Dilatitibialis cognata (Mulsant) Canepari & Gordon & Hanley, 2013

Canepari, Claudio, Gordon, Robert D. & Hanley, Guy A., 2013, South American Coccinellidae (Coleoptera), Part XV: systematic revision of Dilatitibialis Duverger (Coccidulinae; Hyperaspidini), Insecta Mundi 2013 (312), pp. 1-91 : 52-53

44. Dilatitibialis cognata (Mulsant) , new combination

Cleothera cognata Mulsant, 1850: 545 .

Hyperaspis cognata: Crotch 1874: 214 ; Korschefsky 1931: 186; Blackwelder 1945: 27; Gordon 1987: 27.

Description. Male. Length 4.0 mm, width 3.25 mm; body oval, convex. Color bright orange except pronotum irregularly black in basal 1/2; elytron with suture narrowly black except black area dilated anterior to middle, lateral and apical borders narrowly black, irregular black vitta present from behind humeral callus to apical 1/3 ( Fig. 240 ); ventral surface with head, prosternum, meso- and metaventrites black, proleg yellow, meso- and metalegs with femur dark brown in basal 2/3, tibia yellow, abdomen dark brown. Head punctures small, separated by less than a diameter, each puncture as large as 3 eye facets; pronotal punctures same size as on head, separated by less than to twice a diameter; elytral punctures smaller than on pronotum, separated by 2 to 4 times a diameter; metaventral punctures 3 or 4 times as large as on pronotum, separated by a diameter medially, becoming larger and contiguous toward lateral margin. Clypeus very slightly emarginate apically, nearly truncate, lateral angle broadly rounded, surface with dense, long pubescence. Eye canthus about 4 eye facets long, angled forward, apically rounded, yellow. Pronotum narrowed from base to apex, sides feebly rounded, basal and anterior angles rounded, basal margin without bordering line. Epipleuron narrow, grooved, descending externally, deeply emarginate for reception of femoral apices. Protibia narrowly flanged, flange evenly arcuate, narrower than remainder of protibia, outer margin smooth ( Fig. 241 ); sponda slightly wider than flange. Carinae on prosternal process pronounced, narrowly spaced apically, convergent toward base, joined at basal 2/3 of prosternum, connected to prosternal base by single carina. Metaventrite without median setal tuft. Abdomen with primary pores laterally between ventrites 4-5 large, occupying apical 1/3 of ventrite 4, forming slight depression on ventrite 5; postcoxal line on basal abdominal ventrite straight in basal 1/3, extended to apical margin of ventrite at middle, rounded in apical 2/3, extended forward; ventrites 2-4 with sparse, short pubescence and dense, fine punctures, 5th ventrite not depressed medially, unmodified, apical margin broadly, weakly emarginate, surface densely punctate; 6th ventrite short, narrow, depressed in apical 1/2, apical margin broadly, weakly emarginate, lateral angle of emargination projecting, widely rounded, surface densely punctured. Apical tergite densely punctured, with shallow median groove. Genitalia with basal lobe about as long as paramere, asymmetrical, slender, apex obliquely truncate; paramere Psc, wide ( Fig. 242, 243 ); sipho slender, strongly curved in basal 1/2, basal capsule lightly sclerotized, inner arm short, narrowed in apical 1/2, outer arm longer than inner arm, wide, with large accessory piece, basal border broadly emarginate ( Fig. 244, 245 ).

Female. Unknown.

Variation. Unknown.

Type locality. Brazil.

Type depository. MNHP (lectotype here designated).

Geographical distribution. Brazil.

Specimens examined. 1. Brazil. No further data. ( MNHP) ( UMZC).

Remarks. The dorsal color pattern is extremely distinctive, and along with the large size, will serve to distinguish D. cognata from its congeners.

Mulsant (1850) stated that the female type was in the MNHP collection. That lectotype was examined in 1971, but is not presently available, so the description above is from a single specimen in the UMZC that matches the original description quite well. The lectotype is labeled “Museum Paris, Ouest Capit des Mines, 147, Cleothera cognata Muls. , auct. det.” That specimen also bears a 1971 lectotype label that is here validated.

Figures

Figures 235-250. 235-239) Dilatitibialis sherry. 235) Habitus. 236) Phallobase ventral. 237) Phallobase lateral. 238) Sipho. 239) Enlarged siphonal apex. 240-245) Dilatitibialis cognata. 240) Habitus.241) protibia.242) Phallobase ventral. 243) Phallobase lateral. 244) Sipho. 245) Enlarged siphonal apex. 246-250) Dilatitibialis sylvia. 246) Habitus. 247) Phallobase ventral. 248) Phallobase lateral. 249) Sipho. 250) Enlarged siphonal apex.