Pseudobironium horaki, Lobl & Liang Tang, 2013

Löbl, Ivan & Tang, Liang, 2013, A review of the genus Pseudobironium Pic (Coleoptera: Staphylinidae: Scaphidiinae), Revue suisse de Zoologie 120 (4), pp. 665-734 : 701-702

Pseudobironium horaki sp. nov. Figs 52-54

HOLOTYPE Ƌ: MALAYSIA, Sarawak, Kapit Distr., Rumah Ugap village, Sut river , 3.-9.III. 1994, J. Horák ( MHNG).

PARATYPES: INDONESIA, Java, Idjan Plateau, K. O. Blawan, 900-1500m, 4.XII.1933, 10, I., 5.II.1934, and 20.I.1935, 2 Ƌ, 4 ♀, 5 ex. sex not examined leg. H. Lucht ( MHNG, ZMBC) . – W Java Gn. Halimun Nat. Park 5 km W Cipedey ca 1000m, 22.8.1994, 5 ♀ leg. R. Schuh ( NHMW, MHNG) . – MALAYSIA: Sarawak, Kuching Distr., Mt. Serapi , 27.-29.III.1994, 1 ♀ leg. J. Horák ( MHNG) . – NW Borneo, Sarawak, Belaga , 15.III.1990, 1 Ƌ A. Riedel ( NMEC) ; Sabah, Poling nr. Ranau , 28.IV.1980, 1 ♀ leg. M. & A. Sakai ( EUMJ) . – Borneo , Sabah, km 53 road KK-Tambunan Gn. Emas , 1650m, 22.III.- 6.IV.2000, 1 Ƌ, 1 ♀ leg. Bolm ( NHMB, MHNG) .

DESCRIPTION: Length 2.35-2.70 mm, width 1.70-2.00 mm. Head brown. Pronotum reddish-brown, with variably large, usually not well delimited dark median spot, often extended from anterior margin to base and narrowed basally. Pronotal base narrowly darkened. Elytra reddish-brown, each with darkened humeral area, dark, not well delimited spot or transverse band situated posterior elytral mid-length, and narrowly darkened apex; area posterior spot or transverse band lighter than area anterior spot. Prohypomera and abdomen reddish-brown as most of pronotum, apical abdominal segments sometimes lighter, mesoventrite and metaventrite darker brown. Femora and tibiae reddish-brown, tarsi and antennomeres I to VI light reddish or almost yellowish, following antennomeres infuscate. Maxillary palpi with palpomere IV about 2.5 times as long as wide, tapering. Antennae long, length ratio of antennomeres as II 10: III 6: IV 9: V 12: VI 10: VII 13: VIII 12: IX 14: X 13: XI 18; antennomere XI about three times as long as wide. Pronotum with punctation dense and distinct on median area, with punctures usually well delimited and some about as large as puncture interval, punctation inconspicuous on lateral areas. Elytral punctation in average slightly finer than that of pronotum, most puncture intervals about twice to three times as large as puncture diameters. Inferior part of prohypomera finely punctate, superior part impunctate. Mesoventrite punctate in middle and along margin, with short median ridge. Metaventrite with transversely strigulate microsculpture sometimes almost evanescent near lateral margins; with median part convex and distinctly punctate, except along smooth mesal line. Lateral parts of metaventrite very finely punctate. Submesocoxal area about as long as one third of interval to metacoxa. Metacoxal process smooth, lacking stria or tubercle, with margin hardly concave. Protibiae and mesotibiae almost straight, metatibiae slightly curved, mesotibial ventral spur straight. Abdominal sternite 1 very finely punctate, with strigulate microsculpture, lateral, very shallow, foveiform impressions and slightly impressed oblique lines.

Male: Protarsus shorter than protibia, with tarsomeres 1 to 3 moderately widened and bearing short tenent setae. Aedeagus (Figs 52-54) 0.73-0.77 mm long. Median lobe with basal bulb much shorter than apical process, latter weakly inflexed, with ventral side sinuate, tip blunt, rounded. Parameres narrow, weakly curved in dorsal and lateral views, with small apical lobe. Internal sac with mesal guide-sclerites, pair of conic sclerites each including one narrow rod and basally joined by membranes to irregular sclerotized piece; membranes posterior sclerites bearing long spicules.

DISTRIBUTION: Malaysia: Sarawak, Sabah; Indonesia: Java.

ETYMOLOGY: The species is named in honour of Jan Horák, Praha, Czech

Republic, who is one of its collectors.

COMMENTS: The colour pattern of this species is distinctive. Only P. bicolor and P. parabicolor have a similar coloration. In addition to its colour pattern, P. horaki is characterized by the distinctly punctate inferior part of the prohypomera, strigulate microsculpture on metaventrite and on abdominal sternite 1, and weekly lobed apices of the parameres, in combination. The examined aedeagi have the membranes of the internal sac bearing long spicules, similar to those in P. parabicolor , while the parameres are weekly curved and have a small apical lobe. Apically lobed parameres are also in P. bicolor , the aedeagus of latter differs conspicuously by the basal bulb about as long as apical process of median lobe, and by the tip of the median lobe very short and abruptly inflexed ventrally.