Xalama microstyla Muir, 1930: 211

Bartlett, Charles R. & Kennedy, Ashley C., 2018, A review of New World Malaxa (Hemiptera: Fulgoroidea: Delphacidae), Zootaxa 4441 (3), pp. 511-528: 520-523

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Xalama microstyla Muir, 1930: 211


Xalama microstyla Muir, 1930: 211 

( Figures 19–24 View Figure , 27–29 View Figure , 37 View Figure , 39 View Figure )

Type locality. San Fermin , Bolivia. 

Amended diagnosis. Color. Body generally deep brown ( Figs. 19, 21, 23 View Figure ); head, legs and dorsum of pronotum paler; carinae concolorous; vertex and dorsal portion of frons and genae orangish (bearing pale middorsal vitta on posterior vertex, extending posteriorly onto pronotum); ventral portions of frons and genae cream color; antennae stramineous. Disc of pronotum orangish brown except paler median vitta. Legs stramineous except darker proximally; venter of abdomen orangish. Wings clear with strong fuscous markings, one brown bar across apex of clavus to past fork of CuA; a second elongate curved marking extending diagonally along nodal line to wing margin, then diagonally across wing apex to costal margin. Structure. Body slender and elongate; length (from apex of vertex to tip of tegmina) males  = 4.69 mm (4.40–4.88 mm, n=5); females not available. Head narrower than pronotum ( Figs. 19, 23 View Figure ), carinae concolorous. Carinae distinct although median carina of vertex weaker. Frons elongate and parallel-sided ( Fig. 24 View Figure ; l:w  = 2.5:1, range 2.4–2.8:1, n=5), narrowed between eyes (width  = 0.17, n=5), widest near frontoclypeal suture (w  = 0.22, n=5). Vertex narrowing distally, slightly projecting in front of eyes, vertex broader caudally than long along midline (l: [basal] width  = 0.74:1, range 0.65–0.91:1, n=5); submedian carinae distinct, uniting before fastigium, basal compartments longer than wide. Rostrum reaching hind coxae. Antennae cylindrical, segment II long, nearly surpassing apex of clypeus ( Figs. 20, 24 View Figure ); segment I about 1/3 length of II (ratio I:II 0.37:1, range 0.28–0.45:1, n=5), length antennal segment I  = 0.13 mm (range 0.10–0.17, n=5), II  =0.34 (range 0.31–0.37, n=5). Pronotum subequal or just shorter than vertex along midline, lateral carinae usually not attaining hind margin. Mesonotum longer in middle line than vertex and pronotum together (1.91–2.33: 1, n=5). Wings elongate ( = 3.8 mm, range 3.2–4.1, n=5), much longer than abdomen, predominately clear with distinctive dark markings, wing apex rounded ( Fig. 37 View Figure ); nodus at about 2/3 length; Sc, RA and RP unbranched; MP branched near wing apex, CuA 3-branched; junction of PCu + AA near midlength of clavus; fork of MP+CuA at near 2/3 length of clavus; fork RA+SC and RP near claval apex (thus inner subapical cell longer than outer subapical cell). Spinulation of hind leg 5-6-4. Pygofer roughly triangular in lateral view ( Figs. 27 View Figure , 39 View Figure ), elongate ventrally and strongly narrowed dorsally; in caudal view, pygofer opening with midventral and lateral projections; midventral projection large and rounded, scoop-like; lateral projections just below midlength in form of rounded tooth in lateral view. Diaphragm extending from level of lateral projections and broadly concave beneath gonostyli (weak or absent above gonostyli). Gonostyli flattened and curled (apically scoop-like), irregularly parallel-sided, curved medially, bearing low, rounded tooth medially near midlength; apically broader and more flattened, somewhat angular, bearing find, irregular serrulations on inner apical margin fine; irregularly placed setae. Aedeagus elongate and robust, strongly downcurved into long, sclerotized structure, slightly twisted, bearing an irregular row of teeth on left and right sides and long, downward projecting process at curve. Anal tube (anal segment) small and simple.

Remarks. Differences between Xalama  , Lamaxa  , and Malaxa  have been previously provided in the remarks under the later 2 genera. The much shorter and differently proportioned antennae and the very different form of the genitalia (viz. the strongly downcurved aedeagus and the projections on the ventral and lateral opening of the pygofer) are the most diagnostic features of the genus.

Material examined. Type material. Holotype ( Fig. 22 View Figure ) " S. Fermin / Boliv. // N. Holmgren // Typus [red paper] // Malaxa  / microstylus / [male symbol] Muir [handwritten] // 79 / 81 / Riksmuseum / Stockholm / NHRS-GULI / 000011301". Other material examined. PERU: Chanchamayo, San Ramon, 11.12083°S 75.35361°W, 800 m, 17 Oct 2002, R. Rakitov & C. H. Dietrich (6 males, INHSAbout INHS)GoogleMaps  .


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