taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
03CB2A29C513FFDD5D9EF099B8F1FC6B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15214810/files/figure.png	https://doi.org/10.5281/zenodo.15214810	FIGURES 3–4. Inotrechus usholtaensis sp. nov. (3) Habitus of the holotype, dorsal view. (4) Habitus of a paratype, lateral view. Note: setae are marked with white dots.	FIGURES 3–4. Inotrechus usholtaensis sp. nov. (3) Habitus of the holotype, dorsal view. (4) Habitus of a paratype, lateral view. Note: setae are marked with white dots.	2025-03-19	Maghradze, Eter;Barjadze, Shalva;Lohaj, Roman;Faille, Arnaud		Zenodo	biologists	Maghradze, Eter;Barjadze, Shalva;Lohaj, Roman;Faille, Arnaud			
03CB2A29C513FFDD5D9EF099B8F1FC6B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15214812/files/figure.png	https://doi.org/10.5281/zenodo.15214812	FIGURES 5–6. Inotrechus usholtaensis sp. nov. (5) Head of a paratype, lateral view. Note: setae are marked with white dots. (6) Head and ocular area of the holotype, dorsal view.	FIGURES 5–6. Inotrechus usholtaensis sp. nov. (5) Head of a paratype, lateral view. Note: setae are marked with white dots. (6) Head and ocular area of the holotype, dorsal view.	2025-03-19	Maghradze, Eter;Barjadze, Shalva;Lohaj, Roman;Faille, Arnaud		Zenodo	biologists	Maghradze, Eter;Barjadze, Shalva;Lohaj, Roman;Faille, Arnaud			
03CB2A29C513FFDD5D9EF099B8F1FC6B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15214814/files/figure.png	https://doi.org/10.5281/zenodo.15214814	FIGURES 7–12. Aedeagi of Inotrechus species. (7–8) I. kurnakovi: 7. Lateral view, 8. Dorsal view; (9–10) I. injaevae: 9. Lateral view, 10. Dorsal view; (11–12) I. usholtaensis sp. nov.: 11. Holotype, lateral view, 12. Paratype, dorsal view.	FIGURES 7–12. Aedeagi of Inotrechus species. (7–8) I. kurnakovi: 7. Lateral view, 8. Dorsal view; (9–10) I. injaevae: 9. Lateral view, 10. Dorsal view; (11–12) I. usholtaensis sp. nov.: 11. Holotype, lateral view, 12. Paratype, dorsal view.	2025-03-19	Maghradze, Eter;Barjadze, Shalva;Lohaj, Roman;Faille, Arnaud		Zenodo	biologists	Maghradze, Eter;Barjadze, Shalva;Lohaj, Roman;Faille, Arnaud			
03CB2A29C513FFDD5D9EF099B8F1FC6B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15214816/files/figure.png	https://doi.org/10.5281/zenodo.15214816	FIGURES 13–14. Female genitalia (13) Duvalius gregoi sp. nov.; (14) Inotrechus usholtaensis sp. nov. Gonocoxite II: 1— apical nematiform setae, 2—ensiform spines; gonocoxite I: 3—long setae on the inner margin, 4—short randomly distributed setae on the ventral projection; 5—laterotergite setae.	FIGURES 13–14. Female genitalia (13) Duvalius gregoi sp. nov.; (14) Inotrechus usholtaensis sp. nov. Gonocoxite II: 1— apical nematiform setae, 2—ensiform spines; gonocoxite I: 3—long setae on the inner margin, 4—short randomly distributed setae on the ventral projection; 5—laterotergite setae.	2025-03-19	Maghradze, Eter;Barjadze, Shalva;Lohaj, Roman;Faille, Arnaud		Zenodo	biologists	Maghradze, Eter;Barjadze, Shalva;Lohaj, Roman;Faille, Arnaud			
03CB2A29C513FFDD5D9EF099B8F1FC6B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15262730/files/figure.png	https://doi.org/10.5281/zenodo.15262730	MAP 1. Distribution map of the Inotrechus and Duvalius species in Georgias. (1) D. abyssimus (Krubera-Voronya Cave); (2) D. gregoi sp. nov. (Kvedi II cave); (3) D. iljukhini (Souvenir Cave); (4) D. kurnakovi (Baniskhevi valley, Borjomi-Kharagauli National Park); (5) D. megrel (Samegrelo-Zemo Svaneti region, Kvira Mountain); (6) D. sokolovi (Trechus’ Cave); (7) I.injaevae (Sakishore Cave); (8) I. kurnakovi (Prometheus and Melouri caves); (9) I. usholtaensis sp. nov. (Usholta Cave).	MAP 1. Distribution map of the Inotrechus and Duvalius species in Georgias. (1) D. abyssimus (Krubera-Voronya Cave); (2) D. gregoi sp. nov. (Kvedi II cave); (3) D. iljukhini (Souvenir Cave); (4) D. kurnakovi (Baniskhevi valley, Borjomi-Kharagauli National Park); (5) D. megrel (Samegrelo-Zemo Svaneti region, Kvira Mountain); (6) D. sokolovi (Trechus’ Cave); (7) I.injaevae (Sakishore Cave); (8) I. kurnakovi (Prometheus and Melouri caves); (9) I. usholtaensis sp. nov. (Usholta Cave).	2025-03-19	Maghradze, Eter;Barjadze, Shalva;Lohaj, Roman;Faille, Arnaud		Zenodo	biologists	Maghradze, Eter;Barjadze, Shalva;Lohaj, Roman;Faille, Arnaud			
03CB2A29C517FFDD5D9EF449BCC4FACE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15214810/files/figure.png	https://doi.org/10.5281/zenodo.15214810	FIGURES 3–4. Inotrechus usholtaensis sp. nov. (3) Habitus of the holotype, dorsal view. (4) Habitus of a paratype, lateral view. Note: setae are marked with white dots.	FIGURES 3–4. Inotrechus usholtaensis sp. nov. (3) Habitus of the holotype, dorsal view. (4) Habitus of a paratype, lateral view. Note: setae are marked with white dots.	2025-03-19	Maghradze, Eter;Barjadze, Shalva;Lohaj, Roman;Faille, Arnaud		Zenodo	biologists	Maghradze, Eter;Barjadze, Shalva;Lohaj, Roman;Faille, Arnaud			
03CB2A29C517FFDD5D9EF449BCC4FACE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15214812/files/figure.png	https://doi.org/10.5281/zenodo.15214812	FIGURES 5–6. Inotrechus usholtaensis sp. nov. (5) Head of a paratype, lateral view. Note: setae are marked with white dots. (6) Head and ocular area of the holotype, dorsal view.	FIGURES 5–6. Inotrechus usholtaensis sp. nov. (5) Head of a paratype, lateral view. Note: setae are marked with white dots. (6) Head and ocular area of the holotype, dorsal view.	2025-03-19	Maghradze, Eter;Barjadze, Shalva;Lohaj, Roman;Faille, Arnaud		Zenodo	biologists	Maghradze, Eter;Barjadze, Shalva;Lohaj, Roman;Faille, Arnaud			
03CB2A29C517FFDD5D9EF449BCC4FACE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15214814/files/figure.png	https://doi.org/10.5281/zenodo.15214814	FIGURES 7–12. Aedeagi of Inotrechus species. (7–8) I. kurnakovi: 7. Lateral view, 8. Dorsal view; (9–10) I. injaevae: 9. Lateral view, 10. Dorsal view; (11–12) I. usholtaensis sp. nov.: 11. Holotype, lateral view, 12. Paratype, dorsal view.	FIGURES 7–12. Aedeagi of Inotrechus species. (7–8) I. kurnakovi: 7. Lateral view, 8. Dorsal view; (9–10) I. injaevae: 9. Lateral view, 10. Dorsal view; (11–12) I. usholtaensis sp. nov.: 11. Holotype, lateral view, 12. Paratype, dorsal view.	2025-03-19	Maghradze, Eter;Barjadze, Shalva;Lohaj, Roman;Faille, Arnaud		Zenodo	biologists	Maghradze, Eter;Barjadze, Shalva;Lohaj, Roman;Faille, Arnaud			
03CB2A29C517FFDD5D9EF449BCC4FACE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15214818/files/figure.png	https://doi.org/10.5281/zenodo.15214818	FIGURES 15–16. Habitus of Inotrechus species. (15) I. injaevae; (16) I. kurnakovi (dorsal view). Note: setae are marked with white dots.	FIGURES 15–16. Habitus of Inotrechus species. (15) I. injaevae; (16) I. kurnakovi (dorsal view). Note: setae are marked with white dots.	2025-03-19	Maghradze, Eter;Barjadze, Shalva;Lohaj, Roman;Faille, Arnaud		Zenodo	biologists	Maghradze, Eter;Barjadze, Shalva;Lohaj, Roman;Faille, Arnaud			
03CB2A29C517FFD05D9EF52DBDA2FC8B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15214820/files/figure.png	https://doi.org/10.5281/zenodo.15214820	FIGURES 17–18. Duvalius gregoi sp. nov. (17) Habitus of the holotype, dorsal view. Note: setae are marked with white dots. (18) Living specimen in Kvedi II Cave. Photo by Lado Shavadze.	FIGURES 17–18. Duvalius gregoi sp. nov. (17) Habitus of the holotype, dorsal view. Note: setae are marked with white dots. (18) Living specimen in Kvedi II Cave. Photo by Lado Shavadze.	2025-03-19	Maghradze, Eter;Barjadze, Shalva;Lohaj, Roman;Faille, Arnaud		Zenodo	biologists	Maghradze, Eter;Barjadze, Shalva;Lohaj, Roman;Faille, Arnaud			
03CB2A29C517FFD05D9EF52DBDA2FC8B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15214822/files/figure.png	https://doi.org/10.5281/zenodo.15214822	FIGURE 19. Duvalius gregoi sp. nov., head and ocular area of the holotype, dorsal view.	FIGURE 19. Duvalius gregoi sp. nov., head and ocular area of the holotype, dorsal view.	2025-03-19	Maghradze, Eter;Barjadze, Shalva;Lohaj, Roman;Faille, Arnaud		Zenodo	biologists	Maghradze, Eter;Barjadze, Shalva;Lohaj, Roman;Faille, Arnaud			
03CB2A29C517FFD05D9EF52DBDA2FC8B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15214824/files/figure.png	https://doi.org/10.5281/zenodo.15214824	FIGURES 20–21. Aedeagus of Duvalius gregoi sp. nov. (20) Lateral view, the holotype, (21) Dorsal view, a paratype.	FIGURES 20–21. Aedeagus of Duvalius gregoi sp. nov. (20) Lateral view, the holotype, (21) Dorsal view, a paratype.	2025-03-19	Maghradze, Eter;Barjadze, Shalva;Lohaj, Roman;Faille, Arnaud		Zenodo	biologists	Maghradze, Eter;Barjadze, Shalva;Lohaj, Roman;Faille, Arnaud			
03CB2A29C517FFD05D9EF52DBDA2FC8B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15214816/files/figure.png	https://doi.org/10.5281/zenodo.15214816	FIGURES 13–14. Female genitalia (13) Duvalius gregoi sp. nov.; (14) Inotrechus usholtaensis sp. nov. Gonocoxite II: 1— apical nematiform setae, 2—ensiform spines; gonocoxite I: 3—long setae on the inner margin, 4—short randomly distributed setae on the ventral projection; 5—laterotergite setae.	FIGURES 13–14. Female genitalia (13) Duvalius gregoi sp. nov.; (14) Inotrechus usholtaensis sp. nov. Gonocoxite II: 1— apical nematiform setae, 2—ensiform spines; gonocoxite I: 3—long setae on the inner margin, 4—short randomly distributed setae on the ventral projection; 5—laterotergite setae.	2025-03-19	Maghradze, Eter;Barjadze, Shalva;Lohaj, Roman;Faille, Arnaud		Zenodo	biologists	Maghradze, Eter;Barjadze, Shalva;Lohaj, Roman;Faille, Arnaud			
03CB2A29C517FFD05D9EF52DBDA2FC8B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15262730/files/figure.png	https://doi.org/10.5281/zenodo.15262730	MAP 1. Distribution map of the Inotrechus and Duvalius species in Georgias. (1) D. abyssimus (Krubera-Voronya Cave); (2) D. gregoi sp. nov. (Kvedi II cave); (3) D. iljukhini (Souvenir Cave); (4) D. kurnakovi (Baniskhevi valley, Borjomi-Kharagauli National Park); (5) D. megrel (Samegrelo-Zemo Svaneti region, Kvira Mountain); (6) D. sokolovi (Trechus’ Cave); (7) I.injaevae (Sakishore Cave); (8) I. kurnakovi (Prometheus and Melouri caves); (9) I. usholtaensis sp. nov. (Usholta Cave).	MAP 1. Distribution map of the Inotrechus and Duvalius species in Georgias. (1) D. abyssimus (Krubera-Voronya Cave); (2) D. gregoi sp. nov. (Kvedi II cave); (3) D. iljukhini (Souvenir Cave); (4) D. kurnakovi (Baniskhevi valley, Borjomi-Kharagauli National Park); (5) D. megrel (Samegrelo-Zemo Svaneti region, Kvira Mountain); (6) D. sokolovi (Trechus’ Cave); (7) I.injaevae (Sakishore Cave); (8) I. kurnakovi (Prometheus and Melouri caves); (9) I. usholtaensis sp. nov. (Usholta Cave).	2025-03-19	Maghradze, Eter;Barjadze, Shalva;Lohaj, Roman;Faille, Arnaud		Zenodo	biologists	Maghradze, Eter;Barjadze, Shalva;Lohaj, Roman;Faille, Arnaud			
