  L. conspicillatus ,  L. lambirensis ,  L. mocquardi ,  L. penerisensis ,  L. paginatanensis ,  L. separatus ,  L. lanjakensis ,  L. abanghamidi ,  L. batulawensis , and  L. paulyambuni , by dorsal skin of tibia having pattern I of warts (vs. pattern II or III, see Table 3). Additionally,  Limnonectes maanyanorum sp. nov. differs from  L. mocquardi ,  L. paginatanensis ,  L. separatus ,  L. abanghamidi ,  L. batulawensis , and  L. paulyambuni , by having first finger distinctly longer than second finger [vs. first finger as long as or slightly shorter than second: Matsui et al. (2024)].  Limnonectes maanyanorum sp. nov. differs from  L. lambirensis ,  L. mocquardi ,  L. paginatanensis , and  L. abanghamidi , by outer side of third finger having moveable skin flap [vs. weekly moveable or non-moveable: Matsui et al. (2024)]. </p>
            <p> Limnonectes maanyanorum sp. nov. differs from  L. asperatus ,  L. rhacodus ,  L. deinodon Dehling ,  L. hikidai ,  L. cintalubang ,  L. sinuatodorsalis Matsui , and  L. kong , by having fully webbed toes [vs. fourth toe not webbed to base of tip: Inger et al. (1996), Dehling (2014), Matsui &amp; Nishikawa (2014), Matsui et al. (2014b), Matsui (2015), Dehling &amp; Dehling (2017)]; and odontoid processes tip obtuse in male [vs. pointed tusk, except  L. kong : Inger et al. (1996), Dehling (2014), Matsui &amp; Nishikawa (2014), Matsui et al. (2014b), Matsui (2015), Dehling &amp; Dehling (2017)]. Additionally,  Limnonectes maanyanorum sp. nov. differs from  L. kong by absence of minute asperities on male nuptial pads [vs. present: Dehling &amp; Dehling (2017)], and moveable skin flap on the outer side of third finger [vs. non-moveable: Dehling &amp; Dehling (2017)]. </p>
            <p> Limnonectes maanyanorum sp. nov. differs from  L. kuhlii by the absence of thickened skin on the anterior chin in male (vs. present), and nuptial pad present in males (vs. absent).  Limnonectes maanyanorum sp. nov. differs from  L. sisikdagu McLeod, Horner, Husted, Barley &amp; Iskandar , by absence of translucent spinules on ventral skin of anterior chin of males [vs. present: McLeod et al. (2011)], and absence of minute asperities on male nuptial pads [vs. present: McLeod et al. (2011)]. </p>
            <p> Limnonectes maanyanorum sp. nov. differs from  L. utara Matsui, Belabut &amp; Ahmad , and  L. selatan Matsui, Belabut &amp; Ahmad , by smaller of female body size, SVL 40.4–58.5 mm [vs. ≥ 59.2 mm: Matsui et al. (2014a)], few large warts on dorsal skin of tibia [vs. many large wars: Matsui et al. (2014a)], and absence of minute asperities on male nuptial pads [vs. present: Matsui et al. (2014a)]. Additionally,  Limnonectes maanyanorum sp. nov. differs from  L. utara by less confluent dark markings on dorsum [vs. more confluent: Matsui et al. (2014a)]. </p>
            <p> Distribution and natural history.  Limnonectes maanyanorum sp. nov. is distributed within an elevational range of 91–164 m a.s.l. in the northern part of Meratus Mountain Range based on available data. The type series from Mt. Karasik were found in secondary forest, near disturbed areas along small, rocky stream with a width of up to 2 m (Fig. 10A). The holotype was collected from the ground at the edge of the stream at around 20:00 h, while other type specimens have been found on leaves or rocks near the stream. The following anuran species have been found at the Mt. Karasik site with the new species:  Hylarana picturata (Boulenger) ;  Kalophrynus meizon Zug ;  L. leporinus (Andersson) ;  L. paramacrodon (Inger) ;  Leptobrachium abbotti (Cochran) ;  Limnonectes sp. 1 ;  Pelobatrachus kalimantanensis (Munir, Hamidy, Matsui, Iskandar, Sidik, &amp; Shimada) ;  Leptobrachella fritinniens (Dehling &amp; Matsui) ;  Phrynoidis asper (Gravenhorst) ; and  Staurois guttatus (Günther) . </p>
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	https://treatment.plazi.org/id/03D6AC50FF8E8F6A0AF79D64CE53FD7C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Gonggoli, Ade Damara;Shimada, Tomohiko;Matsui, Masafumi;Nishikawa, Kanto;Sidik, Irvan;Kadafi, Ahmad Muammar;Farajallah, Achmad;Hamidy, Amir	Gonggoli, Ade Damara, Shimada, Tomohiko, Matsui, Masafumi, Nishikawa, Kanto, Sidik, Irvan, Kadafi, Ahmad Muammar, Farajallah, Achmad, Hamidy, Amir (2025): Two new species of fanged frog from Southeastern Borneo, Indonesia (Amphibia: Anura: Dicroglossidae). Zootaxa 5575 (3): 387-408, DOI: 10.11646/zootaxa.5575.3.3, URL: https://doi.org/10.11646/zootaxa.5575.3.3
03D6AC50FF828F700AF79C92C8EEFCA4.text	03D6AC50FF828F700AF79C92C8EEFCA4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Limnonectes nusantara Gonggoli & Shimada & Matsui & Nishikawa & Sidik & Kadafi & Farajallah & Hamidy 2025	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Limnonectes nusantara sp. nov.</p>
            <p>(Figs. 7–8)</p>
            <p> Limnonectes kuhlii “Lineage 4”: Matsui et al. (2016) </p>
            <p>
                  Holotype.  
                <a title="Search Plazi for locations around (long 115.42577/lat -2.803344)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=115.42577&amp;materialsCitation.latitude=-2.803344">Adult</a>
                 male (MZB Amph 33516) from  
                <a title="Search Plazi for locations around (long 115.42577/lat -2.803344)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=115.42577&amp;materialsCitation.latitude=-2.803344">Batubalay River</a>
                 ,  
                <a title="Search Plazi for locations around (long 115.42577/lat -2.803344)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=115.42577&amp;materialsCitation.latitude=-2.803344">Lumpangi Village</a>
                 , Loksado District, Hulu Sungai  Selatan Regency , South Kalimantan Province (Borneo), Indonesia at - 2.803344°S, 115.425771°E, elevation 177 m a.s.l., collected by Ade Damara Gonggoli, Adventus Robertino Rangin, and Ayibino on 9 January 2023. 
            </p>
            <p>
                 Paratypes (n = 16). Total of   14 specimens from  Loksado District , Hulu Sungai  Selatan Regency , South Kalimantan Province (Borneo), Indonesia  :   three adult females (MZB  
                <a title="Search Plazi for locations around (long 115.49275/lat -2.794351)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=115.49275&amp;materialsCitation.latitude=-2.794351">Amph</a>
                 33522–24) from Pante River, Loksado Village at - 2.794351°S, 115.492750°E, elevation 270 m a.s.l., collected by Ade Damara Gonggoli, Adventus Robertino Rangin, and Ayibino on 10 January 2023  ;   two males (MZB  Amph 33526, 28) and   two females (MZB  
                <a title="Search Plazi for locations around (long 115.48245/lat -2.794682)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=115.48245&amp;materialsCitation.latitude=-2.794682">Amph</a>
                 33527, 29) from Jaung River, Loksado Village at - 2.794682°S, 115.482456°E, elevation 226 m a.s.l., collected by Ade Damara Gonggoli, Adventus Robertino Rangin, and Ayibino on 11 January 2023  ;   two adult males (MZB  
                <a title="Search Plazi for locations around (long 115.48587/lat -2.799114)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=115.48587&amp;materialsCitation.latitude=-2.799114">Amph</a>
                 25120, 22) from Loksado Village at - 2.799114°S, 115.485869°E, elevation 262 m a.s.l., collected by Irvan Sidik on 18 April 2008  ;   five adult females (MZB  Amph 25110–11, 25291–92, 25405) from Malaris Village collected by Irvan Sidik in April 2008  . 
            </p>
            <p>
                  One adult female (MZB Amph 15256) from  
                <a title="Search Plazi for locations around (long 115.48778/lat -3.028892)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=115.48778&amp;materialsCitation.latitude=-3.028892">Dusun Bancing Village</a>
                 , Paramasan District, Banjar Regency, South Kalimantan Province (Borneo), Indonesia at - 3.028892°S, 115.487781°E, elevation 527 m a.s.l., collected by Amir Hamidy and Abran Slamet on 17 February 2009  .  One adult male (MZB Amph 33520) collected with holotype . 
            </p>
            <p>
                 Referred specimens (n=12). Total of   five specimens from  Loksado District , Hulu Sungai  Selatan Regency , South Kalimantan Province (Borneo), Indonesia  :  one adult male (MZB Amph 33536) and   one adult female (KUHE 60952) from  
                <a title="Search Plazi for locations around (long 115.506195/lat -2.804056)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=115.506195&amp;materialsCitation.latitude=-2.804056">Malaris Village</a>
                 at - 2.804056°S, 115.506195°E, elevation 264 m a.s.l., collected by Tomohiko Shimada and Misbahul Munir on 19 September 2017  ;   one adult female (KUHE 60971)  
                <a title="Search Plazi for locations around (long 115.511536/lat -2.782646)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=115.511536&amp;materialsCitation.latitude=-2.782646">Loksado Village</a>
                 at one the way to Haratai Village, at - 2.782646°S, 115.511533°E, elevation 276 m a.s.l., collected by Tomohiko Shimada and Misbahul Munir on 20 September 2017  ;   two juveniles (MZB  
                <a title="Search Plazi for locations around (long 115.49275/lat -2.794351)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=115.49275&amp;materialsCitation.latitude=-2.794351">Amph</a>
                 33521, 25) from Pante River, Loksado Village at - 2.794351°S, 115.492750°E, elevation 270 m a.s.l., collected by Ade Damara Gonggoli, Adventus Robertino Rangin, and Ayibino on 10 January 2023  . 
            </p>
            <p>
                  One juvenile (MZB Amph 15266) from  
                <a title="Search Plazi for locations around (long 115.48778/lat -3.028892)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=115.48778&amp;materialsCitation.latitude=-3.028892">Dusun Bancing Village</a>
                 , Paramasan District, Banjar Regency, South Kalimantan Province (Borneo), Indonesia at - 3.028892°S, 115.487781°E, elevation 527 m a.s.l., collected by Amir Hamidy and Abran Slamet on 17 February 2009  . 
            </p>
            <p> Total of   three specimens from  Mt. Halau-halau ,  Batang Alai Timur District , Hulu Sungai Tengah Regency (Borneo)  :   one juvenile (KUHE 61013) from  Karuh River ;   one juvenile (KUHE 61015) from  Panitiranggang , and   one juvenile (MZB Amph 33630) from  Jumantir .  Three juveniles (MZB Amph 33517–19) were collected with holotype . </p>
            <p>Etymology. The species epithet refers to the name of the proposed new capital of Indonesia, located in Kalimantan (Borneo Part Indonesia), from where the new species is described. This name is an invariable noun in apposition.</p>
            <p>Local common name. In Loksado village, the local people of Dayak Meratus ethnic call this frog “Lampinik”.</p>
            <p>Suggested English common name. Nusantara Creek Frog.</p>
            <p>Suggested Indonesian common name. Bangkong-tuli Nusantara.</p>
            <p> Diagnosis. The new species clustered within the  L. kuhlii complex in molecular analyses and can be distinguished from its congeners by the following combination of morphological characters: medium size (SVL 52.3–70.6 mm in males, 51.6–61.3 mm in females); toes fully webbed; tympanum hidden; dorsal skin of body weakly rugose; dorsal skin of tibia densely covered by small warts and scattered with low large warts; flap of skin on outer side of third finger well-developed, moveable; inner and outer webbing of fourth toe fairly well excised; disk of third finger not much wider than basal phalanx; disk of fourth toe wider than basal phalanx; nuptial pad on first finger without minute asperities in males; head longer than wide; and throat covered with small irregular dark gray blotches. </p>
            <p>Description of the holotype (measurements in mm). Adult male (SVL 64.7), habitus stocky; head longer (HL 28.9, 44.6 % SVL) than wide (HW 27.3, 42.2 % SVL); rostrum longer (RL 9.9, 15.3 % SVL) than eye (ED 5.7, 8.7 % SVL); nostril dorsolateral on canthus, closer to snout tip (SL 4.4, 6.8 % SVL) than to eye (N-EL 5.5, 8.5 % SVL); internarial distance (IND 5.3, 8.1 % SVL) slightly shorter than interorbital distance (IOD 5.4, 8.3 % SVL); snout obtusely pointed in lateral view; canthus rounded; lore concave; pineal spot distinct; tympanum indistinct; pair of odontoid processes on lower jaw, obtuse in profile; vomerine teeth in oblique groups, groups separated from each another and from choana by half length of one group; vocal sac and vocal slits absent.</p>
            <p>Forelimb stout, relatively short (FLL 31.6, 48.8 % SVL); fingers moderately slender; finger length formula II &lt;I &lt;IV &lt;III; finger webbing absent; finger tips blunt, rounded, without circummarginal grooves; disk of third finger (3TDW 1.1, 1.7 % SVL) not wider than basal phalanx; inner palmar tubercle oval and low (IPTL 4.6, 7.1 % SVL); middle palmar tubercle oval and low, contacting outer palmar tubercle; outer palmar tubercle slightly oval and elongated; proximal subarticular tubercle slightly rounded, elevated; distal subarticular tubercle indistinct; skin flaps present on inner and outer side of second and third fingers, more developed on outer side of third finger than inner, freely moveable; supernumerary metacarpal tubercles absent; medial skin of first finger from base to level of subarticular tubercle yellowish, thickened, forming nuptial pad without minute asperities.</p>
            <p>Hindlimb stout, relatively short (HLL 93.1, 143.9 % SVL), about three times length of forelimb, toe length formula I &lt;II &lt;V &lt;III &lt;IV; foot (FL 29.5, 45.6 % SVL) slightly longer than tibia (TL 29.3, 45.3 % SVL); tibiotarsal articulation reaching between eye and nostril in preserved condition; toes tips blunt, rounded; disk of fourth toe (4TDW 1.6, 2.4 % SVL) wider than basal phalanx; toes fully webbed, webbing formula I 1–1 II 1–1 III 1–1 IV 1–1 V; inner and outer webbing of fourth toe fairly well excised; skin flap on outer side of fifth and inner side of first toe present, moveable; proximal subarticular tubercle distinct, oval; middle subarticular tubercle distinct, oval slightly elongated, longer than proximal subarticular tubercle; distal subarticular tubercle distinct, oval; inner metatarsal tubercle elongated (IMTL 4.7, 7.2 % SVL); outer metatarsal tubercle absent.</p>
            <p>Dorsal skin of head and anterior part of the trunk weakly rugose, wrinkled, without warts; posteriorly rugose with small warts; few small warts and wrinkles on upper eyelids; transverse strip between posterior margins of eyes; dorsolateral fold absent; short dorsolateral ridge behind eyes; strong, curved, supratympanic fold from eye to above arm base; ventral surface of abdomen and thigh smooth; cloaca rugose with small warts; ulnar side of hand with scattered translucent spinules; dorsal skin of tibia covered with numerous small warts and scattered with low large warts, each wart tipped with translucent spinule; tarsus covered with small warts dorsolaterally; skin flap along outer side of fifth toe with translucent spinules; ventral metatarsal skin of fourth toe with irregular translucent spinules.</p>
            <p>Coloration. Dorsal side yellowish-brown with irregularly shaped, confluent dark spots; head with one dark brown interorbital bar; upper and lower lips with three dark brown bands; canthus and supratympanic fold covered by dark gray band; throat white, with small irregular dark gray blotches; abdomen yellowish-white; limbs marked dorsally with dark brown crossbars; thigh ventral side nearly translucent with thin yellowish blotches. In preservative, whole-body coloration turned blackish-red, dark gray spots on throat faded and become cream, and translucent spinules turned whitish.</p>
            <p>Variation. Individuals of the type series were generally similar in proportions. Table 2 shows the individual variations in morphometric values. Males were greater than females in ROH and ROSL; while females were larger than males in ROW (Table 4). One male (MZB Amph 33527) had a larger body (SVL 70.5 mm vs. 64.7 mm) and a thicker nuptial pad than the holotype. One male (MZB Amph 33536) and one female (MZB Amph 33526) had a wide longitudinal white line from the snout to cloaca. One female (MZB Amph 15256) had irregular dark blotches covering the abdomen.</p>
            <p> Comparisons.  Limnonectes nusantara sp. nov. differs from  Limnonectes maanyanorum sp. nov. by larger mean of female body size, SVL 51.6–61.3 mm, mean 49.2 ± SD 7 (vs. 40.4–58.5 mm, mean 49.2 ± SD 7.1); throat covered with small dark blotches, Fig. 7C, F (vs. covered with confluent dark blotches, Fig. 5C, F); male dorsal surface of tibia covered with low warts and relatively larger, Figs. 7A, D, 9C, D (vs. well-elevated warts and relatively smaller, Figs. 5A, D, 9A, B); and females having shorter head, RHL 40.0–43.1 % SVL, 41.8 ± SD 1.0 (vs. longer, 41.3–46.3 % SVL, mean 43.5 ± SD 1.7), shorter nostril-eye length, RN-EL 8.0–9.5 % SVL, mean 8.7 ± SD 0.5 (vs. longer, 9.1–10.6 % SVL, mean 9.8 ± SD 0.6), shorter snout length, RSL 6.0–7.2 % SVL, mean 6.6 ± SD 0.4 (vs. longer, 6.4–8.2 % SVL, mean 7.2 ± SD 0.6), shorter rostrum, RRL 14.1–16.8 % SVL, mean 15.5 ± SD 1.0 (vs. longer, 15.7–18.6 % SVL, mean 17.0 ± SD 0.9), and shorter forelimb RFLL 45.9–55.2 % SVL, mean 50.4 ± SD 2.8 (vs. longer, vs. 49.2–57.8 % SVL, mean 54.0 ± SD 2.8). </p>
            <p> Limnonectes nusantara sp. nov. differs from  L. barioensis and  L. tawauensis by outer and inner webbing of fourth toes fairly well excised, Fig. 9F [vs. little excised, Fig. 9G, H], first finger distinctly longer than second finger [vs. slightly longer: Matsui et al. (2024)], skin flap on outer side of third finger freely moveable [vs. weakly moveable, except  L. tawauensis : Matsui et al. (2024)], male having obtuse odontoid processes tip (vs. sharply pointed, except  L. barioensis : Matsui et al. 2024); and by female having longer first finger, RFFL 16.6–20.0 % SVL (vs. shorter, ≤ 15.6 % SVL: Matsui et al. 2024), and longer first toe, R1TOEL 20.2–21.7 % SVL (vs. shorter, ≤ 16.0 % SVL: Matsui et al. 2024). </p>
            <p> Limnonectes nusantara sp. nov. differs from  L. conspicillatus ,  L. lambirensis ,  L. mocquardi ,  L. penerisensis ,  L. paginatanensis ,  L. separatus ,  L. lanjakensis ,  L. abanghamidi ,  L. batulawensis , and  L. paulyambuni by dorsal tibia having pattern I of warts (vs. pattern II or III, see Table 3). Additionally,  Limnonectes nusantara sp. nov. differs from  L. mocquardi ,  L. paginatanensis ,  L. separatus ,  L. separatus ,  L. abanghamidi ,  L. batulawensis , and  L. paulyambuni , by having first finger distinctly longer than second finger [vs. first finger as long as or slightly shorter than second: Matsui et al. (2024)].  Limnonectes nusantara sp. nov. differs from  L. lambirensis ,  L. mocquardi ,  L. paginatanensis , and  L. abanghamidi by outer side of third finger having moveable skin flap [vs. weekly moveable or non-moveable: Matsui et al. (2024)]. </p>
            <p> Limnonectes nusantara sp. nov. differs from  L. asperatus ,  L. rhacodus ,  L. deinodon ,  L. hikidai ,  L. cintalubang ,  L. sinuatodorsalis , and  L. kong , by having fully webbed toes [vs. fourth toe not webbed to base of tip: Inger et al. (1996), Dehling (2014), Matsui &amp; Nishikawa (2014), Matsui et al. (2014b), Matsui (2015), Dehling &amp; Dehling (2017)]; and odontoid processes tip obtuse in males [vs. pointed, except  L. kong : Inger et al. (1996), Dehling (2014), Matsui &amp; Nishikawa (2014), Matsui et al. (2014b), Matsui (2015), Dehling &amp; Dehling (2017)]. Additionally,  Limnonectes nusantara sp. nov. differs from  L. kong by absence of minute asperities on male nuptial pads [vs. present: Dehling &amp; Dehling (2017)], and moveable skin flap on the outer side of third finger [vs. non-moveable: Dehling &amp; Dehling (2017)]. </p>
            <p> Limnonectes nusantara sp. nov. differs from  L. kuhlii by the absence of thickened skin on the ventral surface of the anterior chin in male (vs. present), and nuptial pad present in males (vs. absent).  Limnonectes nusantara sp. nov. differs from  L. sisikdagu by absence of translucent spinules on ventral surface of anterior chin of males [vs. present: McLeod et al. (2011)], and absence of minute asperities on male nuptial pads [vs. present: McLeod et al. (2011)]. </p>
            <p> Limnonectes nusantara sp. nov. differs from  L. utara and  L. selatan by smaller mean of female body size, 51.6–61.3 mm, mean 55.9 SD ± 3.0 [vs. 59.2–78.7 mm, mean 69.5 ± SD 6.8 in  L. utara ; 61.6–73.0 mm, mean 70.0 SD ± 3.8 in  L. selatan : Matsui et al. (2014a)]; having few large warts on tibia [vs. many large warts: Matsui et al. (2014a)], and absence of minute asperities on males nuptial pads [vs. present: Matsui et al. (2014a)]. Additionally,  Limnonectes nusantara sp. nov. differs from  L. utara by smaller of male body size, 52.3–70.6 mm [vs. 70.0– 79.1 mm: Matsui et al. 2014a], and less confluent dark markings on dorsum [vs. more confluent: Matsui et al. (2014a)]. </p>
            <p> Distribution and natural history.  Limnonectes nusantara sp. nov. is distributed at an altitude of 177–1000 m a.s.l. in the central part of the Meratus Mountain Range. The type specimens were found along a rocky tributary up to 2 m from the water (Fig. 10B). The holotype was collected on the ground at the edge of the river around 19:45 h. Other specimens have been found on leaves and rocks, and the species is rarely seen in water. The species inhabits primary and secondary forests as well as nearby disturbed areas. The following anuran species have been found at the type locality were  Ansonia spinulifer (Mocquard) ;  Chaperina fusca Mocquard ;  Hylarana picturata ;  Ingerophrynus divergens (Peters) ;  Kalophrynus meizon ;  L. leporinus ;  L. paramacrodon ;  Leptobrachium abbotti ;  Limnonectes sp. 2 ;  Microhyla sundaica Trofimets, Dufresnes, Pawangkhanant, Bragin, Gorin, Hasan, Lalremsanga, Muin, Le, Nguyen, Suwannapoom &amp; Poyarkov ;  Microhyla malang Matsui ;  Pelobatrachus kalimantanensis ;  Leptobrachella fritinniens ;  Phrynoidis asper ; and  Staurois guttatus . </p>
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	https://treatment.plazi.org/id/03D6AC50FF828F700AF79C92C8EEFCA4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Gonggoli, Ade Damara;Shimada, Tomohiko;Matsui, Masafumi;Nishikawa, Kanto;Sidik, Irvan;Kadafi, Ahmad Muammar;Farajallah, Achmad;Hamidy, Amir	Gonggoli, Ade Damara, Shimada, Tomohiko, Matsui, Masafumi, Nishikawa, Kanto, Sidik, Irvan, Kadafi, Ahmad Muammar, Farajallah, Achmad, Hamidy, Amir (2025): Two new species of fanged frog from Southeastern Borneo, Indonesia (Amphibia: Anura: Dicroglossidae). Zootaxa 5575 (3): 387-408, DOI: 10.11646/zootaxa.5575.3.3, URL: https://doi.org/10.11646/zootaxa.5575.3.3
