Phoxinus, Rafinesque, 1820

Denys, Gaël P. J., Dettai, Agnès, Persat, Henri, Daszkiewicz, Piotr, Hautecoeur, Mélyne & Keith, Philippe, 2020, Revision of Phoxinus in France with the description of two new species (Teleostei, Leuciscidae), Cybium 44 (3), pp. 205-238 : 226

Key to French species of Phoxinus (male)

1a. – Z4 green; belly continuously red between pectoral and caudalfin bases. .. ... ... ... ... ... ..... Phoxinus csikii

1b. – Z4 yellow or greyish; belly white, grey, black or red, if red, then red colour not extended to caudal fin base...... 2

2a. – Flank bars reaching from Z2 to Z3, forming a stripe along Z3 .. ... ... ... ... ... ... ... ... ... ... ... .... 3

2b. – Flank bars reaching from Z2 to Z4 or Z5, not forming a stripe along Z3. ... ... ... ... ... ... ... ... ... ... .... 4

3a. – Agreen stripe along Z3; Z4 yellow-green; pectoral fin yellow; no yellow dot between mouth and eye; lateral line incomplete, usually reaching to a point anterior of anal fin origin or slightly beyond base of last anal fin ray; 50-74 pored scales along lateral line; snout slightly stout; two scaled patches at pectoral fin bases separated by an unscaled area ............................ Phoxinus dragarum

3b. – Ablack stripe along Z3; Z4 greyish; pectoral fin yel- low-orange with a red blotch behind inner fin base; a yel- low dot between mouth and eye; lateral line complete or incomplete, reaching beyond anal fin base to caudal fin base in some individuals; 73-86 pored scales along lateral line; snout slightly pointed; breast completely covered by scales between pectoral fin bases in some populations (two scaled patches at pectoral fin bases separated by an unscaled area in others). ... ... ... ... ... ... ... .. Phoxinusseptimaniae

4a. – Bars crossing Z1 and Z5 black; belly greyish to black; analfinmarginstraighttoconvex ....... Phoxinusbigerii

4b. – Bars crossing Z3 and Z4 green; belly red, pinkish or pale orange; anal fin margin straight to slightly concave... .............................................. 5

5a. – Z4 bright yellow; flank bars green with sharp margins; lips, pectoral, pelvic and anal fin bases and belly red; pecto- ral fin orange; snout stout and blunt................... ............................... Phoxinusfayollarum

5b. – Z4 greenish yellow; flank bars green with diffuse mar- gin; lips, pectoral, pelvic and anal fin bases pinkish, belly pinkish or pale orange; pectoral-fin yellow; snout slightly pointed. ... ... ... ... ... ... ... .... Phoxinus phoxinus

Key to French species of Phoxinus (female)

1a. – Flank bars reaching from Z2 to Z4, not forming a stripe along Z3; Z4 greyish ................. Phoxinusbigerri

1b. – Flank bars reaching from Z2 to Z3, forming a stripe along Z3; Z4 yellow, greenor white.. ................. 2

2a. – Stripe along Z3 green, Z4 bright yellow; belly below pectoral fin origin red; pectoral, pelvic and anal fin bases red. ........................... Phoxinusfayollarum

2b. – Stripe along Z3 black, Z4 golden, white or greenish; belly completely white; pectoral, pelvic and anal fin bases pinkish. .. ... ... ... ... ... ... ... ... ... ... ... ... .. 3

3a. – Z4 golden; Z5 white; anal fin margin straight to slightly convex. .. ... ... ... ... ... ... ... ... ... Phoxinus csikii

3b. – Z4 white or greenish; Z5 black; anal fin margin straight toslightly concave................................ 4

4a. – Z4 white; breast between pectoral fin origins com- pletely covered by scales in some populations (two scaled patches at pectoral fin bases separated by an unscaled area in others). .. ... ... ... ... ... ... ... Phoxinusseptimaniae

4b. – Z4 greenish; breast between pectoral fin origins with scale patches separated by unscaled area or connected by 1-2 rowsof scales.................................... 5

5a. – Z4 green; lateral line incomplete, reaching to beyond anal fin base, to caudal peduncle; snout slightly pointed.... ................................ Phoxinusphoxinus

5b. – Z4 green-ochre; lateral line incomplete, usually reach- ing to a point anterior of anal fin origin or slightly beyond baseoflastanalfin ray; snoutslightly stout.. ............ ................................ Phoxinus dragarum