Anomala okushimai, Zorn & Kobayashi & Wada, 2017

Zorn, Carsten, Kobayashi, Hirokazu & Wada, Kaoru, 2017, Notes on the genus Anomala SAMOUELLE, 1819 (Coleoptera, Scarabaeidae, Rutelinae) in Vietnam and neighboring regions: eight new species and faunistic records, Beiträge Zur Entomologie = Contributions to Entomology 67 (2), pp. 325-352 : 326-327

publication ID	https://doi.org/ 10.21248/contrib.entomol.67.2.325-352
publication LSID	lsid:zoobank.org:pub:F2CE9642-9537-46FF-BA3F-5D4148AC1A0B
persistent identifier	https://treatment.plazi.org/id/DB65AF09-2AD7-4B48-A64B-0D74C2A8C830
taxon LSID	lsid:zoobank.org:act:DB65AF09-2AD7-4B48-A64B-0D74C2A8C830
treatment provided by	Felipe
scientific name	Anomala okushimai
status	sp. nov.

Treatment

Anomala okushimai spec. nov.

urn:lsid:zoobank.org:act:DB65AF09-2AD7-4B48-A64B-0D74C2A8C830

( Figs 1 View Figs 1–9 , 10 View Figs 10–13 )

Type series: Holotype: , Tamdao VIETNAM 3rd vi. 1993 K. Matsumoto leg. ( NSMT). Paratypes: 5 , Tam Dao, N. Vietnam, 30-IV~ 6-V-1996, Y. Okushima leg. ( MFPC); 1 , Tam Dao, Vinh Phu Prov., Vietnam, IV~ V- 1996 ( MFPC); 1 , Mt. Tam Dao (alt. 900-950 m), Vinh Phu Prov., N. Vietnam, 14~ 18-V-1999, S. Nomura leg. ( NSMT); 1 , VIETNAM Tam Dao VII.97 BERNAUD leg. | Anomala gr. aulax (CZPC) ; 1 , N.- Vietnam Vinh Phu Prov., Tam Dao 01.-07.V.1998 Y. Arita leg. ( CZPC); 1 , 1 , N. VIETNAM ( TONKIN) TAMDAO 12.- 24.5.1989 PACHOLATKO Leg. ( PPPC, CZPC); 1 , Vietnam N, Vinh Phuc Pr. Tam Dao NP, 1000 m, N: 21°27,577'E: 105'38,489 V.2017, M. Pejcha lgt ( CZPC).

Description: Body shape elongate ovoid, convex; body length: 16.7-18.0 mm, width: 8.2-9.5 mm.

Color. Dorsal surface except pygidium and propygidium deep metallic blue-green or green, sometimes with strong reddish reflections, antennae dark brown, basal two segment somewhat lighter; apical half of propygidium and pygidium orange-brown; pronotum with orange-brown lateral margins; ventral surface and legs predominantly dark metallic green; coxae, femora, lateral portions of thoracal segments as well as basal half of abdominal ventrites (sometimes obscured by greenish luster) and a spot at the sides of the abdominal ventrites 1-4 orange-brown.

Head with erect pale yellow setae (0.07-0.13 mm in length) along eyes; interocular distance equals 0.73 times the maximum transverse head width. Antennal club 0.8 times as long as footstalk. Clypeus broadly trapezoidal to semicircular, densely punctate, the punctures partly coalescent and rugoso-punctate along marginal portion; anterior margin distinctly reflexed, with a shallow impression behind the margin at each side; fronto-clypeal suture almost straight. Frons densely punctate, punctures large (ca. 0.05-0.08 mm), partly coalescent in the middle, becoming larger laterad, sparser and smaller posteriad. Vertex with rather sparse and fine punctures.

Pronotum 1.5 times as wide as long; sides convergent in anterior 2/5, almost straight and subparallel in posterior 3/5; widest at base; anterior and lateral margin with weak marginal line; lateral marginal line extending to hind angles; front angles produced and acute, hind angles obtuse; disc with moderately fine, somewhat transverse punctures (ca. 0.05-0.07 mm) which are intermixed with sparse microscopic punctures (visible under 40×); with a vague longitudinal depression in the middle as well as a vague depression near the basal half of lateral margin and medially of the hind angles.

Scutellum 1.4 times as wide as long, apex angulate; with rather dense, somewhat transverse punctation.

Elytra regularly striate-sulcate; intervals moderately convex, primary costae slightly higher than interstices; sutural interval distinctly elevated in posterior 3/4 of elytra; striae not marked by a regular row of punctures; entire surface covered with dense transverse, microscopic-seta-bearing punctures, leaving out only the ridges of primary costae and the humeral protuberance; punctures adjacent to each other, partly coalescing and forming small transverse striae; epipleura rather narrow, extending until apical curvature; marginal membrane starting approximately at level of anterior third of metacoxa.

Pygidium triangular, widely rounded at apex, somewhat tumid; with very transverse punctures which are adjacent to each other or confluate, forming a striolation which is arranged concentrically around the tumidity; each puncture bearing a microscopic seta (50x); longer pale yellow setae (0.13-0.63 mm in length) along lateral and apical portion.

Mesoventrite very narrow and somewhat concave between the approximated mesocoxae.

Metaventrite reticulately rugulose, with long, decumbent yellow setae (0.13-0.38 mm in length).

Abdominal ventrites densely punctate, punctures strongly transverse, partly coalescent, bearing a short adpressed seta; additional transverse rows of semierect yellow setae (0.05-0.13 mm in length) in the apical half of the 2nd to 4th ventrite.

Legs slender and long; protibia bidentate; terminal tooth somewhat prolonged and obtuse at apex in male; protarsal segment 5 with a stout, broad tooth at basal 1/4; inner protarsal claw and outer mesotarsal claw apically incised; the upper branch of the inner protarsal claw small and slender, the lower branch broad with a distinct tooth at its ventral margin; the upper branch of outer mesotarsal claw slightly more slender than the lower one.

Female: Body shape more stout; sides of pronotum slightly more rounded; terminal tooth of protibia spatulate; modified protarsal claw less enlarged, without ventral tooth.

Diagnosis: The similar shape of the aedeagus indicates a close relationship between this species, Anomala imperialis ARROw, 1899 and Anomala harpagophysa PROKOFIEV, 2014 , but the parameres of the three are quite distinctive, and the latter two species lack adpressed setosity at the abdominal ventrites. Additionally, A. harpagophysa is distinctly larger and entirely metallic green with fiery red shine ventrally, while A. imperialis is smaller, and its elytral intervals have rather sparse and simple punctures (dense and transverse in A. okushimai ). Moreover, A. okushimai is similar externally to Anomala aulax ( WIEDEMANN, 1823) and its nearest relatives, but can be separated from those species by the following characters: pronotum narrower, with a transverse depression medially of the posterior angles; base of pronotum without marginal line; abdominal ventrites with distinct adpressed setosity.

Etymology: The new species is named after Dr. Yûichi Okushima, a Japanese entomologist and collector of several paratypes of this species.