taxonID	type	description	language	source
03C38795FFF2FFFAFDB479C0FDB6FBE0.taxon	type_taxon	Type species Ligidium hypnorum (Cuvier, 1792).	en	Recuero, Ernesto, Caterino, Michael S. (2025): Is there anybody (new) out there? Seven new species of Ligidium (Isopoda, Oniscidea, Ligiidae) from the Southern Appalachians, eastern North America. European Journal of Taxonomy 976: 133-170, DOI: 10.5852/ejt.2025.976.2783, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2783/12721
03C38795FFF2FFF9FDBB7EB0FDD1FD43.taxon	description	urn: lsid: zoobank. org: act: CF 5 E 1 EE 6 - 9001 - 45 DA- 9 DB 2 - 5 DE 45 CA 56 D 20 Figs 2 – 3	en	Recuero, Ernesto, Caterino, Michael S. (2025): Is there anybody (new) out there? Seven new species of Ligidium (Isopoda, Oniscidea, Ligiidae) from the Southern Appalachians, eastern North America. European Journal of Taxonomy 976: 133-170, DOI: 10.5852/ejt.2025.976.2783, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2783/12721
03C38795FFF2FFF9FDBB7EB0FDD1FD43.taxon	diagnosis	Diagnosis Ligidium enotahensis sp. nov. can be diagnosed morphologically based on the shape on the male pleopod 2 endopodite, showing a squarish tip with no excrescence, projection, or notch, and on the very broad male pleopod 1 endopodite projection. It also can be identified from all other Appalachian species based on molecular data, representing a distinct evolutionary lineage.	en	Recuero, Ernesto, Caterino, Michael S. (2025): Is there anybody (new) out there? Seven new species of Ligidium (Isopoda, Oniscidea, Ligiidae) from the Southern Appalachians, eastern North America. European Journal of Taxonomy 976: 133-170, DOI: 10.5852/ejt.2025.976.2783, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2783/12721
03C38795FFF2FFF9FDBB7EB0FDD1FD43.taxon	etymology	Etymology From the Cherokee Enotah, a name of the mountain where the species is described, and the Latin suffix ‘ - ēnsis ’, forming an adjective meaning ‘ from Enotah’.	en	Recuero, Ernesto, Caterino, Michael S. (2025): Is there anybody (new) out there? Seven new species of Ligidium (Isopoda, Oniscidea, Ligiidae) from the Southern Appalachians, eastern North America. European Journal of Taxonomy 976: 133-170, DOI: 10.5852/ejt.2025.976.2783, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2783/12721
03C38795FFF2FFF9FDBB7EB0FDD1FD43.taxon	materials_examined	Material examined Holotype USA – Georgia • ♂; Towns Co., Brasstown Bald; 34.8763 ° N, 83.8107 ° W; 1377 m a. s. l.; 17 Nov. 2020; M. Caterino and A. Haberski leg.; GenBank no: OR 172584 (Cox 1); USNM, CUAC 000138051. Other material examined USA – Georgia • 3 imm.; same data as for holotype; CUAC 000171312 to CUAC 000171314.	en	Recuero, Ernesto, Caterino, Michael S. (2025): Is there anybody (new) out there? Seven new species of Ligidium (Isopoda, Oniscidea, Ligiidae) from the Southern Appalachians, eastern North America. European Journal of Taxonomy 976: 133-170, DOI: 10.5852/ejt.2025.976.2783, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2783/12721
03C38795FFF2FFF9FDBB7EB0FDD1FD43.taxon	description	Description Body length of holotype 5.3 mm, body width (at pereonite 4) 2.6 mm. Color of holotype in ethanol (Fig. 2) dorsally brown with large whitish patches on dorsal surface, a dark brown longitudinal medial stripe running from pereonite 1 to telson, epimera dark brown with a large white patch; head, antennae and uropods dark brown, pereopods and pleopods lighter brown. Immatures much lighter and without longitudinal stripe. Antennule (Fig. 3 G) three-segmented; first segment about 2 times as long as wide, with 3 strong spiniform setae on its distal border; second segment slender, about 4 times as long as wide, also bearing 3 strong spiniform setae on the distal border; third segment small, with round apex and a single seta. Antennae (Fig. 3 F) with 5 segmented peduncle, relative size of antennomeres 5> 4> 3> 2> 1; antennal flagellum with 11 articles. Dactylus of pereopods with outer claw longer than inner one; no sexual dimorphism observed in pereopods 1 (Fig. 3 H) and 7 (Fig. 3 I). Male pleopod 1 endopodite (Fig. 3 A) with broad, squarish projection bearing 3 strong, blunt setae, 1.4 times as long as endopodite. Pleopod 1 exopodite (Fig. 3 B) with a flat caudal margin bearing 3 strong, blunt setae, 0.7 times as long as exopodite. Male pleopod 2 exopodite (Fig. 3 C) 2 times as wide as long; inner and most part of frontal margin with dense, hairy setation, as in outer caudal corner. Male pleopod 2 endopodite (Fig. 3 D – E) with squared tip showing a ventral oblique line of small, poorly defined denticles along the interior corner; inner margin with minute spiniform setae, no spines at the base. Telson (Fig. 2) with caudal margin obtusely produced. Uropod (Fig. 3 J) with exopodite broken, endopodite about 1.5 times as long as basipodite; in the examined immatures endopodite 1.5 – 1.6 times as long as exopodite.	en	Recuero, Ernesto, Caterino, Michael S. (2025): Is there anybody (new) out there? Seven new species of Ligidium (Isopoda, Oniscidea, Ligiidae) from the Southern Appalachians, eastern North America. European Journal of Taxonomy 976: 133-170, DOI: 10.5852/ejt.2025.976.2783, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2783/12721
03C38795FFF2FFF9FDBB7EB0FDD1FD43.taxon	distribution	Distribution The species is so far only known from its type locality, near the southern extreme of the southern Appalachian Mountains (Fig. 2). All specimens were collected in mixed forest leaf litter samples.	en	Recuero, Ernesto, Caterino, Michael S. (2025): Is there anybody (new) out there? Seven new species of Ligidium (Isopoda, Oniscidea, Ligiidae) from the Southern Appalachians, eastern North America. European Journal of Taxonomy 976: 133-170, DOI: 10.5852/ejt.2025.976.2783, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2783/12721
03C38795FFF2FFF9FDBB7EB0FDD1FD43.taxon	discussion	Remarks Ligidium enotahensis sp. nov. corresponds with Ligidium sp. 1 in Recuero & Caterino (2024 a). This new species is closely related to L. protuberans sp. nov. and L. schultzi sp. nov., but they can easily be identified based on the shape of male pleopod 2 endopodite, by lacking the conspicuous projection present in L. protuberans, or the marked notch typical of L. schultzi. Ligidium enotahensis also presents a much broader projection of male pleopod 1 endopodite. All three gene fragments analyzed are informative to identify this species, with high genetic p-distances for Cox 1, and low in the conserved NaK and 28 S fragments (Supp. file 1), forming a reciprocally monophyletic clade with respect to either of the other two in all cases.	en	Recuero, Ernesto, Caterino, Michael S. (2025): Is there anybody (new) out there? Seven new species of Ligidium (Isopoda, Oniscidea, Ligiidae) from the Southern Appalachians, eastern North America. European Journal of Taxonomy 976: 133-170, DOI: 10.5852/ejt.2025.976.2783, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2783/12721
03C38795FFF1FFF4FDAD792FFD7BFC83.taxon	description	urn: lsid: zoobank. org: act: A 2 CAA 049 - 6 BCB- 4229 - A 06 D- 49 F 61 B 36 B 030 Figs 4 – 5	en	Recuero, Ernesto, Caterino, Michael S. (2025): Is there anybody (new) out there? Seven new species of Ligidium (Isopoda, Oniscidea, Ligiidae) from the Southern Appalachians, eastern North America. European Journal of Taxonomy 976: 133-170, DOI: 10.5852/ejt.2025.976.2783, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2783/12721
03C38795FFF1FFF4FDAD792FFD7BFC83.taxon	diagnosis	Diagnosis Ligidium nantahala sp. nov. presents a male pleopod 2 endopodite with a robust excrescence occupying the inner part of the tip and projected outwards. The projection of male pleopod 1 endopodite presents a triangular profile. It also can be distinguished from all other Appalachian species based on molecular data, representing a distinct evolutionary lineage with no close relationships among the studied taxa.	en	Recuero, Ernesto, Caterino, Michael S. (2025): Is there anybody (new) out there? Seven new species of Ligidium (Isopoda, Oniscidea, Ligiidae) from the Southern Appalachians, eastern North America. European Journal of Taxonomy 976: 133-170, DOI: 10.5852/ejt.2025.976.2783, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2783/12721
03C38795FFF1FFF4FDAD792FFD7BFC83.taxon	etymology	Etymology From the Cherokee Nantahala, the name of the forests where the species lives. A noun in apposition.	en	Recuero, Ernesto, Caterino, Michael S. (2025): Is there anybody (new) out there? Seven new species of Ligidium (Isopoda, Oniscidea, Ligiidae) from the Southern Appalachians, eastern North America. European Journal of Taxonomy 976: 133-170, DOI: 10.5852/ejt.2025.976.2783, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2783/12721
03C38795FFF1FFF4FDAD792FFD7BFC83.taxon	materials_examined	Material examined Holotype USA – North Carolina • ♂; Clay Co., Nantahala N. F., Chunky Gal Trail; 35.1471 ° N, 83.7144 ° W; 1274 m a. s. l.; 7 Jun. 2021; M. Caterino and E. Recuero leg.; GenBank no: OR 169931 (Cox 1); USNM, CUAC 000180794. Paratype USA – North Carolina • 1 ♂; same data as for holotype; CUAC 000180793. Other material examined USA – North Carolina • 1 imm. ♀; same data as for holotype; CUAC 000171282.	en	Recuero, Ernesto, Caterino, Michael S. (2025): Is there anybody (new) out there? Seven new species of Ligidium (Isopoda, Oniscidea, Ligiidae) from the Southern Appalachians, eastern North America. European Journal of Taxonomy 976: 133-170, DOI: 10.5852/ejt.2025.976.2783, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2783/12721
03C38795FFF1FFF4FDAD792FFD7BFC83.taxon	description	Description Body length of adult males 4.9 – 5.5 mm, width (at pereonite 4) 2.4 – 2.6 mm. Color in ethanol of adult males (Fig. 4) dorsally marbled brown and white, with a conspicuous dark brown longitudinal medial stripe running from pereonite 1 to pereonite 7 and a transverse dark brown band in the caudal margin of pereonites; epimera dark brown with a large white patch at the base; pleonites dark brown caudally, lighter in the central parts (showing in pleonites 4 – 5 in dorsal view); telson dark brown with a white anchor-shaped spot; head marbled brown and white; antennae with basal segments white and brown, distally brown with some white marking; uropod brown with a large white spot in the basipodite; pereopods and pleopods white with brown marbling. Immature specimen with the same color pattern but lighter brown. Antennule (Fig. 5 G) three-segmented; first segment about 1.6 times as long as wide, 3 spiniform setae on its distal border; second segment about 3 times as long as wide, also bearing 3 strong spiniform setae in the distal border; third segment small, with blunt apex and a single seta. Antennae (Fig. 5 F) with 5 segmented peduncle, relative size of antennomeres 5> 4> 3> 2> 1; antennal flagellum with 10 – 11 articles. Dactylus of pereopods with outer claw longer than inner one; no sexual dimorphism observed in pereopods 1 (Fig. 5 H) or 7 (Fig. 5 I). Male pleopod 1 endopodite (Fig. 5 A) with obtuse triangular projection, bearing 2 strong, blunt setae, 0.4 – 0.8 times as long as endopodite. Pleopod 1 exopodite (Fig. 5 B) with rounded caudal margin bearing 2 strong, blunt setae, 0.4 – 0.6 times as long as exopodite. Male pleopod 2 exopodite (Fig. 5 C) 1.6 – 1.7 times as wide as long; inner and frontal margins with limited pilose setation dense, present also along outer caudal corner; caudal margin markedly convex. Male pleopod 2 endopodite (Fig. 5 D – E) with rounded tip and a soft, blunt, robust excrescence occupying most of inner part of tip and projecting outwards; inner margin with minute spiniform setae, no spines at the base. Telson (Fig. 4) with caudal margin obtusely produced, convex in immature female. Uropod (Fig. 5 J) with exopodite and endopodite subequal in length when not broken, 1.5 – 1.7 times as long as basipodite.	en	Recuero, Ernesto, Caterino, Michael S. (2025): Is there anybody (new) out there? Seven new species of Ligidium (Isopoda, Oniscidea, Ligiidae) from the Southern Appalachians, eastern North America. European Journal of Taxonomy 976: 133-170, DOI: 10.5852/ejt.2025.976.2783, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2783/12721
03C38795FFF1FFF4FDAD792FFD7BFC83.taxon	distribution	Distribution The species is so far only known from its type locality (Fig. 4). All specimens were collected in broadleaf forest leaf litter samples.	en	Recuero, Ernesto, Caterino, Michael S. (2025): Is there anybody (new) out there? Seven new species of Ligidium (Isopoda, Oniscidea, Ligiidae) from the Southern Appalachians, eastern North America. European Journal of Taxonomy 976: 133-170, DOI: 10.5852/ejt.2025.976.2783, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2783/12721
03C38795FFF1FFF4FDAD792FFD7BFC83.taxon	discussion	Remarks Ligidium nantahala sp. nov. corresponds with Ligidium sp. 2 in Recuero & Caterino (2024 a). It is not closely related to any of the other species studied here (Fig. 1 A). Morphologically, the presence of a projecting excrescenceat the tip of male pleopod 2 endopodite resembles that observed in L. blueridgensis, L. pacolet sp. nov. and L. whiteoak sp. nov. However, the disposition of the excrescence is different in all four species. In L. nantahala it covers the whole inner part of the tip ventrally, and projects outwards. In L. blueridgensis the excrescence is more slender, and projects from the inner tip corner rising near the margin dorsally, and projecting caudally and inwards. Ligidium pacolet has also a more slender excrescence rising medially or near the inner corner from the end of the tip, projecting caudally or inwards. In L. whiteoak, the excrescence is broad, almost as much as the endopodite stem, rising dorsally and medially from the caudal margin of the tip and projecting caudally and slightly outwards. Ligidium nantahala differs also in the triangular shape of male pleopod 1 endopodite projection, squarish in the other mentioned species. All three gene fragments analyzed are informative to identify this species, with high genetic p-distances for Cox 1 and even for the conserved NaK and 28 S fragments (Supp. file 1), forming a monophyletic clade in all cases.	en	Recuero, Ernesto, Caterino, Michael S. (2025): Is there anybody (new) out there? Seven new species of Ligidium (Isopoda, Oniscidea, Ligiidae) from the Southern Appalachians, eastern North America. European Journal of Taxonomy 976: 133-170, DOI: 10.5852/ejt.2025.976.2783, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2783/12721
03C38795FFFCFFF3FDB8796CFC97FC85.taxon	description	urn: lsid: zoobank. org: act: 65 FAB 813 - B 5 FB- 4865 - 9 F 36 - 925 FC 2 C 1 DCA 2 Figs 6 – 7	en	Recuero, Ernesto, Caterino, Michael S. (2025): Is there anybody (new) out there? Seven new species of Ligidium (Isopoda, Oniscidea, Ligiidae) from the Southern Appalachians, eastern North America. European Journal of Taxonomy 976: 133-170, DOI: 10.5852/ejt.2025.976.2783, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2783/12721
03C38795FFFCFFF3FDB8796CFC97FC85.taxon	diagnosis	Diagnosis This new species is characterized by a male pleopod 2 endopodite distally broadened, with rounded tip presenting a short, squarish projection on its distal margin. It also can be identified from all other Appalachian species based on molecular data, representing a distinct evolutionary lineage.	en	Recuero, Ernesto, Caterino, Michael S. (2025): Is there anybody (new) out there? Seven new species of Ligidium (Isopoda, Oniscidea, Ligiidae) from the Southern Appalachians, eastern North America. European Journal of Taxonomy 976: 133-170, DOI: 10.5852/ejt.2025.976.2783, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2783/12721
03C38795FFFCFFF3FDB8796CFC97FC85.taxon	etymology	Etymology From the latin ‘ protubero ’, meaning ‘ protuberance’, referring to the characteristic projection in the male pleopod 2 endopodite.	en	Recuero, Ernesto, Caterino, Michael S. (2025): Is there anybody (new) out there? Seven new species of Ligidium (Isopoda, Oniscidea, Ligiidae) from the Southern Appalachians, eastern North America. European Journal of Taxonomy 976: 133-170, DOI: 10.5852/ejt.2025.976.2783, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2783/12721
03C38795FFFCFFF3FDB8796CFC97FC85.taxon	materials_examined	Material examined Holotype USA – North Carolina • ♂; Swain Co., Great Smoky Mountains N. P., Payne Creek at Lakeshore Trail; 35.4855 ° N, 83.8028 ° W; 553 m a. s. l.; 12 Apr. 2022; M. Caterino, E. Recuero, A. Haberski and P. Wooden leg.; GenBank no: OR 169915 (Cox 1); USNM, CUAC 000171349. Paratypes USA – South Carolina • 1 ♂; Pickens Co., Chimney Top Gap; 35.0644 ° N, 82.7953 ° W; 781 m a. s. l.; 23 Mar. 2023; C. W. Harden leg.; CUAC 000177090 • 1 ♀; same data as for preceding; CUAC 000177089 • 1 ovi ♀; same data as for preceding; CUAC 000177088. Other material examined USA – North Carolina • 2 ♀♀; Macon Co., Nantahala N. F., Van Hook Glade Campground; 35.0783 ° N, 83.245 ° W; 1006 m a. s. l.: 21 Apr. 2022; S. Payne leg.; CUAC 000171316, CUAC 000171317 • 1 ♀; Clay Co., Nantahala N. F., Tusquitee Bald; 35.1467 ° N, 83.7146 ° W; 1262 m a. s. l.; 1 Sep. 2020; M. Caterino and F. Etzler leg.; CUAC 000138058.	en	Recuero, Ernesto, Caterino, Michael S. (2025): Is there anybody (new) out there? Seven new species of Ligidium (Isopoda, Oniscidea, Ligiidae) from the Southern Appalachians, eastern North America. European Journal of Taxonomy 976: 133-170, DOI: 10.5852/ejt.2025.976.2783, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2783/12721
03C38795FFFCFFF3FDB8796CFC97FC85.taxon	description	Description Body length of males 6.3 – 6.5 mm, of females 6.8 – 8.6 mm; width (at pereonite 4) of males 2.8 – 3, of females 3.3 – 4.6 mm. Color in ethanol (Fig. 6) dorsally marbled brown and white, with a conspicuous dark brown longitudinal medial stripe running from pereonite 1 to pereonite 7; caudal margin of pereonites with no defined dark brown bands; epimera brown with large white spots in the outer surface an at the base; pleonites brown to dark brown with conspicuous white spots; telson with two large white spots laterally, and a white anchor-shaped spot caudally; head marbled brown and white, white above the eyes; antennae brown with white spots in basal segments and a white setae bundle in the flagellum; uropod brown with iner part of basipodite white; pereopods brown and white, pleopods mostly white. Antennule (Fig. 7 G) three-segmented; first segment about 1.4 times as long as wide, 3 spiniform setae on its distal border; second segment about 2.5 times as long as wide, also bearing 3 strong spiniform setae on the distal border; third segment with blunt apex and a single seta. Antennae (Fig. 7 F) with 5 segmented peduncle, relative size of antennomeres 5> 4> 3> 2> 1; antennal flagellum with 11 – 13 articles. Dactylus of pereopods with outer claw longer than inner one; no sexual dimorphism observed in pereopods 1 (Fig. 7 H) and 7 (Fig. 7 I). Male pleopod 1 endopodite (Fig. 7 A) with obtuse triangular projection, bearing 2 – 3 strong, blunt setae, up to 1.5 times as long as endopodite; caudal and inner margins with pilose setae. Pleopod 1 exopodite (Fig. 7 B) with broadly rounded caudal margin bearing 3 – 4 strong, blunt setae, 0.7 – 0.9 times as long as exopodite. Male pleopod 2 exopodite (Fig. 7 C) 2 times as wide as long; inner and most part of frontal margin with dense, hairy setation, as in outer caudal corner; caudal margin with 5 – 6 short but strong setae. Male pleopod 2 endopodite (Fig. 7 D – E) distally broadened, with rounded tip presenting a row of small denticles in the inner corner and a short, squarish terminal projection in the distal margin; inner margin with minute spiniform setae, larger and more conspicuous at base. Telson (Fig. 6) with caudal margin obtusely produced. Uropod (Fig. 7 J) with endopodite 1.3 – 1.4 times as long as exopodite when not broken, and about 2 times as long as basipodite.	en	Recuero, Ernesto, Caterino, Michael S. (2025): Is there anybody (new) out there? Seven new species of Ligidium (Isopoda, Oniscidea, Ligiidae) from the Southern Appalachians, eastern North America. European Journal of Taxonomy 976: 133-170, DOI: 10.5852/ejt.2025.976.2783, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2783/12721
03C38795FFFCFFF3FDB8796CFC97FC85.taxon	distribution	Distribution The species has been found scattered in the southern Blue Ridge Mountains (Fig. 6), at low to midaltitudes. All studied specimens have been collected in leaf litter from hardwood forests.	en	Recuero, Ernesto, Caterino, Michael S. (2025): Is there anybody (new) out there? Seven new species of Ligidium (Isopoda, Oniscidea, Ligiidae) from the Southern Appalachians, eastern North America. European Journal of Taxonomy 976: 133-170, DOI: 10.5852/ejt.2025.976.2783, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2783/12721
03C38795FFFCFFF3FDB8796CFC97FC85.taxon	discussion	Remarks Ligidium protuberans sp. nov. corresponds with Ligidium sp. 3 in Recuero & Caterino (2024 a). This species is closely related to L. enotahensis sp. nov. and L. schultzi sp. nov. (Fig. 1 A), but it can be easily diagnosed from them by the conspicuous short projection present in the male pleopod 2 endopodite. All three gene fragments analyzed are informative to identify this species, with high interspecific p-distances for Cox 1, and low to medium in the conserved NaK and 28 S fragments (Supp. file 1), forming a reciprocally monophyletic clade in all cases.	en	Recuero, Ernesto, Caterino, Michael S. (2025): Is there anybody (new) out there? Seven new species of Ligidium (Isopoda, Oniscidea, Ligiidae) from the Southern Appalachians, eastern North America. European Journal of Taxonomy 976: 133-170, DOI: 10.5852/ejt.2025.976.2783, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2783/12721
03C38795FFFBFFEEFDAB796DFABDFD0C.taxon	description	urn: lsid: zoobank. org: act: 3 EAEAB 27 - 5 C 45 - 4183 - 9 B 5 D- 70 FB 794 D 8094 Figs 8 – 9	en	Recuero, Ernesto, Caterino, Michael S. (2025): Is there anybody (new) out there? Seven new species of Ligidium (Isopoda, Oniscidea, Ligiidae) from the Southern Appalachians, eastern North America. European Journal of Taxonomy 976: 133-170, DOI: 10.5852/ejt.2025.976.2783, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2783/12721
03C38795FFFBFFEEFDAB796DFABDFD0C.taxon	diagnosis	Diagnosis This new species differs from all other Appalachian species in its male pleopod 2 endopodite having an acuminate tip, strongly projected posteriorly and outwards. It has a distinctive male pleopod 1 endopodite, showing a poorly defined, broadly triangular projection, and a male pleopod 1 exopodite with its caudal margin flattened. It also differs from all other Appalachian species based on molecular data, representing a distinct evolutionary lineage with no close relationships among the studied taxa.	en	Recuero, Ernesto, Caterino, Michael S. (2025): Is there anybody (new) out there? Seven new species of Ligidium (Isopoda, Oniscidea, Ligiidae) from the Southern Appalachians, eastern North America. European Journal of Taxonomy 976: 133-170, DOI: 10.5852/ejt.2025.976.2783, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2783/12721
03C38795FFFBFFEEFDAB796DFABDFD0C.taxon	etymology	Etymology A noun in apposition, ‘ gadalutsi ’ is the Cherokee word that originated the name Cataloochee, and consequently the origin of the name of the type locality, Big Cataloochee Mountain.	en	Recuero, Ernesto, Caterino, Michael S. (2025): Is there anybody (new) out there? Seven new species of Ligidium (Isopoda, Oniscidea, Ligiidae) from the Southern Appalachians, eastern North America. European Journal of Taxonomy 976: 133-170, DOI: 10.5852/ejt.2025.976.2783, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2783/12721
03C38795FFFBFFEEFDAB796DFABDFD0C.taxon	materials_examined	Material examined Holotype USA – North Carolina • ♂; Haywood Co., Great Smoky Mountains N. P., Big Cataloochee Mt; 35.6675 ° N, 83.1805 ° W; 1703 m a. s. l.; 14 Jul. 2020; M. Caterino and F. Etzler leg.; GenBank no: PP 737148 (Cox 1); USNM, CUAC 000138053. Other material examined USA – North Carolina • 1 ♀; Clay Co., Nantahala N. F., Chunky Gal Trail; 35.1471 ° N, 83.7144 ° W; 1274 m a. s. l.; 6 Jul. 2021; M. Caterino and E. Recuero leg.; CUAC 000171281. – Tennessee • 1 ♀; Sevier Co., Smoky Mountains N. P., Mount LeConte (Alum Cave Trail); 35.6382 ° N, 83.4387 ° W; 1317 m a. s. l.; 28 Sep. 2021; M. Caterino and E. Recuero leg.; CUAC 000171324.	en	Recuero, Ernesto, Caterino, Michael S. (2025): Is there anybody (new) out there? Seven new species of Ligidium (Isopoda, Oniscidea, Ligiidae) from the Southern Appalachians, eastern North America. European Journal of Taxonomy 976: 133-170, DOI: 10.5852/ejt.2025.976.2783, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2783/12721
03C38795FFFBFFEEFDAB796DFABDFD0C.taxon	description	Description Body length of holotype 4.9 mm, width (at 4 pereonite) 2.4 mm; body length of females 5.5 – 7 mm, width 2.5 – 3.6 mm. Color in ethanol (Fig. 8) of holotype dorsally marbled brown and white, with a conspicuous dark brown longitudinal medial stripe running from pereonite 1 to pereonite 7 and a transversal dark brown band in the caudal margin of pereonites; epimera dark brown with a large white patch at the base; pleonites dark brown with lateral white spots; telson dark brown with a white anchor-shaped spot; head and antennae marbled brown and white, distal segments of antennae darker brown with white setae bundle in flagellum; uropod brown with a white spot in the inner part of basipodite; pereopods and pleopods white with brown marbling. Among females one resembles the holotype in coloration, the other is dorsally almost black with large white spots from head to telson. Antennule (Fig. 9 G) three-segmented; first segment about 1.5 – 1.7 times as long as wide, 2 – 3 spiniform setae in its distal border; second segment about 2.3 – 2.5 times as long as wide, bearing 3 strong spiniform setae in the distal border; third segment with blunt apex and 2 setae. Antennae (Fig. 9 F) with 5 segmented peduncle, relative size of antennomeres 5> 4> 3> 2> 1; antennal flagellum with 11 articles. Dactylus of pereopods with outer claw longer than inner one, inner one reduced in pereopod 1; no sexual dimorphism observed in pereopods 1 (Fig. 9 H) and 7 (Fig. 9 I). Male pleopod 1 endopodite (Fig. 9 A) with short, blunt projection, bearing a single strong, blunt seta, 0.8 times as long as endopodite; caudal and inner margins with pilose setae. Pleopod 1 exopodite (Fig. 9 B) with flat caudal margin bearing 4 strong, blunt setae, up to 0.8 times as long as exopodite. Male pleopod 2 exopodite (Fig. 9 C) 1.9 times as wide as long; inner and half of the frontal margin with dense, hairy setation, as in outer caudal corner; caudal margin evenly convex, with 2 short, strong setae in the outer corner. Male pleopod 2 endopodite (Fig. 9 D – E) with a broad and acuminate tip, projected posteriorly-laterally, with small setae in the interior margin; inner margin of the peduncle with minute spiniform setae, no spines at base. Telson (Fig. 8) with caudal margin slightly concave in the sides and obtusely produced in the middle. Uropod (Fig. 9 J) with endopodite 1.3 – 1.4 times as long as exopodite when not broken, and about 2 times as long as basipodite.	en	Recuero, Ernesto, Caterino, Michael S. (2025): Is there anybody (new) out there? Seven new species of Ligidium (Isopoda, Oniscidea, Ligiidae) from the Southern Appalachians, eastern North America. European Journal of Taxonomy 976: 133-170, DOI: 10.5852/ejt.2025.976.2783, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2783/12721
03C38795FFFBFFEEFDAB796DFABDFD0C.taxon	distribution	Distribution The species has been found in two localities in the Great Smoky Mountains and one at the Nantahala National Forest further south, at mid- to high elevations (Fig. 8). Studied specimens have been collected in leaf litter from conifer and mixed forests.	en	Recuero, Ernesto, Caterino, Michael S. (2025): Is there anybody (new) out there? Seven new species of Ligidium (Isopoda, Oniscidea, Ligiidae) from the Southern Appalachians, eastern North America. European Journal of Taxonomy 976: 133-170, DOI: 10.5852/ejt.2025.976.2783, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2783/12721
03C38795FFFBFFEEFDAB796DFABDFD0C.taxon	discussion	Remarks Ligidium gadalutsi sp. nov. corresponds with Ligidium sp. 4 in Recuero & Caterino (2024 a). It is not closely related to any of the studied Appalachian species (Fig. 1 A). The shape of the male pleopod 2 endopodite somewhat resembles that of L. mucronatum, which could indicate a shared ancestry. However, in L. mucronatum the tip is shorter and blunt, while much longer and acuminate in L. gadalutsi. This new species shows the highest genetic p-distances compared to all other studied species (Supp. file 1).	en	Recuero, Ernesto, Caterino, Michael S. (2025): Is there anybody (new) out there? Seven new species of Ligidium (Isopoda, Oniscidea, Ligiidae) from the Southern Appalachians, eastern North America. European Journal of Taxonomy 976: 133-170, DOI: 10.5852/ejt.2025.976.2783, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2783/12721
03C38795FFE6FFEDFD5E7895FC97FC1D.taxon	description	urn: lsid: zoobank. org: act: 6 A 3 E 7 CC 8 - 1 EF 9 - 4451 - 889 E- 81878 B 6 B 5250 Figs 10 – 11	en	Recuero, Ernesto, Caterino, Michael S. (2025): Is there anybody (new) out there? Seven new species of Ligidium (Isopoda, Oniscidea, Ligiidae) from the Southern Appalachians, eastern North America. European Journal of Taxonomy 976: 133-170, DOI: 10.5852/ejt.2025.976.2783, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2783/12721
03C38795FFE6FFEDFD5E7895FC97FC1D.taxon	diagnosis	Diagnosis This new species is characterized by its male pleopod 2 endopodite presenting a soft, long, slender excrescence rising ventrally in the distal margin of the tip, close to its inner corner, and projected caudally or inwards. It also can be identified from all other Appalachian species based on molecular data, representing a distinct evolutionary lineage.	en	Recuero, Ernesto, Caterino, Michael S. (2025): Is there anybody (new) out there? Seven new species of Ligidium (Isopoda, Oniscidea, Ligiidae) from the Southern Appalachians, eastern North America. European Journal of Taxonomy 976: 133-170, DOI: 10.5852/ejt.2025.976.2783, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2783/12721
03C38795FFE6FFEDFD5E7895FC97FC1D.taxon	etymology	Etymology A noun in apposition, in reference to the Pacolet River area where the species has been found.	en	Recuero, Ernesto, Caterino, Michael S. (2025): Is there anybody (new) out there? Seven new species of Ligidium (Isopoda, Oniscidea, Ligiidae) from the Southern Appalachians, eastern North America. European Journal of Taxonomy 976: 133-170, DOI: 10.5852/ejt.2025.976.2783, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2783/12721
03C38795FFE6FFEDFD5E7895FC97FC1D.taxon	materials_examined	Material examined Holotype USA – North Carolina • ♂; Polk Co., North Pacolet River; 35.2221 ° N, 82.3059 ° W; 384 m a. s. l.; 15 Feb. 2022; M. Caterino, E. Recuero, C. Harden and P. Wooden leg.; GenBank no: OR 169911 (Cox 1); USNM, CUAC 000171345. Paratypes USA – North Carolina • 2 ♂♂; same data as for holotype; CUAC 000171341, CUAC 000171342 • 1 ♀; same data as for holotype; CUAC 000171343.	en	Recuero, Ernesto, Caterino, Michael S. (2025): Is there anybody (new) out there? Seven new species of Ligidium (Isopoda, Oniscidea, Ligiidae) from the Southern Appalachians, eastern North America. European Journal of Taxonomy 976: 133-170, DOI: 10.5852/ejt.2025.976.2783, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2783/12721
03C38795FFE6FFEDFD5E7895FC97FC1D.taxon	description	Description Body length of males 5.5 – 6.1 mm, of female 7.9 mm; width (at pereonite 4) of males 2.6 – 2.9, of female 3.6 mm. Color in ethanol (Fig. 10) dorsally dark brown with large off-white spots, with a conspicuous dark brown longitudinal medial stripe running from pereonite 1 to pereonite 7, frequently broken with light spots in the caudal margin of pereonites, which carry dark brown transverse bands; epimera dark brown with large white spots at the base and sometimes smaller white spots in the external margin; pleonites 1 – 4 with two pairs of off-white spots, pereonite 5 (and rarely 4) with a single pair; telson with two large white spots laterally, merging caudally with the white anchor-shaped spot; head marbled brown, with white areas above the eyes; antennae dark brown with large white spots in basal segments and a white setae bundle in the flagellum; uropod brown with inner part of basipodite white; pereopods brown and white, pleopods mostly white. Antennule (Fig. 11 G) three-segmented; first segment about 1.7 – 1.8 times as long as wide, 3 spiniform setae in its distal border; second segment about 2.5 – 2.6 times as long as wide, also bearing 3 strong spiniform setae in the distal border; third segment with rounded apex and a single seta. Antennae (Fig. 11 F) with 5 segmented peduncle, relative size of antennomeres 5> 4> 3> 2 = 1; antennal flagellum with 11 – 12 articles. Dactylus of pereopods with outer claw longer than inner one; no sexual dimorphism observed in pereopods 1 (Fig. 11 H) and 7 (Fig. 11 I). Male pleopod 1 endopodite (Fig. 11 A) with narrow projection truncated distally, bearing 3 – 4 strong, blunt setae, up to 1.3 times as long as endopodite; caudal and inner margins with pilose setae. Pleopod 1 exopodite (Fig. 11 B) with broadly rounded caudal margin bearing 3 – 5 strong, blunt setae, 0.8 – 0.9 times as long as exopodite; external margin with 2 – 3 short setae, a few short piliform setae in the frontal external corner and in the inner margin. Male pleopod 2 exopodite (Fig. 11 C) 2.1 – 2.3 times as wide as long; inner and most part of frontal margin with dense, hairy setation; caudal margin convex. Male pleopod 2 endopodite (Fig. 11 D – E) with a broad tip, bearing a soft, long, slender excrescence rising ventrally at its distal margin near the inner corner; inner margin of stem with minute spiniform setae, denser at the base and absent well before the tip. Telson (Fig. 10) with caudal margin obtusely produced, with a sinuate profile. Uropod (Fig. 11 J) with endopodite 1.3 – 1.4 times as long as exopodite when not broken, and about 2 times as long as basipodite.	en	Recuero, Ernesto, Caterino, Michael S. (2025): Is there anybody (new) out there? Seven new species of Ligidium (Isopoda, Oniscidea, Ligiidae) from the Southern Appalachians, eastern North America. European Journal of Taxonomy 976: 133-170, DOI: 10.5852/ejt.2025.976.2783, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2783/12721
03C38795FFE6FFEDFD5E7895FC97FC1D.taxon	distribution	Distribution The species is known only from its type locality, by the North Pacolet River near Saluda, North Carolina, along the boundary between the Blue Ridge Mountains and the Piedmont Region (Fig. 10). All studied specimens have been collected in moist leaf litter from deciduous forest.	en	Recuero, Ernesto, Caterino, Michael S. (2025): Is there anybody (new) out there? Seven new species of Ligidium (Isopoda, Oniscidea, Ligiidae) from the Southern Appalachians, eastern North America. European Journal of Taxonomy 976: 133-170, DOI: 10.5852/ejt.2025.976.2783, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2783/12721
03C38795FFE6FFEDFD5E7895FC97FC1D.taxon	discussion	Remarks Ligidium pacolet sp. nov. corresponds with one of the two lineages named as Ligidium sp. 5 in Recuero & Caterino (2024 a). It was clustered together with L. whiteoak sp. nov. in that paper given their morphological and genetic affinities, although species delimitation methods recognized the two species using multilocus data for species delimitation (BPP), or even three with Cox 1 single-locus species delimitation methods (ASAP, mPTP) (Supp. file 1). It belongs to a clade including L. blueridgensis and L. whiteoak, being genetically closer to the latter (Fig. 1 A). They are also morphologically close, all presenting an excrescence projecting from the male pleopod 2 endopodite; in L. pacolet the excrescence is slender and rises medially or near the inner corner from the margin of the tip, projecting caudally or inwards. In L. blueridgensis the excrescence is also slender, projecting from the inner tip corner rising from the dorsal surface near the margin, and projecting caudally and inwards. Ligidium whiteoak presents a broader excrescence, almost as wide as the endopodite stem, rising dorsally and medially from the caudal margin of the tip and projecting caudally and slightly outwards. The stem of male pleopod 2 endopodite in L. pacolet sp. nov. is slightly broadened in the distal third, while in L. blueridgensis and L. whiteoak its width is constant. The projection of male pleopod 1 endopodite is narrower in L. pacolet than in the other two species. All three gene fragments analyzed are informative to identify this species, with high genetic p-distances for Cox 1, moderate to high for NaK and 28 S fragments (Supp. file 1), forming a reciprocally monophyletic clade in all cases.	en	Recuero, Ernesto, Caterino, Michael S. (2025): Is there anybody (new) out there? Seven new species of Ligidium (Isopoda, Oniscidea, Ligiidae) from the Southern Appalachians, eastern North America. European Journal of Taxonomy 976: 133-170, DOI: 10.5852/ejt.2025.976.2783, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2783/12721
03C38795FFE5FFE8FDAA7985FBD0FDAC.taxon	description	urn: lsid: zoobank. org: act: CAB 4 ACD 2 - 999 C- 4 C 0 A- 9 BD 7 - A 196 E 4 B 4 EBDF Figs 12 – 13	en	Recuero, Ernesto, Caterino, Michael S. (2025): Is there anybody (new) out there? Seven new species of Ligidium (Isopoda, Oniscidea, Ligiidae) from the Southern Appalachians, eastern North America. European Journal of Taxonomy 976: 133-170, DOI: 10.5852/ejt.2025.976.2783, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2783/12721
03C38795FFE5FFE8FDAA7985FBD0FDAC.taxon	diagnosis	Diagnosis Ligidium whiteoak sp. nov. is diagnosed from other Appalachian species by the presence in the male pleopod 2 endopodite of a broad excrescence, almost as broad as the endopodite stem, rising dorsally and medially from the caudal margin of the tip and projecting caudally and slightly outwards. It also can be identified from all other Appalachian species based on molecular data, representing a distinct evolutionary lineage.	en	Recuero, Ernesto, Caterino, Michael S. (2025): Is there anybody (new) out there? Seven new species of Ligidium (Isopoda, Oniscidea, Ligiidae) from the Southern Appalachians, eastern North America. European Journal of Taxonomy 976: 133-170, DOI: 10.5852/ejt.2025.976.2783, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2783/12721
03C38795FFE5FFE8FDAA7985FBD0FDAC.taxon	etymology	Etymology A noun in apposition, in reference to the species type locality.	en	Recuero, Ernesto, Caterino, Michael S. (2025): Is there anybody (new) out there? Seven new species of Ligidium (Isopoda, Oniscidea, Ligiidae) from the Southern Appalachians, eastern North America. European Journal of Taxonomy 976: 133-170, DOI: 10.5852/ejt.2025.976.2783, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2783/12721
03C38795FFE5FFE8FDAA7985FBD0FDAC.taxon	materials_examined	Material examined Holotype USA – Tennessee • ♂; Blount Co., Great Smoky Mountains N. P., Whiteoak Sink area; 35.6362 ° N, 83.7412 ° W; 536 m a. s. l.; 27 Oct. 2021; M. Caterino, A. Haberski and P. Wooden leg.; GenBank no: OR 169835 (Cox 1); USNM, CUAC 000171267. Paratypes USA – Tennessee • 3 ♂♂; same data as for holotype; CUAC 000171268, CUAC 000180800, CUAC 000180801 • 5 ♀♀; same data as for holotype; CUAC 000171269 to CUAC 000171273.	en	Recuero, Ernesto, Caterino, Michael S. (2025): Is there anybody (new) out there? Seven new species of Ligidium (Isopoda, Oniscidea, Ligiidae) from the Southern Appalachians, eastern North America. European Journal of Taxonomy 976: 133-170, DOI: 10.5852/ejt.2025.976.2783, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2783/12721
03C38795FFE5FFE8FDAA7985FBD0FDAC.taxon	description	Description Body length of males 4.9 – 5.5 mm, of females 4.9 – 5 mm; width (at pereonite 4) of males 2.3 – 2.5, of females 2.4 – 2.6 mm. Color in ethanol (Fig. 12) dorsally brown to dark brown with off-white and greenish spots, a conspicuous dark brown longitudinal medial stripe running from pereonite 1 to pereonite 7, more or less broken with light spots in the caudal margin of pereonites; not all pereonites with darker transverse bands; epimera brown and off-white, with large white spots at the base; pleonites with a pair of lateral off-white spots, medially brown sometimes with off-white spots; telson with two white spots laterally, and a white anchor-shaped spot; head marbled brown; antennae dark brown with large white spots in basal segments and a white setae bundle in the flagellum; uropod brown, dorsally with inner part of basipodite white; pereopods brown and white, pleopod exopodites brown. Antennule (Fig. 13 G) three-segmented; first segment about 1.7 – 1.8 times as long as wide, 3 spiniform setae in its distal border; second segment about 2.6 – 2.7 times as long as wide, also bearing 3 strong spiniform setae in the distal border; third segment with rounded apex and up to 3 setae. Antennae (Fig. 13 F) with 5 segmented peduncle, relative size of antennomeres 5> 4> 3> 2> 1; antennal flagellum with 10 – 11 articles. Dactylus of pereopods with outer claw longer than inner one; no sexual dimorphism observed in pereopods 1 (Fig. 13 H) and 7 (Fig. 13 I). Pereopod 7 ischium broadened medially. Male pleopod 1 endopodite (Fig. 13 A) with narrow projection truncated distally, bearing 2 – 3 strong, blunt setae, about as long as endopodite; most part of caudal and inner margins with pilose setae. Pleopod 1 exopodite (Fig. 13 B) with rounded caudal margin bearing 2 – 3 strong, blunt setae, 0.6 – 0.7 times as long as exopodite; external margin with 3 – 4 short setae. Male pleopod 2 exopodite (Fig. 13 C) 1.8 – 2 times as wide as long; inner and most part of frontal margin with dense, hairy setation, as in outer caudal corner; caudal margin convex, with 3 – 4 short, but strong setae. Male pleopod 2 endopodite (Fig. 13 D – E) with narrow, rounded tip, bearing a soft, long excrescence rising medially, almost as broad as the stem; inner margin with minute spiniform setae. Telson (Fig. 12) with caudal margin obtusely produced, subtriangular. Uropod (Fig. 13 J) with endopodite 1.4 – 1.5 times as long as exopodite when not broken, and about 2 times as long as basipodite.	en	Recuero, Ernesto, Caterino, Michael S. (2025): Is there anybody (new) out there? Seven new species of Ligidium (Isopoda, Oniscidea, Ligiidae) from the Southern Appalachians, eastern North America. European Journal of Taxonomy 976: 133-170, DOI: 10.5852/ejt.2025.976.2783, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2783/12721
03C38795FFE5FFE8FDAA7985FBD0FDAC.taxon	distribution	Distribution The species is known only from its type locality, at low elevations near the western edge of the Great Smoky Mountains (Fig. 12). All studied specimens have been collected in moist leaf litter from deciduous forest.	en	Recuero, Ernesto, Caterino, Michael S. (2025): Is there anybody (new) out there? Seven new species of Ligidium (Isopoda, Oniscidea, Ligiidae) from the Southern Appalachians, eastern North America. European Journal of Taxonomy 976: 133-170, DOI: 10.5852/ejt.2025.976.2783, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2783/12721
03C38795FFE5FFE8FDAA7985FBD0FDAC.taxon	discussion	Remarks Ligidium whiteoak sp. nov. corresponds with one of the two lineages referred to as Ligidium sp. 5 in Recuero & Caterino (2024 a). See remarks for Ligidium pacolet sp. nov.	en	Recuero, Ernesto, Caterino, Michael S. (2025): Is there anybody (new) out there? Seven new species of Ligidium (Isopoda, Oniscidea, Ligiidae) from the Southern Appalachians, eastern North America. European Journal of Taxonomy 976: 133-170, DOI: 10.5852/ejt.2025.976.2783, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2783/12721
03C38795FFE0FFE7FDA27873FB35F93F.taxon	description	urn: lsid: zoobank. org: act: E 438 B 528 - 4175 - 4 BF 6 - 9308 - EA 9 ECE 8 C 9 D 41 Figs 14 – 15	en	Recuero, Ernesto, Caterino, Michael S. (2025): Is there anybody (new) out there? Seven new species of Ligidium (Isopoda, Oniscidea, Ligiidae) from the Southern Appalachians, eastern North America. European Journal of Taxonomy 976: 133-170, DOI: 10.5852/ejt.2025.976.2783, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2783/12721
03C38795FFE0FFE7FDA27873FB35F93F.taxon	diagnosis	Diagnosis This new species is characterized by the distally broadened male pleopod 2 endopodite, which presents on the caudal margin as a marked notch. Male pleopod 2 exopodite has its frontal margin concave. It also can be distinguished from all other Appalachian species based on molecular data, representing a distinct evolutionary lineage.	en	Recuero, Ernesto, Caterino, Michael S. (2025): Is there anybody (new) out there? Seven new species of Ligidium (Isopoda, Oniscidea, Ligiidae) from the Southern Appalachians, eastern North America. European Journal of Taxonomy 976: 133-170, DOI: 10.5852/ejt.2025.976.2783, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2783/12721
03C38795FFE0FFE7FDA27873FB35F93F.taxon	etymology	Etymology The species is named after George A. Schultz, for his important contributions to our knowledge of the genus Ligidium in North America, and for being the first to illustrate this species from North Carolina.	en	Recuero, Ernesto, Caterino, Michael S. (2025): Is there anybody (new) out there? Seven new species of Ligidium (Isopoda, Oniscidea, Ligiidae) from the Southern Appalachians, eastern North America. European Journal of Taxonomy 976: 133-170, DOI: 10.5852/ejt.2025.976.2783, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2783/12721
03C38795FFE0FFE7FDA27873FB35F93F.taxon	materials_examined	Material examined Holotype USA – North Carolina • ♂; Swain Co., Great Smoky Mountains N. P., off Highway 441, Thomas Divide Trail; 35.5824 ° N, 83.3979 ° W; 1405 m a. s. l.; 12 Mar. 2020; M. Caterino and F. Etzler leg.; GenBank no: OR 169857 (Cox 1); USNM, CUAC 000171290. Paratypes USA – North Carolina • 1 ♂; same data as for holotype; CUAC 000171291 • 4 ♀♀; same data as for holotype; CUAC 000171292 to CUAC 000171295. Other material examined USA – Georgia • 2 ♂♂; Towns Co., Brasstown Bald; 34.8782 ° N, 83.8108 ° W; 1347 m a. s. l.; 17 Nov. 2020; Caterino and F. Etzler leg.; CUAC 000171279, CUAC 000180791 • 1 ♀; same data as for preceding; CUAC 000138052 – North Carolina • 1 ♂; Graham Co., Nantahala N. F., Huckleberry Knob; 35.3210 ° N, 83.9934 ° W; 1674 m a. s. l.; 4 May 2020; M. Caterino and F. Etzler leg.; CUAC 000138050 • 1 ♀, Haywood Co., Balsam Mountain Trail; 35.6425 ° N, 83.2007 ° W; 1565 m a. s. l.; 5 Nov. 2020; M. Caterino and F. Etzler leg.; CUAC 000138054 • 3 ♂♂; Macon Co., Highlands Biological Station, Coker Rhododendron Trail; 35.0558 ° N, 83.1840 ° W; 1191 m a. s. l.; 20 Jun. 2022; P. Marek and L. Kairy leg.; CUAC 000171315, CUAC 000171319, CUAC 000171320 • 1 ♂; Macon Co., Nantahala N. F., Copper Ridge Bald; 35.3270 ° N, 83.3359 ° W; 1535 m a. s. l.; 15 Sep. 2020; M. Caterino and F. Etzler leg.; CUAC 000171277 • 2 ♀♀; same data as for preceding; CUAC 000171278, CUAC 000138046 • 3 ♀♀; Macon Co., Nantahala N. F., Cowee Bald; 35.3270 ° N, 83.3359 ° W; 1503 m a. s. l.; 15 Sep. 2020; F. Etzler, A. Haberski and P. Wooden leg.; CUAC 000138047, CUAC 000171283, CUAC 000171284 • 5 imm.; same data as for preceding; CUAC 000171285 to CUAC 000171289 • 2 ♂♂; Macon Co., Nantahala N. F., Van Hook Glade Campground; 35.0783 ° N, 83.245 ° W; 1018 m a. s. l.; 21 Jun. 2022; CUAC 000171321, CUAC 000171322 • 1 ♀; same data as for preceding; CUAC 000171318 • 2 ♀♀; Transylvania Co., Pisgah Forest; 35.3291 ° N, 82.789 ° W; 910 m a. s. l.; 23 Dec. 2021; E. Recuero and P. C. Rodríguez-Flores leg.; CUAC 000171302, CUAC 000171303 • 1 ♀; Transylvania Co., Pisgah National Forest, Hwy 215; 35.2910 ° N, 82.9133 ° W; 1561 m a. s. l.; 8 May 2018; M. Caterino, R. Kucuk and L. Cushman leg.; CUAC 000171304 • 2 ♂♂; same data as for preceding; CUAC 000171305, CUAC 000171306 • 1 ovi. ♀; Clay Co., Nantahala N. F., Tusquitee Bald; 35.1415 ° N, 83.7273 ° W; 1582 m a. s. l.; 6 Jul. 2021; M. Caterino and E. Recuero leg.; CUAC 000165724. – South Carolina • 1 ♀; Oconee Co., Sumter N. F., Walhalla Hatchery; 34.9853 ° N, 83.0731 ° W; 765 m a. s. l.; 21 Jan. 2023; C. W. Harden leg.; CUAC 000177091 • 1 ♂; same data as for preceding; CUAC 000177092 – Tennessee • 1 ♂; Blount Co., Great Smoky Mountains N. P., Rich Mountain Gap; 35.645 ° N, 83.81 ° W; 600 m a. s. l.; 27 Oct. 2021; M. Caterino, A. Haberski and P. Wooden leg.; CUAC 000171264 • 2 ♀♀; same data as for preceding; CUAC 000171265, CUAC 000171266 • 3 ♂♂; Sevier Co., Great Smoky Mountains N. P., off Highway 441; 35.6240 ° N, 83.4163 ° W; 1394 m a. s. l.; 12 Mar. 2020; M. Caterino and F. Etzler leg.; CUAC 000138081, CUAC 000171297, CUAC 000171298 • 2 ♀♀; same data as for preceding; CUAC 000171300, CUAC 000171301 • 1 imm.; same data as for preceding; CUAC 000171296.	en	Recuero, Ernesto, Caterino, Michael S. (2025): Is there anybody (new) out there? Seven new species of Ligidium (Isopoda, Oniscidea, Ligiidae) from the Southern Appalachians, eastern North America. European Journal of Taxonomy 976: 133-170, DOI: 10.5852/ejt.2025.976.2783, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2783/12721
03C38795FFE0FFE7FDA27873FB35F93F.taxon	description	Description Body length of males 4.6 – 5.7 mm, of females 4.7 – 5.7 mm; width (at pereonite 4) of males 2.5 – 2.8, of females 2.6 – 2.9 mm. Color in ethanol (Fig. 14) dorsally brown with large off-white, a dark brown longitudinal medial stripe running from pereonite 1 to pereonite 7, frequently broadening caudally; pereonites with darker transverse bands in the caudal margin, frequently with clearer spots; epimera brown marbled with off-white, with large white spots at the base; pleonites brown with a pair of more or less defined off-white spots, paramedian in pleonites 1 and 2, more lateral in pleonites 3 – 5; telson with two white spots laterally, and a white anchor-shaped spot; head marbled brown, sometimes with white area above eyes; antennae brown, sometimes with large white spots in basal segments, and a white setae bundle in the flagellum; uropod brown, dorsally with inner part of basipodite white; pereopods brown and white, pleopod exopodites white, often with inner and frontal parts brown. Antennule (Fig. 15 G) three-segmented; first segment about 1.7 – 1.8 times as long as wide, 2 – 3 spiniform setae in its distal border; second segment about 2.5 – 2.6 times as long as wide, bearing 2 – 3 strong spiniform setae in the distal border; third segment with blunt apex and 1 or no setae. Antennae (Fig. 15 F) with 5 segmented peduncle, relative size of antennomeres 5> 4> 3 = 2> 1; antennal flagellum with 10 – 12 articles. Dactylus of pereopods with outer claw longer than inner one; no sexual dimorphism observed in pereopods 1 (Fig. 15 H) and 7 (Fig. 15 I). Male pleopod 1 endopodite (Fig. 15 A) with narrow projection obliquely truncated distally, bearing 2 strong, blunt setae, up to 1.2 times as long as endopodite; part of caudal and inner margins with pilose setae. Pleopod 1 exopodite (Fig. 15 B) with broadly rounded caudal margin bearing 2 – 3 strong, blunt setae, up to 0.5 – 0.7 times as long as exopodite; external margin with 2 – 3 very short but strong setae. Male pleopod 2 exopodite (Fig. 15 C) 1.8 – 1.9 times as wide as long; inner, outer and part of frontal margin with dense, hairy setation; caudal margin strongly convex, with 2 – 4 very short, but strong setae; frontal margin concave. Male pleopod 2 endopodite (Fig. 15 D – E) with unevenly rounded tip, presenting a marked notch or indentation; inner margin with minute spiniform setae denser near the base. Telson (Fig. 14) with caudal margin obtusely produced. Uropod (Fig. 15 J) with long endopodite, 1.6 – 1.8 times as long as exopodite when not broken, and about 2.1 – 2.2 times as long as basipodite.	en	Recuero, Ernesto, Caterino, Michael S. (2025): Is there anybody (new) out there? Seven new species of Ligidium (Isopoda, Oniscidea, Ligiidae) from the Southern Appalachians, eastern North America. European Journal of Taxonomy 976: 133-170, DOI: 10.5852/ejt.2025.976.2783, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2783/12721
03C38795FFE0FFE7FDA27873FB35F93F.taxon	distribution	Distribution The species is widely distributed in the southern part of the Blue Ridge Mountains, with all known records found west of the Asheville Depression (Fig. 14). Studied specimens have been collected in moist leaf litter from conifer, deciduous, and mixed forest, as well as under dead logs and stones, usually in damp places near streams. From 600 m to 1674 m a. s. l., most frequently above 1000 m.	en	Recuero, Ernesto, Caterino, Michael S. (2025): Is there anybody (new) out there? Seven new species of Ligidium (Isopoda, Oniscidea, Ligiidae) from the Southern Appalachians, eastern North America. European Journal of Taxonomy 976: 133-170, DOI: 10.5852/ejt.2025.976.2783, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2783/12721
03C38795FFE0FFE7FDA27873FB35F93F.taxon	discussion	Remarks Ligidium schultzi sp. nov. corresponds with Ligidium sp. 6 in Recuero & Caterino (2024 a). The shape of male pleopod 2 endopodite of this species was first illustrated by Schultz (1982), as a morphological variant of what he identified as L. elrodii, and indicated the need for further taxonomic study to determine the specific status. The new species is closely related to L. enotahensis sp. nov. and L. protuberans sp. nov. (Fig. 1 A; see Remarks for those species). All three gene fragments analyzed are informative to identify this species, with high genetic p-distances among the mentioned taxa for Cox 1, moderate for 28 S and low for NaK (Supp. file 1), forming reciprocally monophyletic clades in all cases.	en	Recuero, Ernesto, Caterino, Michael S. (2025): Is there anybody (new) out there? Seven new species of Ligidium (Isopoda, Oniscidea, Ligiidae) from the Southern Appalachians, eastern North America. European Journal of Taxonomy 976: 133-170, DOI: 10.5852/ejt.2025.976.2783, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2783/12721
03C38795FFEFFFE3FDE77CE3FBBEFDDB.taxon	description	Figs 16 – 17	en	Recuero, Ernesto, Caterino, Michael S. (2025): Is there anybody (new) out there? Seven new species of Ligidium (Isopoda, Oniscidea, Ligiidae) from the Southern Appalachians, eastern North America. European Journal of Taxonomy 976: 133-170, DOI: 10.5852/ejt.2025.976.2783, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2783/12721
03C38795FFEFFFE3FDE77CE3FBBEFDDB.taxon	materials_examined	Material examined USA – Georgia • 1 ♀; Rabun Co., Chattahoochee N. F., Rabun Cliffs; 34.9713 ° N, 83.2978 ° W; 1244 m a. s. l.; 11 May 2021; M. Caterino and A. Haberski leg.; CUAC 000138049 • 2 ♂♂; same data as for preceding; CUAC 000180796, CUAC 000180797 • 1 ♀; Rabun Co., Chattahoochee N. F., Rabun Cliffs; 34.9709 ° N, 83.3004 ° W; 1308 m a. s. l.; 25 Nov. 2019; M. Caterino leg.; CUAC 000138078 • 1 ♂; same data as for preceding; CUAC 000171280. – North Carolina • 1 ♀; Avery Co., Grandfather Mountain; 36.0893 ° N, 81.8388 ° W; 1387 m a. s. l.; 21 Apr. 2022; M. Caterino, E. Recuero and A. Haberski leg.; CUAC 000171348 • 2 ♂♂; same data as for preceding; CUAC 000171346, CUAC 000171347 • 1 ♀; Avery Co., Grandfather Mountain, Bridge Trail; 36.0948 ° N, 81.8311 ° W; 1554 m a. s. l.; 21 Apr. 2022; M. Caterino, E. Recuero and A. Haberski leg.; CUAC 000171339 • 1 ♂; same data as for preceding; CUAC 000171340 • 1 ♀; Avery Co., Grandfather Mountain, Grandfather Trail; 36.0978 ° N, 81.8293 ° W; 1637 m a. s. l.; 21 Apr. 2022; M. Caterino, E. Recuero and A. Haberski leg.; CUAC 000171334 • 3 ♂♂; Caldwell Co., Grandfather Mountain, Boone Scout Trail; 36.1164 ° N, 81.7844 ° W; 1259 m a. s. l.; 6 Oct. 2020; M. Caterino, F. Etzler and A. Haberski leg.; CUAC 000171307, CUAC 000171308, CUAC 000138059 • 1 ♀; same data as for preceding; CUAC 000171309 • 1 ♂; Graham Co., Nantahala N. F., Huckleberry Knob; 35.3210 ° N, 83.9934 ° W; 1674 m a. s. l.; 4 May 2020; M. Caterino and F. Etzler leg.; CUAC 000180792 • 1 ♀; Haywood Co., Black Balsam Knob; 35.3289 ° N, 82.8745 ° W; 1839 m a. s. l.; 20 Oct. 2020; M. Caterino and F. Etzler leg.; CUAC 000138045 • 1 ♂; Madison Co.: Pisgah N. F.: Camp Creek Bald; 36.0220 ° N, 82.7167 ° W; 1445 m a. s. l.; 1 Mar. 2022; M. Caterino, E. Recuero and P. Wooden leg.; CUAC 000171352 • 5 ♀♀; same data as for preceding; CUAC 000171353 to CUAC 000171357 1 ♂; Polk Co., North Pacolet River; 35.2221 ° N, 82.3059 ° W; 384 m a. s. l.; 15 Feb. 2022; E. Recuero, M. Caterino, C. Harden and P. Wooden leg.; CUAC 000171344 • 1 ♂; Polk Co., Tryon, Melrose Falls; 35.2199 ° N, 82.2987 ° W; 407 m a. s. l.; 10 Aug. 2021; E. Recuero, M. Caterino, A. Haberski and P. Wooden leg.; CUAC 000171335 • 2 ♀♀; same data as for preceding; CUAC 000171336, CUAC 000171337 2 ♂♂; Rutherford Co., Chimney Rock; 35.4372 ° N, 82.2506 ° W; 383 m. a. s. l.; 3 Apr. 2023; M. Caterino, E. Recuero and C. W. Harden leg.; CUAC 0001169835, CUAC 0001169838 • 3 ♂♂; Yancey Co., Pisgah N. F., Woody Ridge Trail; 35.8455 ° N, 82.2279 ° W; 1259 m a. s. l.; 19 Oct. 2021; M. Caterino, E. Recuero and A. Haberski leg.; CUAC 000180795, CUAC 000171326, CUAC 000171327 • 6 ♀♀; same data as for preceding; CUAC 000171328 to CUAC 000171333. – South Carolina • 1 ♀; Greenville Co., Chestnut Ridge H. P., South Pacolet River; 35.1503 ° N, 82.2804 ° W; 344 m a. s. l.; 15 Feb. 2022; E. Recuero leg.; CUAC 000171350 • 2 ♂♂; Pickens Co., Sassafras Mountain; 35.0647 ° N, 82.7774 ° W; 1029 m a. s. l.; 11 Jun. 2020; F. E. Etzler leg.; CUAC 000180798, CUAC 000180799 • 1 ♀; same data as for preceding; CUAC 000138055. – Tennessee • 1 ♂; Greene Co., Cherokee N. F., Firescald Knob; 36.0337 ° N, 82.7024 ° W; 1353 m a. s. l; 1 Mar. 2022; M. Caterino, E. Recuero and P. Wooden leg.; CUAC 000171351 • 1 ♀; Sevier Co., Great Smoky Mountains N. P., Mount LeConte; 35.6427 ° N, 83.4426 ° W; 1581 m a. s. l.; 28 Sep. 2021; M. Caterino, E. Recuero, A. Haberski and P. Wooden leg.; CUAC 000171276 • 1 ♂; same data as for preceding; CUAC 000171338 • 1 ♀; Sevier Co., Smoky Mountains N. P., Mount LeConte, Alum Cave; 35.6382 ° N, 83.4387 ° W; 1317 m a. s. l.; 28 Sep. 2021; M. Caterino, E. Recuero, A. Haberski and P. Wooden leg.; CUAC 000171325 • 1 ♂; same data as for preceding; CUAC 000171323 • 1 ♀; Sevier Co., Great Smoky Mountains N. P., off Highway 441; 35.6237 ° N, 83.4163 ° W; 1394 m a. s. l.; 12 Mar. 2020; M. Caterino and F. Etzler leg.; CUAC 000171275 • 1 ♂; same data as for preceding; CUAC 000171299 • 2 imm.; Unicoi Co., Cherokee N. F., Big Bald; 35.9938 ° N, 82.4573 ° W; 1596 m a. s. l.; 5 Aug. 2020; M. Caterino, A. Haberski and P. Wooden leg.; CUAC 000171310, CUAC 000171311.	en	Recuero, Ernesto, Caterino, Michael S. (2025): Is there anybody (new) out there? Seven new species of Ligidium (Isopoda, Oniscidea, Ligiidae) from the Southern Appalachians, eastern North America. European Journal of Taxonomy 976: 133-170, DOI: 10.5852/ejt.2025.976.2783, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2783/12721
03C38795FFEFFFE3FDE77CE3FBBEFDDB.taxon	discussion	Remarks This is the most widely distributed Ligidium species in the southern Appalachian Mountains (Fig. 16), presenting also a broad altitudinal range, from 344 to 1674 m a. s. l. Although the species’ distribution spans important biogeographical barriers, such as the Asheville Depression, it includes several deep mitochondrial lineages that seem to be geographically restricted to particular areas delimited by that barrier and one formed by the Little Tennessee River (Figs 1, 18). This pattern is not common among other litter arthropods in this area (Caterino & Recuero 2023; Recuero & Caterino 2024 b, 2024 c). This suggests an old presence and diversification within these mountains. See Remarks under Ligidium pacolet sp. nov. and L. whiteoak sp. nov. and Fig. 17 for diagnostic information.	en	Recuero, Ernesto, Caterino, Michael S. (2025): Is there anybody (new) out there? Seven new species of Ligidium (Isopoda, Oniscidea, Ligiidae) from the Southern Appalachians, eastern North America. European Journal of Taxonomy 976: 133-170, DOI: 10.5852/ejt.2025.976.2783, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2783/12721
03C38795FFEBFFE1FD8B7844FC9AFB98.taxon	description	Figs 19 – 20	en	Recuero, Ernesto, Caterino, Michael S. (2025): Is there anybody (new) out there? Seven new species of Ligidium (Isopoda, Oniscidea, Ligiidae) from the Southern Appalachians, eastern North America. European Journal of Taxonomy 976: 133-170, DOI: 10.5852/ejt.2025.976.2783, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2783/12721
03C38795FFEBFFE1FD8B7844FC9AFB98.taxon	materials_examined	Material examined USA – West Virginia • 1 ♂; Pocahontas Co., Pocahontas Campground; 38.1026 ° N, 79.9666 ° W; 756 m a. s. l.; 13 Oct. 2022; M .. Caterino and E. Recuero leg.; CUAC 000174439 • 1 ♀; Randolph Co., Monongahela N. F.; 38.5595 ° N, 79.9262 ° W; 1135 m a. s. l.; 12 Oct. 2022; M. Caterino and E. Recuero leg.; CUAC 000174441 • 2 ♂♂; same data as for preceding; CUAC 000174440, CUAC 000174442 • 1 imm.; same data as for preceding; CUAC 000174443.	en	Recuero, Ernesto, Caterino, Michael S. (2025): Is there anybody (new) out there? Seven new species of Ligidium (Isopoda, Oniscidea, Ligiidae) from the Southern Appalachians, eastern North America. European Journal of Taxonomy 976: 133-170, DOI: 10.5852/ejt.2025.976.2783, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2783/12721
03C38795FFEBFFE1FD8B7844FC9AFB98.taxon	discussion	Remarks The studied specimens from West Virginia have been collected in leaf litter from deciduous and conifer forests at mid-elevations (Fig. 18). They differ slightly from the ones illustrated from North Carolina (Schultz 1970, 1982). Particularly, males from North Carolina present ten scale-like spines on the distal margin of the tip of male pleopod 2 endopodite, while those from West Virginia present four (Fig. 19 D – E). Also, the illustration of a male pleopod 2 exopodite from North Carolina shows a large spiniform or plumose seta on the inner corner of the caudal margin, not present in the samples from West Virginia (Fig. 19 C). These differences could indicate different species, but we refrain from proposing any more new names for this complex, until the true status of Ligidium elrodii and L. longicaudatum may be clarified. This species is not closely related to any of the other taxa included in our analyses (Fig. 1 A). Four subspecies of Ligidium elrodii have been described from caves in the states of Virginia, Tennessee and Georgia, based on differences in the male pleopod 2 endopodite (Schultz 1970). One of them, L. e. leensis, from Bowling Cave, Lee Co., Virginia (see map in Fig. 18), is relatively similar to our specimens and to those illustrated from North Carolina, differing in a more markedly squarish process on the inner margin of male pleopod 2 endopodite, with only two scale-like spines on its distal margin. Indeed, it could represent a different species, but until L. elrodii is revised we prefer to maintain its status. The other three have more pronounced differences in this particular structure, which are indicative of specific status. We propose to raise them to full species.	en	Recuero, Ernesto, Caterino, Michael S. (2025): Is there anybody (new) out there? Seven new species of Ligidium (Isopoda, Oniscidea, Ligiidae) from the Southern Appalachians, eastern North America. European Journal of Taxonomy 976: 133-170, DOI: 10.5852/ejt.2025.976.2783, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2783/12721
03C38795FFE9FFE1FDE87E0BFD1BFAF4.taxon	discussion	Remarks In the form of male pleopod 2 endopodite this species resembles Ligidium elrodii, but having a constricted tip, and only five scale-like spines on the distal margin of the tip. The species is so far known only from its type locality, Blowing Spring Cave, 2.5 miles NE Cloudland, Chattooga Co., Georgia (see map in Fig. 18).	en	Recuero, Ernesto, Caterino, Michael S. (2025): Is there anybody (new) out there? Seven new species of Ligidium (Isopoda, Oniscidea, Ligiidae) from the Southern Appalachians, eastern North America. European Journal of Taxonomy 976: 133-170, DOI: 10.5852/ejt.2025.976.2783, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2783/12721
03C38795FFE9FFE1FDE47FBFFE66F940.taxon	discussion	Remarks In the form of male pleopod 2 endopodite this species resembles Ligidium nantahala sp. nov., but its tip has a much shorter process and a squared section. The species is so far known only from its type locality, Cantwell Valley Cave, Hancock Co., Tennessee (see map in Fig. 18).	en	Recuero, Ernesto, Caterino, Michael S. (2025): Is there anybody (new) out there? Seven new species of Ligidium (Isopoda, Oniscidea, Ligiidae) from the Southern Appalachians, eastern North America. European Journal of Taxonomy 976: 133-170, DOI: 10.5852/ejt.2025.976.2783, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2783/12721
03C38795FFE9FFDEFD8D7CD2FE96FEDE.taxon	discussion	Remarks In the form of male pleopod 2 endopodite this species resembles L. gadalutsi sp. nov., but the tip is shorter and blunt, not acuminate. They also differ in the shape of male pleopod 1 endopodite. The species is so far known only from its type locality, Coley Cave # 2, in Virginia, Scott Co. (see map in Fig. 18).	en	Recuero, Ernesto, Caterino, Michael S. (2025): Is there anybody (new) out there? Seven new species of Ligidium (Isopoda, Oniscidea, Ligiidae) from the Southern Appalachians, eastern North America. European Journal of Taxonomy 976: 133-170, DOI: 10.5852/ejt.2025.976.2783, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2783/12721
