Bryaxis goliath ( Jeannel, 1922 )

Hlaváč, Peter, 2009, Taxonomic notes on the Bryaxis splendidus species group (Coleoptera: Staphylinidae: Pselaphinae), with the description of a new species from the Ukraine, Acta Entomologica Musei Nationalis Pragae 49 (2), pp. 651-659 : 655-656

Bryaxis goliath ( Jeannel, 1922) , stat. restit.

( Figs. 5 , 8 )

Megalobythus goliath Jeannel, 1922: 233 . Type locality: Romania, cave Corobana Mandraţului, comm. de Scărişoara, jud. Turda-Arieş.

Bryaxis splendidus goliath Jeannel : MACHULKA (1935: 131).

Note. MACHULKA (1935) treated this species as a variety of B. splendidus . I had not the posibility to see the species but the shape of the apices of the parameres as illustrated in JEANNEL (1955: 40, Fig. 17b) and reproduced in Fig. 5 combined with the large body size indicate that B. goliath is a distinct species.

Distribution. Known only from the type locality, Corobana Mandraţului cave in Munti Apuseni, Romania

Bryaxis monstrosetibialis ( Stolz, 1923)

( Figs. 6 , 8 )

Bythinus monstrosetibialis Stolz, 1923: 71–72 . Type locality: Bad Rosenau in Ungarn (Komitat Gömör [Gemer region in southern Slovakia]).

Bythinus Trojani Fleischer, 1925:17–18 . Type: Slovakia, ‘okraj Demänovských jeskyň [edge of Demänovská cave]’, synonymised by MACHULKA (1929c: 120–121).

Megalobythus Trojani Fleischer : MACHULKA (1929a: 22; 1929b: 23–24).

Bythinus (Macrobythus) monstrosetibialis Stolz : ROUBAL (1930: 507–508).

Bythinus monstrosetibialis Stolz var. ocularis Machulka, 1935: 131 . Type locality: Slovakia, ‘okolie dobšinské jeskyně [surroundings of Dobšiná cave]’, synonymised by LÖBL & BESUCHET (2004: 307).

Balcanobythus (Megalobythus) monstrosetibialis Stolz : KARAMAN (1957: 187, 208, Fig. 6 (aedeagus)).

Bryaxis monstrosetibialis Stolz : BESUCHET (1974: 349, Fig. F. Basis 18: 26 (first two antennal segments)); HLAVÁČ, (2000: 37, new records, biology, aedeagus and habitus drawings); FRANC & MLEJNEK (2000: 31–34, new records,

biology). New records (all data provided by Martin Švarc; all specimens deposited in his collection). SLOVAKIA: 13 JJ 17

♀♀: Hrochoť , Hrochoťská dolina [valley], sifting of leaf-litter in cave, Švarc lgt .; 4 JJ 2 ♀♀: Liptovský Mikuláš , Demänovská jaskyňa [cave], under stones and sifting leaf-litter in cave, Švarc lgt .; 2 spec.: Muráňska planina, jaskyňa Maretkina [cave], Orszulík lgt .; 1 J: Slovenský raj, jaskyňa Duča [cave], trap, Košel lgt .; 3 JJ: Závadka nad Hronom , Veľká Stožka, Machnatá dolina [valley], sifting of litter near spring, Krátky lgt .; 1 ♀: Muráň , Javorníkova dolina [valley], sifting Fagetum, Mantič lgt .; 1 J: Kremnické vrchy, Diera pod oknom, 1190m, Mlejnek lgt .; 1 J: Nízke Tatry Mts. , jaskyňa nad Patočinami [cave above Patočiny], 3,700 m a.s.l., Mlejnek lgt .; 1J: Malá Fatra , Veľký Rozsutec, Brestovanský lgt .; 2 ♀♀: Nízke Tatry Mts. , Demänovská dolina [valley], Orszulík lgt .

Records from the literature. ROUBAL (1930) and MACHULKA (1929a,b,c, 1935) summarized old data from the following localities: near Rožňava, near Košice, Gemer (exact locality not available), Krupina, Nová Baňa, Muráň, and Dobšiná. Additional recent record was published from Zádieľ in the eastern Slovakia by MAJZLAN & RYCHLÍK (1994).A decade ago I summarized all old data and added some other localities: Slovenský kras (exact locality not available), Tisovec env. (Michňová jama), Handlová, Plešivecká planina (Lastovičia jama) ( HLAVÁČ 2000). Finally, FRANC & MLEJNEK (2000) studied the ecology and distribution of the species and mentioned the following localities: Malá Fatra (lúky nad Hornými dierami [meadows above Horné diery]); Sokolec; Fačkovské sedlo [saddle], Plešivecká planina (Lastovičia priepasť [abyss]), Revúcka vrchovina (jaskyňa [cave] Burda,Veľká drienčanská jaskyňa [cave]), Poľana (Hrochoťská jaskyňa [cave]), Horehronské podolie: Valaská (jaskyňa v kameňolome [cave in stone-pit] Potôčky), Horehronské podolie: Červená skala (Márnikova jaskyňa [cave]), Starohorské vrchy (jaskyňa pri Uľanke [cave near Uľanka]), Nízke Tatry (Demänovská dolina [valley], Tunelová jaskyňa [cave]).

Distribution. Slovakia.

Figures

Figs. 2–7. 2–4 – Bryaxis cepliki sp. nov. 2 – scape and pedicel; 3 – aedeagus, dorsal aspect; 4 – aedeagus, ventral aspect. 5 – B. goliath (Jeannel, 1922), aedeagus, dorsal aspect (reproduced from JEANNEL 1955: 40). 6 – B. monstrosetibialis (Stolz, 1923), aedeagus, dorsal aspect (from HLAVÁČ 2000: 38). 7 – B. splendidus (Croissandeau, 1891), aedeagus, dorsal aspect. Scale bars = 0.2 mm.

Fig. 8. Bryaxis splendidus species-group, distribution (black circle – B.monstrosetibialis (Stolz, 1923); white circle – B. cepliki sp. nov.; black square – B. splendidus (Croissandeau, 1891); white square – B. goliath (Jeannel, 1922)).