Cloeodes incus (Waltz & McCafferty) Waltz & McCafferty, 2008

Nieto, Carolina & Richard, Barton, 2008, The genus Cloeodes (Ephemeroptera: Baetidae) in Argentina with new generic synonymy and new species, Zootaxa 1727, pp. 1-21 : 11

Cloeodes incus (Waltz & McCafferty) n. comb.

( Figs. 31–42 )

Bernerius incus Waltz & McCafferty, 1987a: 181 . Baetis (?) sp. B. Berner, 1980: 190.

Diagnosis. Cloeodes incus n. comb., known from nymphs, can be distinguished from the other species of the genus by the following combination of characters: 1) labrum ( Fig. 31 ) with one subapical seta centrally and two setae near lateral margin, anterior margin with basally bifid setae near midline and apically bifid setae near lateral margin; 2) mandibles without setae between prostheca and mola; 3) left mandible with thumb of molar area in the same plane as anterior margin ( Fig. 32 ); 4) right mandible with incisors cleft in two sets ( Fig. 33 ); 5) lingua with a rounded projection ( Fig. 34 ); 6) maxillary palpi long, 1.5 times the length of galealacinia, two segmented, segment II with a constriction ( Fig. 35 ); 7) labium with segment III of palpi rounded ( Fig. 36 ); 8) hind wing pads present; 9) dorsal edge of femora ( Fig. 37 ) with a row of pointed spines and apically rounded (as in Fig. 63 ), femora, tibiae and tarsi with bipectinate spines; 10) tarsal claws with minute denticles basally ( Fig. 38 ); 11) posterior margins of abdominal terga with spines ( Fig. 39 ), sterna with long fine setae randomly distributed throughout; 12) gills pointed apically ( Fig. 40 ); 13) posterior margin of paraprocts with spines apically ( Fig. 41 ); 14) caudal filaments with simple setae, and with whorl of spines on each segment ( Fig. 42 ).

Material examined. Paratypes: 2 nymphs: BOLIVIA, trib. Río Umalo o Grande, below Calamarca on La Paz, Oruro Rd, 18/ VII/ 1977, S. S. Roback & L. Berner colls. Material housed in FAMU.

Figures

FIGURES 31 – 42. Cloeodes incus n. comb. Nymph. Mouthparts (Figs. 31 – 36): 31, labrum d. v., left d. v., right v. v.; 32, left mandible v. v.; 33, right mandible v. v.; 34, hypopharynx v. v.; 35, maxilla v. v.; 36, labium, left d. v., right v. v. 37, leg I; 38, tarsal claw I. 39, posterior margin of tergum IV. 40, gill IV. 41, paraproct. 42, terminal filament, left cercus, right caudal filament.

FIGURES 55 – 67. Cloeodes penai. Nymph. 55, antennae. Mouthparts (Figs. 56 – 61): 56, labrum, left d. v., right v. v.; 57, left mandible v. v.; 58, right mandible v. v.; 59, hypopharynx v. v.; 60, maxilla v. v.; 61, labium, left d. v., right v. v. 62, leg I; 63, femur I apical margin; 64, tarsal claw I. 65, posterior margin of tergum IV. 66, gill IV. 67, paraproct.