Morbora DISTANT 1899
publication ID |
https://doi.org/ 10.5281/zenodo.12996779 |
DOI |
https://doi.org/10.5281/zenodo.13715146 |
persistent identifier |
https://treatment.plazi.org/id/0380878F-FF91-FFAD-FDAB-FDB3C332FBBD |
treatment provided by |
Luisschmitz |
scientific name |
Morbora DISTANT 1899 |
status |
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Morbora DISTANT 1899 ( Figs 2d, e View Fig , 11 View Fig , 12 View Fig , 13, 14, Table 5 View Table 5 )
Morbora DISTANT 1899: 47 (gen. nov.); DISTANT 1899: 47 (description); BERGROTH 1904: 355 (note); SCHOUTEDEN 1904: 86 (description); KIRKALDY 1909: 265 (catalogue); GROSS 1975: 88 (description); MCDONALD & CASSIS 1984: 538, 547 (key; revision); CASSIS & GROSS 2002: 586 (catalogue)
Type species: Morbora australis DISTANT 1899 monotypy
Diagnosis: Morbora is recognised by the following characters: body oval; head semicircular, with dentate margins ( Figs 11 View Fig a-c, 12a); body with fanlike (Figs 13c, h, i) or clove-like (Figs 13a, b, f) setae intermixed with curly sericeous setae; pronotal, corial and connexival margins dentate ( Fig. 11 View Fig ); male androconial glands present on abdominal sterna IV-VI ( Figs 2d, e View Fig ); ejaculatory reservoir small, oval ( Figs 14c, d View Fig ); ventral conducting canal straight; CAI membraneous, medially fused ( Figs 14c, d View Fig ); CAII strongly sclerotized, U-shaped ( Figs 14c, d View Fig ); CAIII present or absent ( Figs 14c, d View Fig ); vesica, elongate, arcuate or S-shaped ( Figs 14c, d View Fig ); female gonocoxae I tripartite; interlocking rami absent; spermathecal fecundation canal short; spermathecal reservoir elliptoid; and spermathecal bulb bilobed.
Description: Body oval; small species, males 4.51-5.37 mm, females 4.41-5.88 mm; dorsum moderately convex; cryptozoic colouring, yellow-brown to dark brown, with patterned darker markings; dorsum with elongate clove-shaped (Figs 13a, b, f) to fanlike setae (Figs 13c, h, i), intermixed with curly sericeous setae (Figs 13a, e, h); densely punctate. Head: semicircular, strongly deflexed ( Figs 12a, b View Fig ); lateral mar-gins of jugae explanate, strongly denticulate, surpassing apex of clypeus ( Figs 12a, b View Fig ); bucculae moderately tumescent, medial margins subparallel ( Fig. 12c View Fig ); eyes small ( Figs 12 View Fig a-c); antennae: AI elongate, either longest subequal or subequal to AIV; labium: reaching metasternum; either LI or LII longest segment, LIII & LIV small, either subequal or LIII longest. Pronotum: subtrapezoidal, moderately convex in profile ( Figs 11 View Fig a-c); anterior margin excavate; with broad, explanate paranotal lobes, lateral margins convex, explanate, denticulate, anterolateral angles anteriorly produced, extending beyond posterior margins of eyes ( Figs 11 View Fig a-c, 12a-c); posterior margins rounded to truncate ( Figs 11 View Fig a-c). Scutellum: broad, U-shaped to subhemispherical ( Figs 11 View Fig a-c), strongly convex in profile; dentate corial and abdominal connexival margins visible ( Figs 11 View Fig a-c). Thoracic sterna: prosternal margins and anterior margin of proepisternum jointly raised, moderately explanate ( Fig. 12c View Fig ). Thoracic pleura: external efferent system of metathoracic glands moderately developed ( Fig. 12c, d View Fig ); ostiole small; evaporative area of metathoracic gland extending onto posterior portion of mesopleuron and beyond peritreme; peritreme short, tongue-like, raised, and wrinkled ( Figs 12c, d View Fig ). Pregenital abdomen: broad, strongly convex; lateral margins spinose; male abdominal SIV-VI with paired, suboval regions of androconial glands ( Figs 2 View Fig d-f). Male Genitalia: ventral surface of pygophore terminal in orientation ( Fig. 12e View Fig ), ventral margin entire ( Fig. 14a View Fig ); genital opening small to moderately sized ( Figs 12f View Fig , 14a View Fig ); parameres short, crown small, hook-like, tip blunt ( Fig. 14b View Fig ); aedeagus broad; phallotheca conical, without processes, weakly sclerotized ( Figs 14c, d View Fig ); ductus seminis proximalis thin, membraneous ( Figs 14c, d View Fig ); ejaculatory apparatus simple ( Figs 14c, d View Fig ), with single, straight, ventral conducting canal, without convolutions; ejaculatory reservoir short, suboval; dorsal conducting canal simple, not expanded; vesica broad, elongate, weakly S-shaped ( Figs 14c, d View Fig ); CAI large, membraneous, medially fused, distally divided, without sclerotization ( Figs 14c, d View Fig ); CAII large, strongly sclerotized, U-shaped, with subdistal spur ( Figs 14c, d View Fig ); CAIII membraneous, medially fused, distally free, without sclerotization ( Figs 14c, d View Fig ). Female Terminalia: female paratergites IX Sshaped; female gonocoxae I divided, tripartite; interlocking rami absent. Spermatheca: fecundation canal short; spermathecal reservoir weakly dilated, simple; and spermathecal bulb bilobed, with proximal and distal flanges.
Diversity and distribution: Morbora is endemic to continental Australia, comprised presently of three species.
Included species: M. australis DISTANT 1899 Australia M. hirtula BERGROTH 1904 Australia M. schoutedeni BERGROTH 1904 Australia
Remarks: DISTANT (1899) first described Morbora , with SCHOUTEDEN (1904) and GROSS (1975) providing subsequent redescriptions. MCDONALD & CASSIS (1984) redescribed the species, without providing any additional observations for the genus or comments on its relationships.
GROSS (1975) placed Morbora in the Odontotarsinae , on the basis of observations by Fred McDonald of the male genitalia, the latter worker suggesting that they resembled members of the Odontotarsinae . Our observations confirm these findings, and have found further evidence that Morbora is nest-ed within the odontotarsines. As stated above, Morbora has male abdominal androconial glands, in common with some Palearctic odontotarsinae taxa ( Table 3 View Table 3 ). Moreover, we have found that some species of Psacasta (e.g., P. exanthematica SCOPOLI ) have the first gonocoxae divided (bipartite), which is a putative homology to the tripartite GXI condition found in all the species of Morbora .
Table 5: Morbora australis, M. hirtula, and M. schoutedeni: diagnostic measurements in millimetres. N = sample size.
Male | Female | |||||
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N | Mean±SD | Range | N | Mean±SD | Range | |
M. australis | ||||||
Length | 1 | 5.37 | - | 4 | 5.40±0.22 | 5.27-5.73 |
Pronotal width | 1 | 4.41 | - | 4 | 4.17±0.21 | 3.85-4.29 |
Width between eyes | 1 | 1.59 | - | 4 | 1.66±0.13 | 1.54-1.84 |
Antennal segment length | ||||||
I | 2 | 0.64 | 0.61-0.67 | 7 | 0.60±0.03 | 0.54-0.63 |
II | 2 | 0.30 | 0.29-0.30 | 8 | 0.33±0.02 | 0.30-0.36 |
III | 2 | 0.22 | - | 8 | 0.23±0.02 | 0.22-0.26 |
IV | 2 | 0.35 | - | 7 | 0.32±0.02 | 0.28-0.35 |
V | 2 | 0.62 | 0.62-0.63 | 7 | 0.56±0.03 | 0.50-0.59 |
Rostral segment length | ||||||
I | 1 | 0.62 | - | 2 | 0.59 | 0.57-0.61 |
II | 1 | 0.51 | - | 3 | 0.45 | 0.39-0.54 |
III | 1 | 0.45 | - | 2 | 0.45 | 0.42-0.48 |
IV | 1 | 0.34 | - | 2 | 0.33 | - |
Morbora hirtula | ||||||
Length | 3 | 5.06 | 4.97-5.12 | 2 | 4.90 | 4.41-5.39 |
Pronotal width | 3 | 3.64 | 3.58-3.72 | 2 | 3.55 | 3.31-3.80 |
Width between eyes | 3 | 1.53 | 1.47-1.57 | 2 | 1.58 | 1.54-1.62 |
Antennal segment length | ||||||
I | 5 | 0.58±0.04 | 0.53-0.63 | 3 | 0.61 | 0.61-0.62 |
II(a) | 5 | 0.33±0.03 | 0.30-0.36 | 2 | 0.36 | 0.35-0.36 |
II(b) | 4 | 0.23±0.02 | 0.21-0.25 | 2 | 0.22 | 0.22-0.23 |
III | 3 | 0.40 | 0.35-0.42 | 2 | 0.42 | 0.41-0.42 |
IV | 5 | 0.65±0.03 | 0.62-0.70 | 2 | 0.64 | 0.63-0.65 |
Labial segment length | ||||||
I | 1 | 0.82 | - | 1 | 0.85 | - |
II | 1 | 0.97 | - | 1 | 0.91 | - |
III | 1 | 0.36 | - | 1 | 0.42 | - |
IV | 2 | 0.37 | 0.36-0.38 | 1 | 0.45 | - |
Morbora schoutedeni | ||||||
Length | 2 | 4.74 | 4.51-4.97 | 5 | 5.47±0.34 | 5.00-5.88 |
Pronotal width | 2 | 3.79 | 3.65-3.92 | 5 | 4.41±0.28 | 4.17-4.85 |
Width between eyes | 2 | 1.40 | 1.35-1.45 | 5 | 1.70±0.09 | 1.59-1.84 |
Antennal segment length | ||||||
I | 4 | 0.54±0.02 | 0.51-0.57 | 9 | 0.68±0.06 | 0.61-0.76 |
II(a) | 4 | 0.25±0.03 | 0.22-0.28 | 9 | 0.28±0.03 | 0.23-0.33 |
II(b) | 4 | 0.23±0.01 | 0.23-0.24 | 10 | 0.22±0.01 | 0.21-0.24 |
III | 4 | 0.29±0.02 | 0.27-0.31 | 10 | 0.32±0.04 | 0.24-0.39 |
IV | 4 | 0.49±0.03 | 0.46-0.52 | 9 | 0.55±0.04 | 0.46-0.61 |
Labial segment length | ||||||
I | 2 | 0.77 | 0.75-0.79 | 3 | 0.75 | 0.73-0.79 |
II | 2 | 0.77 | 0.76-0.79 | 2 | 0.79 | 0.73-0.85 |
III | 2 | 0.36 | - | 2 | 0.41 | 0.36-0.46 |
IV | 2 | 0.38 | 0.36-0.39 | 1 | 0.43 | - |
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Morbora DISTANT 1899
Gerry Cassis & Loren Vanags 2006 |
Morbora
DISTANT 1899: 47 |