Chthonius

Zaragoza, Juan A., 2017, Revision of the Ephippiochthonius complex in the Iberian Peninsula, Balearic Islands and Macaronesia, with proposed changes to the status of the Chthonius subgenera (Pseudoscorpiones, Chthoniidae), Zootaxa 4246 (1), pp. 1-221 : 19-20

Key to adults of the Chthonius -related genera (coxae II and III with coxal spines)

1 Intercoxal tubercle absent.............................................................................. 2

- Intercoxal tubercle present, bisetose.......................................................................4

2 Trichobothrium sb of movable chelal finger placed approximately halfway between trichobothria b and st .............. 3

- Trichobothrium sb of movable chelal finger distinctly closer to st than to b, forming with st and t a separate group with respect to b ....................................................................... Spelyngochthonius Beier, 1955

3 Chela not lagyniform, fixed finger sigmoid in lateral view. Distal marginal seta on pedipalpal coxa distinctly longer than distal marginal seta on coxa I.......................................................... Microchthonius Hadži, 1933

- Chela lagyniform, fixed finger straight in lateral view. Distal marginal seta on pedipalpal coxa (dcs) markedly shorter than the

distal marginal seta on coxa I (dps).................................................. Cantabrochthonius n. gen. 4 Male genitalia with median hiatus dividing each row of internal guard-setae into two groups of two setae................................................................................. Hesperochthonius Muchmore, 1968 , n. stat.

- Male genitalia without median hiatus dividing each row of internal guard-setae................................... 5

5 Coxal spines of coxa II arranged in a transverse row............................... Neochthonius Chamberlin, 1929

- Coxal spines of coxa II irregularly arranged in a rounded patch................................................ 6

6 Chela lagyniform; chelal fingers mostly with large, upright and well-spaced teeth; apodeme of movable chelal finger usually well developed and strongly sclerotized.................................................................. 7

- Chelal not lagyniform, hand fusiform; chelal fingers usually with more or less close-set teeth, distinctly reclined backwards; apodeme of movable chelal finger absent or poorly developed............................ Chthonius C.L. Koch, 1843

7 Proximal portion of chelal hand with 4 setae in adults and tritonymphs, seta hp3 present. Tip of fixed chelal finger of male, tritonymph and deutonymph usually without a deep hollow on paraxial face and subapical protuberance (sp) (sometimes present in Globochthonius ); without a medial protuberance (ip) between chelal condyles............................... 8

- Proximal portion of chelal hand with 3 setae in adults and tritonymphs, seta hp3 absent. Tip of fixed chelal finger of male, tritonymph and deutonymph with a deep hollow on paraxial face and subapical protuberance (sp); medial protuberance (ip) present between chelal condyles.......................................... Ephippiochthonius Beier, 1930 , n. stat.

8 Chelal hand distad of trichobothria ib / isb with a rounded hump and a marked hollow before base of movable finger; base of chelal hand without a pronounced hump dorso-paraxially; third tooth of normal row (mt) of fixed chelal finger modified in shape and deviated in orientation with respect to the other teeth; movable chelal finger lacking lyrifissures ma1 and ma2 ..................................................................................... Occidenchthonius n. gen.

- Chelal hand without a hump distad of trichobothria ib / isb, gradually constricted distad of trichobothria ib / isb and without hollow before base of movable finger; base of chelal hand with a pronounced hump dorso-paraxially; third tooth of normal row (mt) of fixed chelal finger neither modified in shape nor deviated in orientation with respect to the other teeth; lyrifissures ma1 and ma2 present on movable chelal finger...................................... Globochthonius Beier, 1931 , n. stat.