Conocephalus (Anisoptera) maculatus (Le Gouillou, 1841)

Nagar, Rajendra & Swaminathan, R., 2016, Notes on the genus Conocephalus (Orthoptera: Tettigoniidae; Conocephalinae; Conocephalini) from India and description of two new species, Zootaxa 4126 (1), pp. 1-43: 13-20

publication ID

http://doi.org/10.11646/zootaxa.4126.1.1

publication LSID

lsid:zoobank.org:pub:71409CFF-5A38-493F-80D7-56D34D2739D2

persistent identifier

http://treatment.plazi.org/id/038A8781-0B2A-FF85-A7CE-7778FAF3FAFC

treatment provided by

Plazi

scientific name

Conocephalus (Anisoptera) maculatus (Le Gouillou, 1841)
status

 

Conocephalus (Anisoptera) maculatus (Le Gouillou, 1841) 

( Figs. 3View FIGURE 3, 11View FIGURE 11, 19, 27, 35, 42, 49, 56, 63, 70 & 78; Plate VI, VII & VIII) http://lsid.speciesfile.org/urn:lsid: Orthoptera  .speciesfile.org:TaxonName: 17438

Xiphidium maculatum Le Guillou, 1841: 294  ; Locusta (Xiphidium) lepida De Haan, 1842: 189  ; Locusta (Xiphidium) continuum Walker, 1869: 271  ; Locusta (Xiphidium) maculatum Walker, 1869: 275  ; Xiphidium sinense Walker, 1871: 35  ; Anisoptera maculatum Kirby, 1906: 278  ; Xiphidion maculatum Karny, 1907: 93  ; Xiphidium maculatum Matsumura et Shiraki, 1908: 51  ; Xiphidion neglectum Bruner, 1920: 123  ; Conocephalus (Xiphidion) maculatus Karny, 1912: 11  ; Conocephalus maculatus Bolivar, 1913: 8  ; Conocephalus sinensis Jin et Xia, 1994: 33  .

Material examined. (371 Specimens, 171 ♂ & 200 ♀). Paratypes: India: Rajasthan; 18.v. 2006, Coll. R. Swaminathan, (Jalore) (2); 14.vi. 2006, Coll. D. P. Choudhary (Banswara) (1); 14.vi. 2006, Coll. B. K. Patidar (Banswara); 19.ii. 2006, Coll. B. K. Patidar (Jhalawar); 21.ix. 2006, Coll. B. K. Patidar (Banswara); 04.xi. 2006, Coll. D. P. Choudhary (Banswara); 28.x. 2006, Coll. G. S. Chundawat (RCA, Udaipur) (2); 08.viii. 2006, Coll. D. P. Choudhary, Kanpur (Udaipur); 08.viii. 2006, Coll. G. S. Chundawat, Kanpur (Udaipur); 26.ix. 2006, Coll. D. P. Choudhary ( CTAE, Udaipur); 08.x. 2006, Coll. B. K. Patidar (Indore); 06.vii. 2006, Coll. B. K. Patidar (ARS, Kota); 08.x. 2006, Coll. D. P.Choudhary (Indore) (1); 08.x. 2006, Coll. B. K. Patidar (Jabalpur) (1); D. P. Choudhary; 10.viii. 2006, Coll. D. P. Choudhary, (Karbadia, Udaipur); 19.ii. 2007, Coll. B. K. Patidar, Jhadol (Udaipur) (5); 15.ii. 2007, Coll. B. K. Patidar, Jhadol (Udaipur); 18.vi. 2007, Coll. B. K. Patidar, Metwala (Banswara) 19.ii. 2007, Coll. P. S. Rathore, Jhadol (Udaipur) (4); 17.iv. 2007, Coll. S. L. Jat (RCA, Udaipur); 02.iv. 2007, Coll. Lekha (RCA, Udaipur); 19.v. 2007, Coll. S. L. Jat (RCA, Udaipur); 30.i. 2007, Coll. P. S. Rathore (Sirohi) (1); 30.ix. 2007, Coll. S. L. Jat (RCA, Udaipur); 30.ix. 2007, Coll. Lekha (RCA, Udaipur); 01.viii. 2007, Coll. S. L. Jat, Jobner (Jaipur); 17.vii. 2007, Coll. B. K. Patidar (Dungarpur); 23.iii. 2007, Coll. R. S. Meena (RCA, Udaipiur); 17.iv. 2007, Coll. S. L. Jat (Hort. Farm, Udaipur); 14.iv. 2007, Coll. S. L. Jat (RCA. Farm, Udaipur) (4); 13.iv. 2007, Coll. B. K. Patidar (RCA, Udaipur) (5); 30.iv. 2007, Coll. B. K. Patidar (RCA, Udaipur); 14.iv. 2007, Coll. R. S. Meena (RCA, Udaipur); 24.iii. 2007, Coll. Lekha (RCA, Udaipur); 30.ix. 2007, Coll. Lekha (RCA, Udaipur); 14.iv. 2007, Coll. Lekha (RCA, Udaipur), (11); 13.iv. 2007, Coll. Lekha (RCA, Udaipur) (4); 03.ix. 2007, Coll. Lekha (RCA, Udaipur); 12.ix. 2007, Coll. Lekha (RCA, Udaipur); 27.x. 2007, Coll. Lekha (RCA, Udaipur); 24.viii. 2007, Coll. P. S. Rathore (Dungarpur) (1); 04.x. 2007, Coll. P. S. Rathore, Rao Madra (Udaipur); 15.x. 2007, Coll. P. S. Rathore (RCA, Udaipur) (3); 29.ix. 2007, Coll. P. S. Rathore (RCA, Udaipur); 16.ix. 2007, Coll. P. S. Rathore, ARS-Mandor (Jodhpur) (1); 03. xi. 2 0 0 7, Coll. P. S. Rathore, Bagawas (Pali) (1); 13.x. 2007, Coll. P. S. Rathore, Sisarama (Udaipur); 30.i. 2007, Coll. P. S. Rathore (Sirohi) (4); 27.x. 2007, Coll. P. S. Rathore (Bhilwara) (3); 01.ix. 2007, Coll. P. S. Rathore (Banswara); 11.ix. 2007, Coll. P. S. Rathore, Mavali (Udaipur); 28.xi. 2007, Coll. P. S. Rathore, Fatehnagar (Udaipur); 27.viii. 2007, Coll. Lekha ( NRCS, Indore); 13.x. 2007, Coll.R. Swaminathan, Sisarama (Udaipur); 17.vi. 2007, Coll. R. Swaminathan, Barda(Dungarpur); 25.xii. 2007, Coll. R. Swaminathan (Jhalawar); 27.x. 2007, Coll. R. Swaminathan (Bhilwara) (4); 24.viii. 2007, Coll. S. L. Jat (Dungarpur); 02.xi. 2007, Coll. S. L. Jat, Ghatol (Udaipur); 17.vi. 2007, Coll. R. S. Meena, Barda (Dungarpur); 17.vi. 2007, Coll. R. Swaminathan, Paldewara (Dungarpur); 14.iv. 2007, Coll. R. S. Meena (RCA, Udaipur) (1); 13.iv. 2007, Coll. R. S. Meena (RCA, Udaipur); 18.iv. 2007, Coll. S. L. Jat (RCA, Udaipur) (3); 02.iv. 2007, Coll. R. S. Meena (RCA, Udaipur), 18.v. 2007, Coll. R. S. Meena (RCA, Udaipur); 03.x. 2007, Coll. S. L. Jat, Revdhar (Udaipur) (2); 15.xi. 2007, Coll. P. S. Rathore (RCA, Udaipur); 30.i. 2007, Coll. P. S. Rathore (Bhilwara); 13.x. 2007, Coll. Coll. Hansa, Sisarama (Udaipur); 29.x. 2007, Coll. S. L. Jat (RCA, Udaipur); 20.vii. 2007, Coll. S. L. Jat (RCA, Udaipur); 20.i. 2007, Coll. G. S. Chundawat (Khedbrama); 26.vii. 2007, Coll. B. K. Patidar, KVK Piprone (Neemuch); 19.viii. 2008, Coll. A. Pareek (Banswara) (2); 19.viii. 2008, Coll. P. S. Rathore (Banswara); 06.xi. 2008, Coll. P. S. Rathore, Chikalwas (Udaipur); 17.ii. 2008, Coll. S. L. Jat (Navsari) (1); 04.xi. 2008, Coll. P.S. Rathore, Bhuasa (Banswara); 04.xi. 2008, Coll. A. Pareek, Bhuasa (Banswara) (3); 15.vi. 2008, Coll. A. Pareek (Udaipur); 02.vi. 2013, Coll. R. Swaminathan (Larbang); 02.vi. 2013, Coll. R. Swaminathan (Mawflang) (1); 09.vi. 2013, Coll. Jhabar Mal (Bhoirymbong) (2); 02.vi. 2013, Coll. Rajendra Nagar (Mawflang) (7); 09.vi. 2013, Coll. R. Swaminathan (Bhoirymbong) (2); 01.vii. 2013, Coll. R. Swaminathan (Anand); 06.vi. 2013, Coll. Rajendra Nagar (Dawki) (2); 02.vi. 2013, Coll. Jhabar Mal (Mawflang) (2); 04.vi. 2013, Coll. Jhabar Mal, Umiam (Ri-bhoi); 04.vi. 2013, Coll. R. Swaminathan, Umiam (Ri-bhoi); 09.vi. 2013, Coll. Rajendra Nagar (Bhoirymbong); 02.vi. 2013, Coll. Rajendra Nagar (Mawbri); 02.vi. 2013, Coll. Jhabar Mal (Mawbri); 04.vi. 2013, Coll. Rajendra Nagar (Khasi Hill West); 02.vi. 2013, Coll. Jhabar Mal (Khasi Hill West); 01.xi. 2013, Coll. R. Swaminathan (KVK, Sirohi); 01.viii. 2013, Coll. Rajendra Nagar (Rajasmand); 01.xi. 2013, Coll. Devendra (KVK, Sirohi); 07.ix. 2013, Coll. Rajendra Nagar (RCA, Udaipur) (1); 01.v. 2013, Coll. Jhabar Mal (ARS, Banswara) (1); 31.x. 2014, Coll. Rajendra Nagar (ARS, Banswara); 01.xi. 2012, Coll. Devendra (KVK, Chitorgarh); 20 iv. 2013, Coll. Shivangi (RCA, Udaipur); 01.viii. 2012, Coll. R. Swaminathan (RCA, Udaipur) (1); 01.x. 2012, Coll. Devendra (RCA, Udaipur); 01.x. 2012, Coll. S. L. Jat (Banswara); 01.v. 2012, Coll. R. Swaminathan ( TNAU, Coimbatore) (3); 01.v. 2012, Coll. Tatiana ( TNAU, Coimbatore) (1); 12.vii. 2012, Coll. Subhash (Anand); 01.xi. 2012, Coll. Rajendra Nagar (Mandsaur) (2) 01.i. 2012, Coll. Jhabar Mal (Jaipur); 25.ii. 2014, Coll. S. K. Yadav (RCA, Udaipur); 04.iii. 2014, Coll. N. Y. Chanu (RCA, Udaipur); 26.ix. 2012, Coll. Rajendra Nagar (Alsigarh, Udaipur) (4); 13.x. 2013, Coll. Rajendra Nagar (Khemli, Udaipur); 08.ix. 2015, Coll. Rajendra Nagar (Kotda, Udaipur); 10.viii. 2006, Coll. G. S. Chundawat, Kavita (Udaipur); 04.. x. 2013, Coll. Jhabar Mal (Chikaldhara); 16.iii. 2015, Coll. Rajendra Nagar( ARSS, Vallabhanagar); 26.ix. 2012, Coll. Rajendra Nagar, Alsigarh (Udaipur); 13.iv. 2007, Coll. B. K. Patidar (RCA, Udaipur); 25.v. 2007, Coll. P.S. Rathore (Jaisamand); 04.xi. 2006, Coll. D.P. Choudhary (Banswara); 20.ix. 2015, Coll. Rajendra Nagar, RARI, Durgapura (Jaipur); 14.iv. 2007, Coll. Lekha (RCA, Udaipur); 14.iv. 2007, Coll. R. S. Meena (RCA, Udaipur); 21.i. 2014, Coll. Rajendra Nagar (Thiruvanthapuram) (6); 21.i. 2014, Coll. Jhabar Mal (Thiruvanthapuram) (5); 22.i. 2014, Coll. Rajendra Nagar ( TNAU, Coimbatore) (22); PLATE VI. C. maculatus  male 1–6: 1. Male habitus; 2. Female habitus; 3. lateral view of hind femur; 4. Palpi; 5. Mid tibia; 6. Fore tibia.

PLATE VII. C. maculatus  male male 7–14: 7. Dorsal view of head with pronotum; 8. Lateral view of pronotum; 9. Fastigium verticis; 10. Mirror on right tegmina; 11. Dorsal view of stridulatory file; 12. Ventral view of stridulatory file on left tegmina; 13. Tubercle spines; 14. Tegmina.

PLATE VIII. C. maculatus  male 15–19: 15. Dorsal view of cerci; 16. Ventral view of subgenital plate; 17. Ventral view of left cerci; 18. Lateral view of genitalia; 19. Dorsal view of tenth abdominal tergite; Female 20–22: 20. Dorsal view of tength abdominal tergite with cerci; 21. Lateral view of ovipositor; 22. Ventral view of subgenital plate.

22.i. 2014, Coll. Jhabar Mal ( TNAU, Coimbatore) (20); 20.i. 2014, Coll. Rajendra Nagar ( TNAU, Coimbatore) (5); 20.i. 2014, Coll. Jhabar Mal ( TNAU, Coimbatore), (4); 19.i. 2014, Coll. Jhabar Mal ( TNAU, Coimbatore); 04.. x. 2013, Coll. Rajendra Nagar (Chikaldhara) (4); 05.x. 2013, Coll. Rajendra Nagar (Chikaldhara) (6); 07.. x. 2013, Coll. Rajendra Nagar (Chikaldhara) (2); 04.x. 2013, Coll. Jhabar Mal, Patur (Akola) (15); 01.x. 2013, Coll. Rajendra Nagar (KVK, Chaitorgarh) (1); 01.x. 2013, Coll. Jhabar Mal (KVK, Chaitorgarh) (1); 01.ix. 2013, Coll. Rajendra Nagar ( ARSS, Vallabhanagar); 01.ix. 2013, Coll. Jhabar Ma ( ARSS, Vallabhanagar); 01.xi. 2012, Coll. Rajendra Nagar (Mandsaur) (1); 21.xii. 2013, Coll. Rajendra Nagar (RCA, Udaipur) (1); 20.x. 2015, Coll. R. Swaminathan, Ponmudi (Thiruvanthapuram), (2); 20.x. 2015, Coll. Rajendra Nagar, Ponmudi (Thiruvanthapuram) (4); 20.x. 2015, Coll. T. Swaminathan, Ponmudi (Thiruvanthapuram) (1); 21.x. 2015, Coll. Rajendra Nagar, ICAR, Myladumpara (Idukki) (7); 22.x. 2015, Coll. R. Swaminathan, ICAR, Myladumpara (Idukki) (1); 22.x. 2015, Coll. Rajendra Nagar, ICAR, Myladumpara (Idukki) (1); 23.x. 2015, Coll. Rajendra Nagar, Elappara, Vagamon Heights (Idukki) (1); 24.x. 2015, Coll. T. Swaminathan, Elappara, Vagamon Heights (Idukki) (2); Karnataka, 07.xi. 2013, Coll. Abhay Kumar Sharma (1), Bangalore ( GKVK); 10.v. 2011, Coll. C. Latha, Bangalore ( GKVK); 29.xi. 2010, Coll. P. Nirmala, Bangalore ( GKVK); 28.v. 2008, Coll. S. Murthy, Bangalore ( GKVK); 21.11. 2013, Coll. Geoby George, Bangalore ( GKVK); 21.11. 2013, Coll. A. N. Reddy, Bangalore ( GKVK) 25.vi. 2010, Coll. S. Murli, Bangalore ( GKVK); 0 7.11. 2013, Coll. Shwetha, Bangalore ( GKVK); 06.xi. 2010, Coll. Revanna Revannavar, Mudigere ( ZHRS); 08.x. 2014, Coll. Gavash Ragesh, Mudigere ( ZHRS).

Description. Male — Fastigium of vertex moderately wide, anterior margin of pronotum concave, posterior margin rounded. Prosternum bispinose, macropterous, forewing pigmented with comparatively large dark spots; no marking in costal and percostal area, cross veins of costal and perocostal area of forewings irregular to almost regular. Stridulatory file area of male left tegmen large, stridulatory file elevated on thickened vein, slightly rounded, about 52 large and 28 small teeth with main teeth very robust, which are large and spaced in the middle, gradually becoming narrower towards the apex. Hind femora unarmed ventrally. Tibia marked with spot towards base, dark on mid tibia, often faint on other. Hind tibiae with 6 apical spurs. Tenth tergite slightly broad, basal margin arched, concave, posterior margin with a pair of obtuse lobes. Cerci conical with spines in middle and at the apex small spine-like projection. Subgenital plate long with middle carinae, posterior margin incision in the middle; styli long, cylindrical, apices obtuse.

Female: Tenth abdominal tergite small, incision in middle with apical margins rounded. Epiproct small, triangular, rounded with apex obtuse. Cerci conical at base, suddenly narrowing, apex obtuse, hirsute. Subgenital plate slightly broader anteriorly and posterior narrowing posteriorly, flat, apical lobe slightly rounded. Ovipositor moderately straight and slightly curved upward, dorsal and ventral margins smooth, dorsal valve longer than ventral.

Distribution. India: Rajasthan; Karnataka; Kerala; Meghalaya; Madhya Pradesh; Gujarat; Maharashtra.

TNAU

Tamil Nadu Agricultural University

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Dryophthoridae

Genus

Conocephalus

Loc

Conocephalus (Anisoptera) maculatus (Le Gouillou, 1841)

Nagar, Rajendra & Swaminathan, R. 2016
2016
Loc

Conocephalus sinensis

Jin et Xia 1994: 33
1994
Loc

Xiphidion neglectum

Bruner 1920: 123
1920
Loc

Conocephalus maculatus

Bolivar 1913: 8
1913
Loc

Conocephalus (Xiphidion) maculatus

Karny 1912: 11
1912
Loc

Xiphidium maculatum

Matsumura et Shiraki 1908: 51
1908
Loc

Xiphidion maculatum

Karny 1907: 93
1907
Loc

Anisoptera maculatum

Kirby 1906: 278
1906
Loc

Xiphidium sinense

Walker 1871: 35
1871
Loc

Locusta (Xiphidium) continuum

Walker 1869: 271
1869
Loc

Locusta (Xiphidium) maculatum

Walker 1869: 275
1869
Loc

Locusta (Xiphidium) lepida

De Haan 1842: 189
1842
Loc

Xiphidium maculatum

Le Guillou 1841: 294
1841