Gondwanodromia mikae, Sinclair & Saigusa, 2022

Sinclair, Bradley J. & Saigusa, Toyohei, 2022, A New Aquatic Associated Genus of Trichopezinae from the Southern Hemisphere (Diptera: Empidoidea: Brachystomatidae), Records of the Australian Museum (Rec. Aust. Mus.) 74 (3), pp. 75-98 : 84-87

publication ID

https://doi.org/ 10.3853/j.2201-4349.74.2022.1797

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scientific name

Gondwanodromia mikae

sp. nov.

Gondwanodromia mikae sp. nov.


Figs 16 View Figures 13–16 , 19–33 View Figures 19–23 View Figures 24–29 View Figures 30–33 , 52 View Figures 50–52

Holotype ♂, labelled: “Arr.[oyo] Casa de Piedra [ca 41°08'05"S 71°27'12"W]/ 15kmW of Bariloche / Rio Negro, Argentina / 16.xi.2004 / T. SAIGUSA col.”; “ HOLOTYPE / Gondwanodromia / mikae/ Sinclair & Saigusa” ( KUMF) GoogleMaps . Paratypes: ARGENTINA. Río Negro: 1 ♀, Arroyo Torrontegui [ca 41°15'12"S 71°27'14"W], Lago Gutiérrez, Bariloche , 18.xi.2004, T GoogleMaps . Saigusa ( KUMF) ; 1 ♀, same data except, M. Sugimoto ( KUMF) . Neuquen: 1 ♂, Arroyo de la Estacada [ca 40°50'06"S 71°32'12"W], Lago Nahuel Huapi , 17.xi.2004, T GoogleMaps . Saigusa ( KUMF) .

Possible additional material. CHILE. Region X (Los Lagos): 3 ♀, PN Puyehue, Rio Anticura above Salto Rio Anticura , 6.xii.2013, 40°40.268'S 72°10.215'W, 395 m, G. R GoogleMaps . Curler, ex. riparian vegetation ( CNC).

Diagnosis. The species is readily distinguished by bare eyes, male midlegs not modified, epandrium not visibly divided, with apical surstylus and hypandrium with narrow, upright apical half forming brace for phallus.

Description. Wing length 2.7–3.0 mm. Male. Head ( Fig. 30 View Figures 30–33 ). Eye bare, without ommatrichia. Frons brown. Face with bluish pruinescence; narrower than width of antennal sockets. Ocellar setae long and divergent, inserted anterior to lateral ocelli; pair of shorter anterior and posterior setae; 3 vertical setae with outer seta arched over margin of eye, shorter than ocellar seta; occipital setae shorter and thinner than vertical setae. Antenna ( Figs 21 View Figures 19–23 , 30 View Figures 30–33 ) with scape slightly longer than pedicel; postpedicel less than 1.5× height of eye. Proboscis subequal in length to head height; palpus pale brown, one-third length of proboscis.

Thorax. Dark brown, with thin blue pruinescence laterally on scutum and pleura; faintly bivittate between rows of acrostichals and dorsocentrals. Antepronotum with several pairs of short, strong setae. Acrostichal setae with pair of strong anterior (scapular) setae, uniserial posteriorly with setae alternating in direction, more than half length of dorsocentral setae; 8–10 uniserial dorsocentral setae, anterior seta longer, set off laterally from row; 1 long postpronotal seta and 1–2 shorter anterior setae; 1 presutural supra-alar seta; 3 notopleural setae, middle seta stronger and twice length of other setae; 3–4 postsutural supra-alar setae; 1 postalar seta; 2 pairs of scutellar setae, outer pair about half length of inner pair. Laterotergite with several short setae. Wing ( Fig. 20 View Figures 19–23 ). Infuscate; basal costal seta long and slender. Halter brown. Legs. Coxae pale brown with bluish pruinescence, apical setae brown, sometimes apex of coxae and trochanters yellowish brown; remaining leg segments increasingly darker apically. Femora short and slightly swollen; fore femur with ventral seta on proximal third, nearly as long as width of femur; mid femur with row of short posteroventral setae; hind femur with short, erect, strong dorsal setae near base. Fore tibia without apical spine-like setae. Mid tibia slender with row of short anteroventral and posteroventral setae; without pair of peg-like apicoventral setae. Hind tibia with strong ventral and dorsal setae, shorter than width of tibia. Mid tarsomere 1 without row of strong setae; without erect posteroventral seta at basal fifth.

Abdomen. Tergite 6 without dorsal setae longer than on other tergites. Sternite 7 not prolonged posterolaterally into narrow lobe. Tergite 8 trapezoidal, half-length of sternite 8; posterior margin wider than anterior margin, bearing numerous setae, denser on lateral margin; sternite 8 trapezoidal, with setae longer than anterior sternites. Terminalia ( Figs 24–27 View Figures 24–29 , 31–33 View Figures 30–33 ). Cercus thinly sclerotized narrow, rectangular with truncate apex, bearing long setae along inner margin and apex. Epandrium triangular, tapered in lateral view with short oblong lobe along posteromedial margin; without dorsal bridge. Surstylus at apex of epandrium, without distinct separation; triangular; inner margin with short, pointed lobe and pair of long medially directed setae mounted on short tubercles, lower seta nearly touching opposite pair. Hypandrium with broad base and narrow apical half bracing phallus; gonocoxal apodeme broad, extending beyond anterior margin of hypandrium. Phallus narrow, tubular, upright beyond hypandrium, with expanded apex; ejaculatory apodeme nearly as broad as gonocoxal apodeme, articulated at base of phallus.

Female. Similar to male except as follows: scape subequal in length to pedicel; postpedicel subequal in length to eye height. Terminalia ( Figs 28, 29 View Figures 24–29 ). Tergite 8 with short dorsomedial apodeme on anterior margin. Syntergite 9+10 (acanthophorites) with 1 row of strong, erect setae, apical setae longer and strong setae scattered dorsally. Spermatheca spherical ( Fig. 29 View Figures 24–29 ).

Etymology. This species is named after Mika Sugimoto, who cooperated with Saigusa during field work of torrenticolous Empididae in northern Patagonia.

Distribution. This species is currently confirmed from several cascading streams around lakes Gutiérrez and Nahuel Huapi ( Figs 1, 2 View Figures 1–4 ) in Argentina ( Fig. 52 View Figures 50–52 ).

Remarks. Three female specimens listed above from Chile are possibly conspecific, but males from this region (G. sp., Fig. 52 View Figures 50–52 ) will need to be collected to confirm their identification.


Tavera, Department of Geology and Geophysics


Kasetsart University Museum of Fisheries


Departamento de Geologia, Universidad de Chile


Canadian National Collection of Insects, Arachnids, and Nematodes













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