Orcevia yahaha Yu, Maddison & Zhang, 2023

Yu, Kun, Maddison, Wayne P. & Zhang, Junxia, 2023, Taxonomic revision of Orcevia Thorell, 1890, with description of fifteen new species (Araneae, Salticidae, Euophryini), Zootaxa 5384 (1), pp. 1-79 : 17-18

Orcevia yahaha Yu, Maddison & Zhang , sp. nov. (呀ëĸ尔¤)

Figs 7 , 315–331

Type material. Holotype: ♂ ( UBCZ; KYU-SAL700), MALAYSIA: Sarawak, Mulu Nat. Pk., headquarters area, 4.042°N, 114.814°E, 50 m elev., 25–27 March 2012, leg. W.P. Maddison, E.K. Piascik & A. Ang, WPM#12-094 . Paratypes: 1♂ 1♀ ( UBCZ; KYU-SAL701~KYU-SAL702), same data as holotype ; 1♂ ( UBCZ; KYU-SAL703), MALAYSIA: Sarawak, Mulu Nat. Pk., Canopy Walk , 4.032°N, 114.819°E, 80 m elev., 26 March 2012, leg. W.P. Maddison, E.K. Piascik & A. Ang, WPM#12-099 .

Etymology. The specific epithet is a noun in apposition from “Yahaha” (also known as “Korok”), a very cute pixie in the game The Legend of Zelda: Breath of the Wild. Yahaha often hide in tree trunks, bushlands, or rocks, and if you find them, they will say “Yahaha!” and share with you some “fruits” as a gift. Collecting Orcevia specimens is very like searching for Yahaha.

Diagnosis. It can be distinguished from all other congeners by its long and baffy-like embolus ( Figs 322 , 326 ), and the small and ligulate embolic disc which is well defined from the base of embolus ( Fig. 329 ). Females resembles O. calcicola sp. nov. in the relatively large atria close to genital furrow; however, they can be distinguished by the copulatory ducts that extend upward first, further down and then connect with spermathecae ( Fig. 331 ; vs. CDs extend upwards and directly connect with spermathecae in O. calcicola sp. nov.).

Description. Male. Habitus as shown in Fig. 319 . Measurements of holotype: carapace length 1.92; abdomen length 1.64; measurements of eyes: AME 0.40, ALE 0.25, PME 0.06, PLE 0.21; measurements of legs: I 4.21 (1.24, 0.83, 1.03, 0.68, 0.43), II 2.99 (0.93, 0.56, 0.63, 0.53, 0.34), III 3.12 (0.95, 0.49, 0.58, 0.73, 0.37), IV 3.32 (1.00, 0.47, 0.72, 0.75, 0.38); leg formula 1432. Dorsal body covered with dense white setae; legs I–II obviously darker than III–IV.

Palp ( Figs 321–323 ): embolus long and baffy-like, distal half with many small protrusions on retrolateral surface; embolic disc small and ligulate, well defined from embolus ( Fig. 329 ); lobe-like protrusion of tegulum well developed; tegular lobe well defined from main part of tegulum, with prolateral and retrolateral constrictions close to tegulum; RTA narrow in retrolateral view, bent medially; ventral femoral bump hook-like as in O. zabkai sp. nov. ( Fig. 342 ), distal part blunt.

Female. Habitus as shown in Fig. 320 . Measurements of paratype: carapace length 1.55; abdomen length 1.66; measurements of eyes: AME 0.33, ALE 0.22, PME 0.04, PLE 0.17; measurements of legs: I 2.41 (0.75, 0.45, 0.53, 0.37, 0.31), II 2.06 (0.68, 0.34, 0.40, 0.37, 0.27), III 2.50 (0.82, 0.36, 0.49, 0.50, 0.33), IV 2.71 (0.87, 0.37, 0.60, 0.55, 0.32); leg formula 4312. Dorsal body covered with dense golden setae; legs relatively light-colored, melanic area indistinct.

Epigynum ( Fig. 324 ): Atria close to each other and close to genital furrow; outer edge of copulatory ducts faintly visible. Vulva ( Fig. 325 ): Accessory glands rather small, hiden in ventral side of spermathecae and invisible in dorsal view of vulva; copulatory ducts flat and rolled, weakly sclerotized; spermathecae far from each other, fertilization ducts attached on small mound on spermathecae.

Distribution. Malaysia ( Sarawak).

Figures

FIGURES 1–12. Comparison of male palpal bulbs of some euophryines, in ventral view. 1–8. Orcevia spp. [O. bokoblin Yu, Maddison & Zhang sp. nov. (1), O. nietzschei Yu, Maddison & Zhang sp. nov. (2), O. jinping Yu & Zhang, sp. nov. (3), O. mercuryi Yu, Maddison & Zhang, sp. nov. (4), O. shuyuani Yu & Zhang, sp. nov. (5), O. zabkai Yu, Maddison & Zhang, sp. nov. (6), O. yahaha Yu, Maddison & Zhang, sp. nov. (7) and O. wuliang Yu & Zhang, sp. nov. (8)]; 9. Laufeia concava Zhang & Maddison, 2012; 10. Amoenema erhai Yu & Zhang, 2023; 11. Lokina nyuewa Yu & Zhang, 2023; 12. Chalcovietnamicus vietnamensis (Żabka, 1985). Black dotted lines indicate retrolateral edge of membrane between salticid radix and tegulum, white arrows point to the retrolateral extension, and yellow lines indicate the proximal margin of tegulum.

FIGURES 315–318. Living male (315–317) and female (318) of Orcevia yahaha Yu, Maddison & Zhang, sp. nov.. Copyright © 2015 W.P. Maddison, released under a Creative Commons Attribution (CC-BY) 3.0 license.

FIGURES 319–325. Orcevia yahaha Yu, Maddison & Zhang, sp. nov., holotype male (319, 321–323) and paratype female (320, 324–325). 319–320. Habitus; 321–323. Palp; 324. Epigynum; 325. Vulva; in dorsal (319–320, 325), prolateral (321), ventral (322, 324) and retrolateral (323) view.

FIGURES 326–331. Orcevia yahaha Yu, Maddison & Zhang, sp. nov., holotype male (326–327), paratype male (328–329) and paratype female (330–331), male palp (326–327), expanded palpal bulb (328–329), 330–331. Transparent epigynum; in ventral (326, 330–331), retrolateral (327) and dorsal (328) view. Abbreviations: AG, accessory gland; CO, copulatory opening; CD, copulatory ducts; DH, distal haematodocha; E, embolus; ED, embolic disc; LPT, lobe-like protrusion of tegulum; RTA, retrolateral tibial apophysis; SR, salticid radix; ST, subtegulum; T, tegulum; TL, tegular lobe.

FIGURES 338–345. Orcevia zabkai Yu, Maddison & Zhang, sp. nov., holotype male (338), paratype male (340–342) and female (339, 343–345). 338–339. Habitus; 340–342. Palp, with arrows indicating the row of serrula on embolus, and the small bump near base of RTA; 343–344. Epigynum; 345. Vulva; in dorsal (338–339, 345), prolateral (341), ventral (340, 343–344) and retrolateral (342) view. Abbreviations: AE, apophysis of embolic disc; AG, accessory gland; CD, copulatory duct; E, embolus; ED, embolic disc; LPT, lobe-like protrusion of tegulum; pH, pro-ventral hump of male palpal tibia; RTA, retrolateral tibial apophysis; S, spermatheca; SR, salticid radix; T, tegulum; TL, tegular lobe; VFB, ventral femoral bump of male palp.