Cyanus reuterianus var. phrygius (Bornm.) Negaresh

Negaresh, Kazem, 2022, A checklist of the genus Cyanus (Asteraceae, Cardueae) in Turkey, with notes on taxonomy and distribution, Phytotaxa 531 (1), pp. 18-40 : 30-31

14b. Cyanus reuterianus var. phrygius (Bornm.) Negaresh , comb. nova.

Basionym: Centaurea reuteriana var. phrygia Bornmüller (1909: 474) . Type :— TURKEY. Phrygia : Akscheher (Wilajet Konia), in regione alpina montis Sultandagh, in rupestr. et saxosis, 2000 m, 21 June 1899, Bornmüller 5201 (lectotype B 10 0367361! designated here, isolectotypes B 10 0367362!, G00074351!, G-BOIS! (G00074351), JE00015820!, W19000001934!).

Taxonomic notes: — Centaurea reuteriana var. phrygia was first described by Bornmüller (1909) from Turkey. According to him, it is distinguished from C. reuteriana var. reuteriana by creamy-white flowers (vs. pink- to violetpurple). Currently this taxon is accepted as a distinct variety by subsequent authors such as Wagenitz (1975), Boršić et al. (2011), Şirin et al. (2019) and Çitak et al. (2021). This variety has still not been transferred to the genus Cyanus , so I propose the new combination.

Six duplicates of the type of Centaurea reuteriana var. phrygia ( Cyanus reuterianus var. phrygius ) were found at B, G, G-BOIS, JE, and W, which are to be treated as syntypes (Art. 40 Note 1 of Turland et al. 2018). According to McNeill (2014), the existence of a holotype cannot be established in this case because there was more than one specimen for this taxon, housed in more than one institution. The sheet B 10 0367361 is designated here as the lectotype, because it is the original material that was annotated by Bornmüller and contains capitula and flowers that are better developed than in the other specimens.

On the label of the lectotype is handwritten with the locality as above. On the label of all the isolectotypes the following text is printed: “Phrygia: Akscheher (Wilajet Konia), in regione alpina montis Sultandagh, 2000 m, 21 June 1899, Bornmüller ” and handwritten: “5201 Centaurea reuteriana Boiss. var. phrygia ”.

Distribution: — Cyanus reuterianus var. phrygius is endemic to Isparta, Sivas, Zonguldak, Konya, Karaman, Amasya, Denizli and Adana Provinces, S to SW and N Turkey. Its distribution is wider in comparison to the other variety ( Fig. 7 ).

Selected specimens examined:— TURKEY. C3 Isparta: in pascuis saxosis montis Davros Da. , 1950 m, May 1845, Heldreich s.n. (E!, G!, G-DC!, W!); 29 May 1845, Heldreich 746 (G-BOIS!); mounad-Dagh (Phrygie), 28 June 1857, Blanche 122 (G-BOIS!); Barla Dağ, Ostgipfel, Seehöhe, 1600–2100 m, 13 June 1988, Speta s.n. (W!). A6 Sivas: Koyulhisar, 1858, Tchihatcheff 817 (G-BOIS!). A4 Zonguldak: Keltepe, above Karabük, 1950 m, 3 August 1962, Davis & Coode D38916 (E!). B3 Konya: Sultan Da. near Akşehir, 1890–1910 m, Huber-Morath 8673 ( Hb. Hub. - Mor. !); 2 km from Camlik village , Akseki-Beysehir area , 1360–1370 m, 3 June 2003, Reeve & Adigüzel 2680 (E!) . Amasya: Akdag, above Zefe Koy , Grid A 6, 1900 m, 26 June 1965, Tobey 1208 (E!); in cacumine montis Sana-dagh, 1500–1600 m, 15–16 July 1899, Bornmüller 2246 (B!); in montis Ak-dagh regione alpine, 1500–1900 m, 18 June 1889, Bornmüller 539 (B!); Abadschi-dagh, in reg. alpina, 1300 m, 13 May 1890, Bornmüller s.n. (B!). s.c., Whittall 544 (E!); Whittall 545 (E!) .

Figures

FIGURE 7. Distribution of Cyanus reuterianus var. reuterianus (circles), C. reuterianus var. phrygius (squares), and C. raimondoi (triangle) in Turkey.