Paramaka Savage & Peters, 1992

Salles, F. F., Domínguez, E., Mariano, R., Boldrini, R., Clavier, S. & Lima, L. R. C., 2025, A review of Paramaka Savage & Peters, 1992 (Ephemeroptera: Leptophlebiidae) with the description of two new species, Zootaxa 5590 (3), pp. 401-421 : 402-403

Paramaka Savage & Peters, 1992

Thraulus [partim]; Spieth, 1943: 10.

Hermanella [partim]; Demoulin, 1966: 12.

Paramaka Savage & Domínguez, 1992: 244 ; Blanco-Belmonte et al., 2003: 117; Sartori, 2005: 243; Domínguez et al., 2006: 463; Mariano, 2011: 46; Domínguez et al., 2014: 308.

Diagnoses. Male imago: 1) eyes meeting on meson of head ( Figs. 3C, 4A, 7A); 2) cross vein above fork of MA slanted ( Fig. 8A); 3) fork of MA and MP asymmetrical ( Fig. 8A); 4) penes divided, each lobe with ( Figs. 2E, 4D) or without ( Figs. 7E, 8C, 8D) a long spine ventromedially directed [telopenis according to Kluge (2007)]; 5) posteromedial projection of styliger plate single, short triangular ( Figs. 2E, 4D) to long and lanceolate ( Figs. 7E, 8C); and 6) female sternum IX with distal margin widely cleft.

Nymph: 1) labrum greatly expanded laterally, 1.3–2.2x maximum width of clypeus ( Figs. 2A, 2B, 3B, 10A); 2) maxilla with short tusk at distomedial margin ( Fig. 10D); 3) gills plate-like, apically rounded to obtuse, with or without a pointed projection; 4) gills present on abdominal segments I to VI and vestigial or absent on abdominal segment VII; 5) posterolateral projections present on segments VII or VIII to IX.

Distribution. Neotropical, from Southeastern Brazil (Minas Gerais) to Northern South America (Central Colombia to Northern Venezuela) ( Fig. 1).

Species included. Paramaka antonii Sartori, 2005 ; Paramaka convexa ( Spieth, 1943) (type-species); Paramaka incognita Domínguez, Grillet, Nieto, Molineri & Guerrero, 2014 ; P. froehlichi Lima & Salles, 2023; Paramaka pearljam Mariano, 2011 ; Paramaka takari sp. nov.; and Paramaka lucimarae sp. nov.