identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03C587C9FF81FF8CFEBA7D2A3954FA48.text	03C587C9FF81FF8CFEBA7D2A3954FA48.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Berkanella Taybi & Glöer & Mabrouki 2024	<div><p>Berkanella gen.n.</p><p>Figs 2–4.</p><p>TYPE SPECIES:  Islamia tifertiensis Glöer et al., 2020 (see below)  .</p><p>DIAGNOSIS: The new genus is characterised by the following characters: the minute valvatiform transparent shell (when fresh), the colourless body and atrophy of eye spots (adapted to a subterranean or stygobiotic mode of life), the penis has a knob at the left side near the basis, at the distal end tapered.</p><p>DIFFERENTIAL DIAGNOSIS:  Berkanella gen.n. may be confused with the only three genera of stygobionts and valvatiformes known to occur in North Africa, namely:  Rifia Ghamizi, 2020, which has a bifurcated penis; the recently described Maroccohoratia Ghamizi et Falniowski, 2024 which has a proportionally minute penis, bent and triangular in shape, with a broad and slightly marked swelling distally on its right side, and finally Maroccoarganiella Ghamizi et Falniowski, 2024 which has a broad and straight penis flat and massive, with a broad and not prominent outgrowth distally on its right side. The “outgrowths” in the genera of Ghamizi et al. (2024) should be better named as “swelling”.  Berkanella gen.n. is the only stygobiont and valvatiform hydrobiid having a penis with a knob at the left side near the basis and the tapered distal end. While the other stygobiont genera i.e.  Heideella Backhuys et Boeters, 1974;  Atebbania Ghamizi et al., 1999 and therecentlydescribed  Mahrazia Mabrouki et al.,2022 have elongated conical shells.The penis morphology of  Berkanella gen.n. is a unique characteristic for the stygobiont and valvatiform microsnails, not known among any other hydrobiid genera.This feature justifies the description of a new genus, as it is acknowl- edged that hydrobiid genera can be identified by the shell shape in combination with the penis morphology (Radoman, 1983; Szarowska,2006; Glöer,2022).The other valvatiform genera of NorthAfrica are crenobi - ont gastropods living in surface waters, have eyespots and pigmented bodies. The new genus resembles the recently described crenobiont  Ainiella Taybi, Glöer et Mabrouki 2022 . However, the latter has larger shells, pigmented bodies and the largest eyespots among the valvatiform hydrobiid microsnails of North Africa. Whilst  Berkanella gen.n. is an obligate stygobiont.</p><p>DERIVATIO NOMINIS. The new genus was named after Berkane province, where it was first found the first time.</p><p>The new genus contains two species, one of them is described below as new for science.</p></div>	https://treatment.plazi.org/id/03C587C9FF81FF8CFEBA7D2A3954FA48	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Taybi, A. F.;Glöer, P.;Mabrouki, Y.	Taybi, A. F., Glöer, P., Mabrouki, Y. (2024): Description of Berkanella selouensis gen. et sp. n., a new subterranean snail from Morocco and redescription of Islamia tifertiensis Glöer, Mabrouki et Taybi, 2020 (Gastropoda: Hydrobiidae). Invertebrate Zoology 21 (2): 147-156, DOI: 10.15298/invertzool.21.2.03, URL: https://doi.org/10.15298/invertzool.21.2.03
03C587C9FF80FF8AFF3A7E0D3AC1F91A.text	03C587C9FF80FF8AFF3A7E0D3AC1F91A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Berkanella tifertiensis (Gloer, Mabrouki et Taybi 2020) Taybi & Glöer & Mabrouki 2024	<div><p>Berkanella tifertiensis (Glöer et al., 2020) comb.n.</p><p>Islamia tifertiensis Glöer, Mabrouki et Taybi, 2020: Glöer et al., 2020: 3–4; fig. 2.</p><p>MATERIAL EXAMINED. Holotype. Empty shell deposited in the collection of the Zoological Museum Hamburg (ZMH 140648). Paratypes.  Three empty shells from the type locality deposited in the collection of the  Zoological Museum Hamburg (ZMH 140649) and three ethanol-fixed specimens kept in the personal collection of Glöer and 10 ethanol-fixed specimens in the personal collection of Mabrouki and Taybi (Figs 2–4)  .</p><p>TYPE LOCALITY.  Tiffert spring, Berkane prov- ince (the oriental region of Morocco), adjacent to the Moulouya River (2′16.800″N, 2°25′36.000″W), located inside the   SBEI ( Site of Biological and  Ecological Interest) of the  Moulouya, listed also as a Ramsar site. The live specimens (topotypes) were collected on 20 March, 2023  .</p><p>DESCRIPTION:</p><p>CONCHOLOGICAL FEATURES. The translucent and colourless shell has 3 fast-growing glossy whorls with a slightly striated surface (Fig. 2A–D). Shell surface smooth,with growth lines that are slightly visible. The small spire is slightly raised up; the body whorl is prominent.The aperture is rather rhomboidal (or almost quadrangular) and prosocline (leaning forwards relative to the direction of growth), the peristome is situated at the bottom in umbilical view. The shell is 0.92–0.96 mm high and 1.31–1.42 mm broad. Shell measurements are presented in Table 1.</p><p>OPERCULUM. The corneous operculum is smooth and very thin, pale yellow.</p><p>SOFT PARTS. The species is eyeless and colourless, body translucent when alive (Fig. 4), white when preserved, pigment-less; cephalic tentacles very long and thin (no pallial tentacle), without eyes. The penis has a knob at the left side near the basis and the distal part is tapered (Fig. 2E).</p><p>REMARKS. This species has originally been places in the genus  Islamia because we had only empty shells and were unable to study its anatomy (Glöer et al., 2020). The  Islamia species of Morocco were described only based on shell criteria and assigned provisionally to this genus without knowledge of its anatomy, pending the discovery of males. In  Islamia, the penis is bilobed and the bursa copulatrix is absent.</p><p>GEOGRAPHIC RANGE. This new genus and species is known only from eastern Morocco (Fig. 1). It’s type locality, which is a spring located in the plain of Triffa, is bounded to the east by Oued Kiss (Algeria), to the north by the Mediterranean Sea, to the west by Oued Moulouya and to the south by the Beni-Snassen ranges. It covers an area of 61,000 ha and is one of the most fertile and productive areas in eastern Morocco (Yahya et al., 2017). The plain of Triffa (Eastern Morocco) has great socio-economic importance in the region; the local economy is based specially on agriculture, which consumes the total water use. Indeed, the plain is characterized by two aquifers:afreeoneofsecondaryandquaternaryforma- tions; and a confined aquifer of the Liassic formation (El Mandour et al., 1998).</p></div>	https://treatment.plazi.org/id/03C587C9FF80FF8AFF3A7E0D3AC1F91A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Taybi, A. F.;Glöer, P.;Mabrouki, Y.	Taybi, A. F., Glöer, P., Mabrouki, Y. (2024): Description of Berkanella selouensis gen. et sp. n., a new subterranean snail from Morocco and redescription of Islamia tifertiensis Glöer, Mabrouki et Taybi, 2020 (Gastropoda: Hydrobiidae). Invertebrate Zoology 21 (2): 147-156, DOI: 10.15298/invertzool.21.2.03, URL: https://doi.org/10.15298/invertzool.21.2.03
03C587C9FF86FF88FC847C7E381BFC92.text	03C587C9FF86FF88FC847C7E381BFC92.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Berkanella selouensis Taybi & Glöer & Mabrouki 2024	<div><p>Berkanella selouensis sp.n.</p><p>Figs 5–6.</p><p>MATERIAL EXAMINED. Holotype. Ethanol-fixed specimen (Fig. 5A, B, D), shell is 1.42 mm high and 1.31 mm wide, deposited in the collection of the Zoological Museum Hamburg (ZMH 141508). Paratypes. Two paratypes deposited in the collection of the  Zoological Museum Hamburg (ZMH 141509), ethanol-fixed (Figs. 5C, 6). In addition, 10 ethanol-fixed specimens are kept in the personal collections of Mabrouki and Taybi  .</p><p>TYPELOCALITY. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-2.911778&amp;materialsCitation.latitude=35.06561" title="Search Plazi for locations around (long -2.911778/lat 35.06561)">AwelllocatedinsidetheMultidisciplinary Faculty of Nador</a>, Selouan, Nador province, the oriental region of Morocco (35°03′56.2″N 2°54′42.4″W) belongstothewatershedoftheMarchica lagoon, the latter classified as SBEI (Site of Biologi- cal and Ecological Interest) and a Ramsar site. The groundwater of the region belongs to the aquifer of Garet. Collected on 20 March, 2023  .</p><p>DESCRIPTION:</p><p>CONCHOLOGICAL FEATURES. The translucent, glossy and colourless shell has 3 fast-growing glossy whorls with a smooth surface,nearly planispiral and thin-walled. Shell growth lines are hardly visible (Fig. 5A–D). The small spire is slightly raised up; the body whorl is prominent. The aperture is prosocline and rounded, the peristome is not sinuated at the bottom from umbilical view. The umbilicus wide, with the earlier whorls visible inside. The shell is 0.63 mm high and 1.35 mm broad. Shell measurements are presented in Table 2.</p><p>OPERCULUM.The very thin corneous operculum is pale yellow.</p><p>SOFT PARTS. The species is eyeless and colourless (Fig. 6), cephalic tentacles very long and thin (no pallial tentacle); the penis has a large knob at the left side near the basis and the distal part is tapered (Fig. 5E).</p><p>DIFFERENTIATING CHARACTERS.The new species differs from  Berkanella tifertiensis (Glöer et al., 2020) comb.n. by a lower shell, almost planispiral, the smooth shell surface, the non-sinuated peristome and the rounded aperture. In addition, the penis in  Berkanella selouensis sp.n. is larger, widened at the basis and tapered at the distal end with a long pointed penis tip, and the knob is larger too.</p><p>DERIVATIO NOMINIS. The specific epithet refers to the city of Selouane, where the new species was found. It is located in the northeast, in the province of Nador, in the eastern Rif and the oriental region of Morocco.</p><p>GEOGRAPHIC RANGE. This new species is known only from the type locality and groundwater from a well located in north-eastern Morocco (Fig. 1).  The Garet-Bouareg aquifer consists of a free aquifer, which extends continuously and generally under the two plains of  Garet and Bouareg. This aquifer extends over an area of 570 km 2 (ADS, 2005). The overall quality of waters in this aquifer is poor due to the high mineralization of the waters (high salinity) and the high levels of nitrates. This is due to the geological nature of the formations from this aquifer and intensive agricultural use  .</p><p>NOMENCLATURAL ACTS: This work and the nomenclatural acts it contains have been registered in ZooBank. The ZooBank Life Science Identifiers (LSID) for this publication are:</p><p>http://zoobank.org/ urn:lsid:zoobank. org:act: E4690E62-79DE-4F94-9DF6- 17E28787A558</p><p>http://zoobank.org/ urn:lsid:zoobank. org:act: 3FA0FF29-3C01-495E-9009- 11B761D8B2B7</p></div>	https://treatment.plazi.org/id/03C587C9FF86FF88FC847C7E381BFC92	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Taybi, A. F.;Glöer, P.;Mabrouki, Y.	Taybi, A. F., Glöer, P., Mabrouki, Y. (2024): Description of Berkanella selouensis gen. et sp. n., a new subterranean snail from Morocco and redescription of Islamia tifertiensis Glöer, Mabrouki et Taybi, 2020 (Gastropoda: Hydrobiidae). Invertebrate Zoology 21 (2): 147-156, DOI: 10.15298/invertzool.21.2.03, URL: https://doi.org/10.15298/invertzool.21.2.03
