Hispanomys moralesi, López-Antoñanzas & Peláez-Campomanes & Álvarez-Sierra & García-Paredes, 2010

López-Antoñanzas, Raquel, Peláez-Campomanes, Pablo, Álvarez-Sierra, M. Ángeles & García-Paredes, Israel, 2010, New species of Hispanomys (Rodentia, Cricetodontinae) from the Upper Miocene of Batallones (Madrid, Spain), Zoological Journal of the Linnean Society 160 (4), pp. 725-747 : 729-732

SPECIES HISPANOMYS MORALESI SP. NOV.

( FIGS 3 , 4 )

Chresonymy: Cricetidae indet. Morales et al., 1992: 80; Cricetidae indet. Morales et al., 2000: 186; Hispanomys sp. Morales et al., 2004: 140; Hispanomys sp. Morales et al., 2008: 76; Hispanomys peralensis López-Antoñanzas et al., 2008 .

Etymology: In honour of Jorge Morales (MNCN-CSIC, Madrid) to whom we are indebted for the development of the Batallones research programme.

Holotype: BAT10′07-G3-23: left maxillary fragment with M2–M3, left maxillary fragment with M1, right maxillary fragment with M1, and right maxillary fragment with the M3 ( Fig. 4A–D ). All these fragments come from the same individual.

Hypodigm: Fragmentary skulls with complete maxillae: BAT3′07-728; BAT3M′08-31; complete maxillae: BAT1′03-D3a; BAT10′07-F4-9-a; BAT10′07-G4-60, BAT3M′06-5; BATM 4227-6a; BAT3M′08-30; BAT10′08 F3-4; maxillary fragments: BAT10′07-G3- 23a, with M1; BAT10′07-G3-23a, with M2-M3; BAT10′07-G3-23b, with M1; BAT10′07-G3-23b, with M3; BATM 4227-6b, with M2; BATM 4227-6b, with M3; BAT10′08G4-50, with M1; BAT3M′08-33, with M2–M3; M1: BAT1M′02-E5-1; BAT1M′02-D7-2; BAT5M′02-3; BAT1M′03-D3-4; BAT10′07-F4-9; M2: BAT1M′02-D7-7; BAT5M′02-8; BAT1′01-D4-19; BAT3M′08-32; M3: BAT1M′01-D4-24; complete mandibles: BAT1′03F6-60; BAT1′03-F6-79a; BAT1′03-F6- 79b; BAT10′07-G4-36; BAT3M′06-17; BAT10′07-E3-4; BAT10′07-F4-72; BAT1′03-F4-30; BAT3M′08-34; BAT3M′08-35; BAT3M′08-37; BAT10′08 G4-37; BAT10′08 F3-12; mandibular fragments: BAT1′05-E8- 15, with m1–m2; BAT10′07-G3-29, with m1–m2; BAT10M′07-H4-14, with m1; BAT10M′07-H4-15, with m1–m2 and the m3 isolated, BAT10M′07-G4-16, with broken m1 and m2–m3; BAT10M′07-G4-24, with m2–m3; BAT10′07-G4-58, with m1–m3; BAT3M′08-36 with m1–m3; BAT10′08 F3-9 with m1–m2; BAT10′08 F2-19, with m1–m2; BAT10′08 G4-59, with m1–m2; m1: BAT1M′05-C3-9, BAT1M′05-E3-10, BAT1′03- F5-11, BAT1M′02-D7-12, BAT1M-E5-13, BAT1′01- D4-18; BAT3M′08-38; BAT3M′08-39; BAT3M′08-40; BAT3M′08-41; BAT10′08 G4-49; BAT10M′07-G4-47; m2: BAT5M′06-I16-18, BAT1M′04-D6-19, BAT1M′02- D7-20, BAT5M′02-21, BAT1M′02-E5-22, BAT1M′ 05-E3-23, BATM′91-25, BAT1M′03-F5-26, BAT1M′03- F5-27; BAT3M′08-42; BAT3M′08-43; BAT3M′08-44; BAT3M′08-45; BAT3M′08-46; m3: BAT1M′03-F5-28, BAT1M′03-F5-29; BAT3M′08-48. Size data are provided in Table 1 and Appendix 1.

Repository institution: MNCN-CSIC collections.

Type locality: Batallones 10 (Madrid, Spain).

Age: Late Vallesian, Late Miocene (MN10).

Other localities: Batallones 1, Batallones 3, Batallones 5 (Madrid, Spain).

Diagnosis: Hispanomys species of medium size, with high values of the (mean LM1)/(mean LM3) and (mean Lm1)/(mean Lm3) ratios; upper molars lacking cingula on the labial valleys as well as mesoloph, four- or five-rooted M1; M1, and M2 with almost complete ectolophs, M3 relatively small; lower molars lacking cingula on the lingual valleys, either absent or short mesolophids, and long, short or absent anterolophid.

Differential diagnosis: Differing from Hispanomys castelnovi , H. decedens , H. bijugatus , H. aguirrei , H. daamsi , H. dispectus , H. lavocati , H. aragonensis , and H. baixasi in having four- or five-rooted M1; differing from H. castelnovi , H. decedens , H. bijugatus , H. aguirrei , H. daamsi , H. dispectus , H. lavocati , H. aragonensis , and H. baixasi in having well-developed ectolophs on the M1 and M2; differing from H. decedens , H. dispectus , H. thaleri , and H. aragonensis in lacking enamel-coated valley on the M1–M2; differing from H. decedens , H. aguirrei , H. daamsi , H. dispectus , H. lavocati , and H. nombrevillae in lacking labial cingula on the M1–M2; differing from H. decedens , H. aguirrei , H. dispectus , H. lavocati , H. thaleri , and H. aragonensis in lacking any trace of mesoloph on the upper molars; differing from H. castelnovi , H. decedens , H. bijugatus , H. aguirrei , H. daamsi , H. dispectus , H. lavocati , H. thaleri , H. aragonensis , H. peralensis , and H. baixasi in having larger LM3/LM1 ratio; differing from H. castelnovi , H. decedens , H. bijugatus , H. aguirrei , H. daamsi , H. dispectus , H. lavocati , H. nombrevillae , H. thaleri , H. aragonensis , H. mediterraneus , and H. baixasi in lacking labial anterolophid in about half the m1; differing from H. decedens , H. bijugatus , H. aguirrei , H. lavocati , H. nombrevillae , H. mediterraneus , and H. baixasi in lacking double metalophulid on the m1; differing from H. freudenthali and H. adroveri in having labial anterolophid in about half the m1 and very short or absent mesolophids in more than half the specimens; differing from H. aragonensis , H. thaleri , H. nombrevillae , and H. aguirrei in having the m2 three-rooted; differing from H. castelnovi , H. decedens , H. bijugatus , H. aguirrei , H. daamsi , H. dispectus , H. lavocati , H. thaleri , H. aragonensis , and H. baixasi in having larger Lm3/Lm1 ratio.

DESCRIPTION

Figures

Figure 3. Lower molars of Hispanomys moralesi sp. nov. A, left m1 (BAT1M′02-D7-12); B, right m2 (BAT1M′02-D7- 20); C, right m3 (BAT1M0′07-H4-15); D, right m1 (BAT10′07-G4-58); E, right m2 (BAT10′07-G4-58); F, right m3 (BAT10′07-G4-58); G, left m1 (BAT1M′03-F5-11); H, left m2 (BAT1M′03-F5-26); I, left m3 (BAT1M′03-F5-28); J, left mandible with m1–m3 (BAT1′03F6-60). Scale bar = 1 mm.

Figure 4. Upper molars of Hispanomys moralesi sp. nov. A, right maxilla fragment with M2–M3 (BAT10′07-G3-23a); B, right M1 (BAT10′07-G3-23a); C, left M1 (BAT10′07-G3-23b); D, left M3 (BAT10′07-G3-23b); E, left M1 (BAT1M′02- E5-1); F, left M1 (BAT1M′02-D7-2); G, right M2 (BAT1M′02-D7-7); H, right M3 (BAT1M′01-D4-24); I, left maxilla with M1–M3 (BAT10′07-F4-9); J, right maxilla with M1–M3 (BAT1M′03-D3-4). BAT10′07-G3-23 is the holotype. Scale bar = 1 mm.