Ropalidia impetuosa, (Smith)

Carpenter, James M. & Brown, Graham R., 2022, Catalogue of the Australian social wasps (Hymenoptera: Vespidae), Zootaxa 5214 (4), pp. 451-495 : 462-463

impetuosa (Smith)

Icaria impetuosa Smith, 1860 [1861]: 131, female - “Batchian; Amboyna” (lectotype Oxford); 1863 [1864]a: 41 (distr.); 1863 [1864]b: 138 (distr.); 1871: 380 (cat.). - Dalla Torre, 1894: 118 (cat.); 1904: 73 (cat.). - van der Vecht, 1941: 104 (syn. of R. gregaria (de Saussure)) . - Kojima, 2001a: 9 (syns.: I. spilocephala Cameron , I cohni du Buysson , R. gregaria inquieta Cheesman , R. gregaria var. tolerans Cheesman , R. gregaria auctt.).

Icaria spilocephala Cameron, 1906b: 230 , female - [New Guinea:] “Etna Bay” (lectotype Naturalis). - van der Vecht, 1941: 104 (syn. of R. gregaria (de Saussure)) . - Kojima & Carpenter, 1997a: 5 (designation of lectotype). - Kojima & van Achterberg, 1997a: 166 (list). - Kojima, 1999b: 367, 371 (syn. of gregaria (de Saussure)) ; 2001a: 9 (syn. of R. impetuosa (Smith)) .

Icaria Cohni du Buysson, 1909: 306 , male - “ Bougainville ” (lectotype Paris). - van der Vecht, 1941: 104 (syn. of R. gregaria (de Saussure)) . - Richards, 1978: 86 ( kohni [!]; syn. of R. gregaria spilocephala (Cameron)) . - Kojima, 2001a: 5 (designation of lectotype), 9 (syn. of R. impetuosa (Smith)) .

Icaria picta ; du Buysson, 1913 [1911]: 229 (distr.).

Ropalidia gregaria ; Tillyard, 1925: 297. - Williams, 1928: 79 (prey: Perkinsiella saccharicida Kirkaldy ). - van der Vecht, 1941: 104 (syns.: Icaria cohni du Buysson , I. impetuosa Smith , I. spilocephala Cameron ), 111 (key), 149 (distr. [including subspecies spilocephala ]). - Cheesman, 1952: 3, 20 (male). - van der Vecht, 1962: 27 (distr.; nest). - Baltazar, 1966: 291 (in subgenus Anthreneida ; cat.). - Iwata, 1976: 295 (in subgenus Anthreneida ; ethology). - Richards, 1978: 50, 53, 61 (key), 65 (larva in key), 83, Fig. 26 (in subgenus Icariola ; redescription; nest; ethology). - Carpenter, 1981 [1982]: 34 (phylogenetic relationships). - Kojima, 1982a: 272 (in subgenus Icariola ; nest); 1982b: 17 (ethology); 1984a: 52, Fig. 2 (larva); 1984b: 525 (key), 528, Figs 6, 34, 39, 44, 49 (in subgenus Icariola ; male genitalia; Philippines). - Spradbery, 1991b: 349 (colony population). - Wenzel, 1991: 487 (nest). - Itô, 1993: 87 (in subgenus Icariola ; ethology). - Naumann, 1993, in Naumann et al.: 180 (distr.). - Macalintal & Starr: tables 1–2 (in subgenus Icariola ; morphology). - Wenzel, 1998: 26 (nest examined). - Kojima, 1999b: 367, 371 (syn.: I. spilocephala (Cameron) , erroneously marked “Syn. nov.”; W. A., N. T., Qld., N. S. W., Vic.); 1999c: 381 (key), 386 (distr.), Figs 17, 21, 25, 36.

Ropalidia impetuosa ; Bequaert, 1932: 51. - Richards, 1978: 58 (in subgenus Icariola ; list), 86 (probable syn.: Ropalidia gregaria spilocephala (Cameron)) . - Kojima, 2001a: 9 (syns.: I. spilocephala , I. cohni , R. gregaria inquieta , R. gregaria var. tolerans ); 2001b: 32, 56 (syn.; distr.). - Saito & Kojima, 2005a: 273–274 (not established south of latitude 30°S). - Nugroho et al., 2012 [2011]: 152 (list).

Ropalidia cohni ; Bequaert, 1932: 51.

? Ropalidia cyathiformis ; Bequaert, 1932: 51 (recorded from New Guinea; syns.: variegata Smith , picta de Saussure ) [misidentification?].

Ropalidia spilocephala ; Bequaert, 1932: 51.

Ropalidia gregaria inquieta Cheesman, 1952: 3 , 20, female, male - “Papua,...Mafulu, 4000 ft. ” (lectotype male London); also from three other localities in New Guinea. - Richards, 1978: 86 (designation of lectotype [error in locality: Mafalu]; syn. of R. gregaria spilocephala ). - Kojima, 2001a: 9 (syn. of R. impetuosa (Smith)) .

Ropalidia gregaria var. tolerans Cheesman, 1952 ): 3 (as subspecies), 21 (as var.), female - “Cyclops Mts. Sabron” (lectotype London). - Richards, 1978: 86 (designation of lectotype; syn. of R. gregaria spilocephala ). - Kojima, 2001a: 9 (syn. of R. impetuosa (Smith)) .

Ropalidia gregaria gregaria ; Richards, 1978: 53 (nest), 62 (key), 86 (in subgenus Icariola ; “two [subspecies] can be recognized in the Australia-New Guinea region. The forms of Malaya will require separate study.”). - Cardale, 1985: 214 (in subgenus Icariola ; cat.). - Itô, 1986b: 313 (ethology). - Kojima, 1987a: 145, Figs 52–58 (larva). - Spradbery, 1991b: 351 (colony population). - Itô & Yamane, 1992: 63–74 (ethology). - Itô, 1993: 27, 46, 66, 68 (in subgenus Icariola ; ethology). - Borsato, 1994 [1993],: 250 (in subgenus Icariola ; N. T.). - Kojima & Carpenter, 1997a: 27 (cat.). - Brown, 2009: 30 (North Coast).

Ropalidia gregaria spilocephala ; Richards, 1978: 53, 62 (key), 86 (syns.: Icaria kohni [!] du Buysson, Ropalidia gregaria inquieta Cheesman , Ropalidia gregaria var. tolerans Cheesman ; probably syn. of Icaria impetuosa Smith ; “I examined the holotype male [!] in the Zoological Museum, Amsterdam.”; redescription; nest). - Cardale, 1985: 215 (in subgenus Icariola ; cat.). - Spradbery & Kojima, 1989: 636 (nest; colony population). - Spradbery, 1991b: 351 (colony population). - Borsato, 1994 [1993]: 250 (in subgenus Icariola ; Qld.). - Itô et al., 1996: 231 (ethology). - Kojima & Carpenter, 1997a: 27 (cat.).

Distribution: NE coastal, Murray-Darling basin; N. T., Qld., N. S. W.; also New Guinea; New Ireland; Solomon Is.; Fiji; Sulawesi; Ceram; Bacan; Ambon; Philippines.