Bartlett, Charles R. & Kennedy, Ashley C., 2018, A review of New World Malaxa (Hemiptera: Fulgoroidea: Delphacidae), Zootaxa 4441 (3), pp. 511-528: 516-517

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Lamaxa  , new genus

( Figures 7–18View FIGURES 7–14View FIGURES 15–18, 25–26 View Figure , 36 View Figure , 38, 40–42 View Figure )

Type species. Malaxa occidentalis Muir, 1926  .

Description. Color. Brownish-orange with brown and paler markings; wings clear with characteristic dark (and sometimes pale) markings. Structure. Body slender and elongate; length (from apex of vertex to tip of tegmina) males x‾ = 4.97 mm (4.71–5.33 mm, n=5); females x‾ = 5.31 mm (5.10–5.62 mm, n=6). Head much narrower than pronotum ( Figs. 7, 10 View Figure ), carinae concolorous. Lateral carinae of clypeus, frons (~metope) and vertex (~coryphe) distinct, other carinae obscure, especially near fastigium. Frons elongate and parallel-sided, widest near frontoclypeal suture, narrowed to fastigium ( Figs. 9, 12 View Figure ; l:w x‾ = 2.6:1). Vertex longer than broad at base (l:w x‾ = 1.4:1), distally narrowing and projected slightly in front of eyes; submedian carinae uniting before fastigium, basal compartments longer than wide. Rostrum reaching or exceeding metathoracic trochanters. Antennae cylindrical, both segments very long ( Figs. 10, 12 View Figure ), surpassing apex of clypeus (and apex of mesonotum), segment I close to half length of II (ratio I:II 0.4–0.5:1), length antennal segment I x‾ = 0.47 mm, II  = 1.05 mm. Pronotum shorter than vertex in middle line, lateral carinae usually not attaining hind margin. Mesonotum longer in middle line than vertex and pronotum together. Wings elongate, much longer than abdomen, nodus at about 2/3 length, apex acutely rounded ( Fig. 36 View Figure ). Spinal formula of hind leg 5-7-5 (or 5-6-5). Calcar thick, concave on inner surface, without teeth along the hind margin, with an apical tooth. Pygofer irregularly quadrilateral in lateral view ( Fig. 26 View Figure ). In caudal view, opening carinate, with midventral forked process ( Figs. 13 View Figure , 25 View Figure ), broad lamellate medioventral processes absent. Dorsocaudal margins of pygofer expanded and inflected to partially enclose anal tube ( Fig. 42 View Figure ), diaphragm evident, armature absent. Gonostyli broad basally, basal angles prominent, diverging and narrowed distally to apically to blunt or acute apex ( Figs. 25 View Figure , 40 View Figure ). Aedeagus elongate and tubular with large, flattened, poorly sclerotized subapical process ( Figs. 14 View Figure , 38 View Figure ; representing a flagellum?). Anal tube (anal segment) small, without processes.

Remarks. Lamaxa  differs from Malaxa acutipennis  in having a parallel-sided wing with a rounded apex (vs. pointed with an arched leading wing edge in Malaxa acutipennis  ); antennae are similar in proportion but nearly twice as long in Lamaxa  than Malaxa acutipennis  (and lacking the ventral carina on I); the genitalia of Lamaxa  bear a forked projection on the pygofer opening, whereas Malaxa acutipennis  is simple; and the dorsocaudal margin of the pygofer is expanded in Lamaxa  but not in Malaxa  . The aedeagus of Lamaxa  is slightly curved and caudally directed, bearing weakly sclerotized apical projections (vs. downcurved in Malaxa acutipennis  ).

Lamaxa  differs from the Chinese Malaxa  in structural proportions along with distinctive features of the male genitalia. Lamaxa  is larger than the Chinese Malaxa  , the vertex is more elongate, the lateral carinae of the pronotum are laterally curved, usually not reaching the hind margin, and the spinulation of the hind leg is 5-7-5 (or 5-6-5) (instead of 5-6-4). More distinctly, in Lamaxa  , the anal tube is without processes, whereas in Chinese Malaxa  , the anal tube bears a single (sometimes 2), large usually strongly asymmetrical process(es). In Lamaxa  , the dorsocaudal angle of the pygofer is projected (simple in Chinese Malaxa  ), and the midventral portion of the pygofer opening bears a forked process, whereas in Chinese Malaxa  , there are “two broad lamellate medioventral processes, between of them with a V-like emargination” ( Hou et al. 2013: 867), although these are reduced in some species. In Chinese Malaxa  , the aedeagus is downcurved and C-shaped, whereas in Lamaxa  the aedeagus is slightly curved and caudally directed, bearing weakly sclerotized apical projections, missing from Chinese Malaxa  .

Species included are Lamaxa gracilis ( Fennah, 1945)  and Lamaxa occidentalis ( Muir, 1926)  . Two female specimens, probably representing additional species of Lamaxa  were examined (listed below under ‘females tentatively assigned to Lamaxa  ’). One was a specimen from Honduras that was similar to the coloration of Lamaxa occidentalis  , except that the face (frons, genae, clypeus) were broadly black except for a strongly contrasting pale stripe above the frontoclypeal suture, also bearing more substantially embrowned wings. The second, from southern Brazil, was very pale yellowish with weakly marked wings, a dark longitudinal line along both segments of the antennae, and partially embrowned legs.

Etymology. The new genus name “ Lamaxa  ” is an anagram of “ Malaxa  ”, and is to be understood as feminine in gender.